| 0 | PI 664881 | 6 | Prunella vulgaris L. | Iowa, United States | NC7 | | | 2006 | COLLECTED | 10/02/2006 | Near the east end of Reindeer Ridge, Ledges State Park, NE 1/4 of NE 1/4 of NW 1/4 of Section 21, T83N, R26W, Worth Township, Boone County. | 41.99388889, -93.88444444 | 329 | Large slump below sandstone cliff outcrop. Few feet up from Pea's creek floodplain. Partial sun. Northwest facing. Moist-wet clay soil. | Wild material | Plants 12 inches tall. | 1712926 | PI 664881 |
| 1 | PI 664882 | 9 | Prunella vulgaris L. | Iowa, United States | NC7 | | | 2006 | COLLECTED | 09/22/2006 | 300 yards south of Carl Fritz Henning Shelter House, 300 yards northeast of Lost Lake parking lot, Lower Ledges Road, Ledges State Park, NW 1/4 of NE 1/4 of SW 1/4 of Section 21, T83N, R26W, Worth Township, Boone County. | 41.98555556, -93.88833333 | 329 | Cleared woods for powerline right-of-way. Moist-wet mostly clay soil. Plants scattered over entire hilltops. Site approximately 250 yards by 30 feet. | Wild material | Plants 16-18 inches tall. | 1712928 | PI 664882 |
| 2 | PI 664883 | 10 | Prunella vulgaris L. | Iowa, United States | NC7 | | | 2006 | COLLECTED | 09/15/2006 | Downhill, east-southeast of primitive campground parking lot, Ledges State Park, NE 1/4 of SW 1/4 of SW 1/4 of Section 15, T83N, R26W, Worth Township, Boone County. | 41.99638889, -93.87111111 | 329 | 3-4 small springs at bottom of west facing hillside and clay slump on west side of a deep gulley. Dense woods. Very wet soil. | Wild material | Plants up to 11 inches tall. | 1712929 | PI 664883 |
| 3 | PI 656805 | 001 | Cornus alternifolia L. f. | Iowa, United States | NC7 |  | | 2005 | COLLECTED | 07/14/2005 | Ledges State Park, Boone County. | 41.99666667, -93.88166667 | 328 | Mostly shaded, exposed over rock cliffs, next to small creek. 55% slope with a northern aspect. Well drained soil, predominately rock substrate. | Wild material | Plants 6-10 feet in height, clean foliage, some canker. | 1685173 | PI 656805 |
| 4 | PI 649661 | 23 | Diervilla lonicera Mill. | Iowa, United States | NC7 |  | | 2003 | COLLECTED | 09/15/2003 | Steamboat Rock County Park just southwest of Steamboat Rock, Steamboat Rock Quad. T88N R19W N edge of SE 1/4 SW 1/4 Sec. 28 | 42.40138889, -93.07305556 | 299 | Eroding sandstone cliff. Mostly shaded area with 50% slope with a northern aspect. Sand and organic matter just above sandstone rock cliff, well drained. | Wild material | | 1649456 | PI 649661 |
| 5 | PI 656820 | Ames 27186 | Physocarpus opulifolius (L.) Maxim. | Iowa, United States | NC7 |  | | 2003 | COLLECTED | 08/24/2003 | Along Old Indian Trail, overlooking Pease Creek, Ledges State Park, S 1/2 of SE 1/4 of SW 1/4 of Section 16, T83N, R26W, Madrid NW Quad, Boone County. | 41.99500000, -93.88583333 | 289 | Sandstone ridge. | Wild material | Shrubs to 2 meters tall. | 1648564 | PI 656820 |
| 6 | Ames 36073 | JDC/UT/2022/028/1009 | Ulmus thomasii Sarg. | Iowa, United States | NC7 |  | | 2023 | COLLECTED | 06/05/2023 | Ledges State Park approximately 4.75 miles south of Boone, T83N R26W (Madrid NW Quad.) near center of SE ¼ Sec. 16 and also NW ¼ SW ¼ Sec. 21. | 41.99780200, -93.87967900 | 288 | Moist deciduous woods typically on mid-slopes to base of slopes and also well-drained floodplains. | Wild material | Ulmus thomasii (Rock elm) is an underutilized tree species both in conservation and in the urban landscape. Charles Sprague Sargent (first director of the Arnold Arboretum of Harvard University, Boston, MA) described the wood of Rock Elm as "heavy, hard, very strong and tough, closely grained, and susceptible of receiving a beautiful polish". Historically noted for a variety of uses in woodworking including furniture, heavy agricultural implements, timber frame buildings, pianos, hockey sticks, and hand tools. U. thomasii is extremely drought tolerant, adaptable to very calcareous soils, and compared to U. americana or U. rubra has a very conservative growth rate requiring minimal attention in regards to pruning. Rock elm naturally tends to develop into a nice structural specimen in the landscape, unlike other fast growing U. americana cultivars in the nursery trade needing frequent attention. Rock elm has very fissured bark and is well known for maintaining a strong central leader resulting in a very attractive, pyramidal form. Despite many literature sources stating that "wings" on the stems are a key identifiying characteristic, this trait is not reliable and many genotypes may completely lack "wings" on the stems. While growth habit and morphology of dormant buds (very pointy) are good traits to help aid in identification, fruits are the best way to confirm identity. Fruits are densly pubescent, often described as "inflated" and generally reach maturity a couple weeks after U. rubra and U. americana. U. thomasii flowering in central Iowa typically occurs the last week in April into the first week in May and are rather distinguishable from other Ulmus in being pendulous. Fruits are extremely prized treats for many birds, where fruits are often "robbed" prior to full maturity. Fruits are also highly prized by many other animals such as mice, squirrels, chipmunks, turkeys, etc. and are generally consumed within hours after fruit abscission. The native range of U. thomasii is concentrated in the northeastern United States and into Canada with disjunct populations documented in northwestern Missouri and northwestern Arkansas (Little, E. L. Jr. 1971). In the 1895 Silva of North America (vol. 7) it was stated that, “The value of the wood of the Rock Elm threatens its extinction and most of the large trees have already been cut.” Today, since rock elm is rather rare, especially in areas near the edges of its range, many states list the species as endangered or threatened warranting preservation efforts. Genetics of this accession would be representative of the species southwestern edge of its native range. Out of approximately 100 specimens noted throughout Ledges State Park only 1-2% of specimens had characteristic "wings" on the stems. Research was executed (Moore, P., 2024) involving the core sampling of U. thomasii and U. americana specimens in Ledges State Park to make growth ring widths and basal area increment comparisons. Results determined that the average age of cored trees U. thomasii (n=4) and U. americana (n=4) of both species was roughly 80 years, excluding the exceptional case of tree #007 (Ames 36073 23ncao02 SD). Tree #007 was aged at 300 years of age and currently (2025) ranks as the 14th largest U. thomasii in Iowa measuring 21" DBH with a height of 75' 5" and a spread of 43' 5" totalling 152.21 points (Rouw, M., personal communication, 4 May 2022). The computed mean density at 𝑀𝑀 = 12% for U. thomasii was 0.80 g/cm3, and for U. americana was 0.672 g/cm3. These values are both higher than the representative values inferred from the Woods Database which are 0.755 g/cm3 for U. thomasii and 0.56 g/cm3 for U. americana. To date, there is little to no information regarding U. thomasii resistance to Dutch Elm Disease. Collections of U. thomasii at the Arnold Arboretum have over-time succumbed to DED (Pruka, B., 2016). However, data recorded by Moore (2024) indicates that U. thomasii growing in central Iowa can surpass 300 years in age verifying it's long-term utilization in the landscape. | 2142915 | Ames 36073 |
| 7 | Ames 35996 | AS/QR/2022/067/199 | Quercus rubra L. | Iowa, United States | | | Historic | 2022 | COLLECTED | 09/27/2022 | Ledges State Park, approximately 4.5 miles south of Boone, T83N R26W Sec. 20 | 41.99504700, -93.88304400 | 330 | Hardwood forest. | Wild material | | 2141429 | Ames 35996 |
| 8 | Ames 35734 | Ledges State Park | Teucrium canadense L. | Iowa, United States | NC7 |  | | 2021 | COLLECTED | 08/23/2021 | Ledges State Park located approximately 4.75 miles south of Boone T83N R26W Sec. 16 & 21 | 41.99262400, -93.87724800 | 330 | Edges of woods | Wild material | | 2120338 | Ames 35734 |
| 9 | WLP 1685 | JDC/RS/2019/036/915 | Rosa setigera Michx. | Iowa, United States | NA |  | | 2019 | COLLECTED | 10/17/2019 | Ledges State Park located approximately 4.75 miles south of Boone, just south of the Oakwoods Shelter/Picnic area, T83N R26W NE1/4 NE1/4 and also NW1/4 NE1/4 Sec. 21, Boone County | 41.99262400, -93.87724800 | 330 | Edges of woods (adjacent to prairie) and also within 20-25 yards from woods in prairie. Noted in east, west, and south-facing edges of woods (no specimens noted on north-facing woodland edges) | Wild material | Woody shrub/vine stretching up to 12' in length. Alternate compound leaves usually consisting of five lealets. Leaf rachis and stipule margins glandular. Mature hypanthium and pedicel with stipitate glandular hairs. Mature hypanthium mostly bright to dark red in color with a few dark orange noted at time of harvest (17 October 2019). Leaves rather to somewhat glossy. This collection represents genetics from the northwestern edge of the species range, likely suggesting good cold-hardiness (USDA Zone 5a). | 2097769 | WLP 1685 |
| 10 | Ames 35228 | JDC/AS/2019/031/910 | Agastache scrophulariifolia (Willd.) Kuntze | Iowa, United States | NC7 |  | | 2019 | COLLECTED | 09/30/2019 | Ledges State Park located approximately 4.75 miles south of Boone, just east and south of the Carl Fritz Henning Shelter, T83N R26W SW1/4 SE1/4 NW1/4 Sec. 21, Boone County | 41.98786200, -93.88824500 | 276 | Moist, semi-shaded, deciduous woods. Recently disturbed. | Wild material | Plants measuring approximately 6-8' in height with 4-5" spikes rather white or extremely pale pink in color. Peak flowering occurred circa 26 August 2019 in nature. | 2097764 | Ames 35228 |
| 11 | Ames 33736 | JDC/BA/2016/028/743 | Betula alleghaniensis Britton | Iowa, United States | NC7 |  | | 2016 | COLLECTED | 09/02/2016 | Fallen Rock State Preserve approximately 1.9 miles southwest of Steamboat Rock, T88N R19W NW ¼ Sec. 32. | 42.39459000, -93.09423600 | 310 | High-quality, native, mesic deciduous woods on loose, loamy sandstone amongst very large boulders. North-facing with variable 10-50 degree slopes. Majority of specimens located at base of bluff or near water’s edge (Iowa River). | Wild material | The native range of Betula alleghaniensis is concentrated in the northeastern U.S. with disjunct populations in Iowa (Little, 1971). This accession represents a collection of Betula alleghaniensis germplasm from the extreme western edge of the species native range. The next closest occurrence is in Mitchell County, Iowa (60 miles). As of 2025, two trees at Fallen Rock State Preserve currently represent the state of Iowa's 1st and 2nd largest yellow birch specimens measuring 22.76" DBH; 70.83' height; 40' average crow spread (151.83 points) and 17.18" DBH; 65.33' height; 33.75' average crow spread (127.76 points), respectively (Rouw, M., personal communication, 13 May 2018). | 1942689 | Ames 33736 |
| 12 | Ames 32758 | Fallen Rock | Pinus strobus L. | Iowa, United States | NC7 |  | | 2014 | COLLECTED | 08/21/2014 | Fallen Rock State Preserve and adjacent private property just north of Eldora, T88N R19W SW ¼ Sec. 28 and N ¼ Sec. 32 and also T87N R19W N ½ Sec. 5. | 42.39548900, -93.09165200 | 311 | Cool moist woods. Slopes averaging 60 degrees. Northern and eastern aspects. Localized to bluffs above Iowa River. | Wild material | The native range of Pinus strobus is concentrated in the northeastern United States extending south into the northern edge of Georgia and west to central Minnesota. In Iowa, scattered native populations exist in a few northeastern counties as well as small disjunct populations in central and southeastern counties (Little, 1971). This accession includes genetics from legacy mother trees located in Fallen Rock State Preserve and nearby private property to the south. This area is near Pine Lake State Park, which in 2009 was devastated by a severe hailstorm of "supersized softballs" resulting in significant cambial damage to many species of trees. Damage to the majority of P. strobus specimens throughout the area was so severe that losses occurred immediately. Additional losses in subsequent years continued to occur likely due to additional stress by drought and destruction by bark beetles. The loss of approximately 75% of the P. strobus specimens at Pine Lake State Park was attributed to the 2009 hailstorm prompted efforts to preserve genetics of the white pines here. In 2014, specimens in parts of Pine Lake State Park and also one specimen at Fallen Rock State Preserve were sampled (Carstens, J.) as seeds from specimens ranging in size from 9.25 inches to 31.75 inches d.b.h. However, in 2021, it was determined (Place, A. pers. comm. 24 February 2021 and Carstens, J. & Sherwood, A., pers. observations 1 March 2021) that likely most if not all of the specimens in Pine Lake State Park are planted specimens of unknown origin because P. strobus specimens located at the far east end of Lower Lake (south side) were noted (Carstens and Sherwood, pers. observations 1 March 2021) growing with uncharacteristic associated taxa including Acer saccharinum and Gymnocladus dioicus. Two documents that support these observations includes: 1. Letter signed by Louis H. Pammel (March 25, 1925) to G.B. McDonald of the Iowa State University Forestry Department asking for 200 white pine to be planted at Pine Lake. 2. "Trees to Be Set Out by CCC Workers" in the Eldora Herald (May 9, 1935) referencing the CCC (Civilian Conservation Corps) to plant “1,000 white pines at Pine Lake State Park". Therefore, if obtained, increment borer measurements of specimens that are 94 years of age at present (2014) or older would suggest those as being true, native remnants. Documents that supports the position of remnant specimens at Fallen Rock State Preserve includes: 1. "Picturesque Drives About Eldora", in the Eldora Herald (December 8, 1910) by Clara Estabrook stating "Miss Stoddard of the Iowa Falls school says these (only 11 trees left) are the only native pines (those restricted to the sandstone cliffs) found in Hardin County" and suggested a movement to have them preserved. 2. Forest Flora of Hardin County (1899) by L. H. Pammel stating that the Iowa valley serves as the western limits for white pine and also refers to David D. Owen's geological report from 1839 indicating the occurrences of white pines on the summits of the hills along the Iowa River. In addition, a photo of white pines growing at Steamboat Rock taken by L. H. Pammel was featured on the front page of the 1919 report of the State Board of Conservation titled "Iowa Parks: Conservation of Iowa Historic, Scenic and Scientific Areas." Due to the significant presence of planted specimens of unknown origin in Pine Lake State Park and surrounding private properties, samples collected in 2014 as seeds may likely contain genetics (pollen donors) from specimens of unknown origin. This resulted in the resampling of specimens in 2021 (USFS - IAHA#'s) and 2023 as dormant scions that are from locations known to be untouched by the CCC and thus represent true, remnant specimens. To date (2021), there are only eight legacy specimens left in this population (four specimens at Fallen Rock State Preserve and four specimens at the nearby private property). The largest specimen of P. strobus (Ames 32758) is #IAHA-08 located at (42.395489, -93.091652) measuring (13 May 2018) 35.65" DBH with a height of 78.33' and a spread of 26' totalling 197 points (Rouw, M., personal communication, 8 February 2023). This germplasm represents the southwestern most population of native P. strobus in Iowa and one of the western most populations in North America. Germplasm likely is adapted to Midwestern climatic extremes and potentially very drought tolerant in comparison to other provenances from the eastern or northeastern U.S. | 1921270 | Ames 32758 |
| 13 | Ames 32093 | JDC/CC/2013/048/492 | Carpinus caroliniana Walter | Iowa, United States | NC7 | | | 2013 | COLLECTED | 10/25/2013 | Ledges State Park mostly from specimens adjacent to Peas Creek, NE ¼ of SE ¼ of Sec. 16, T83N R26W, Des Moines Quad. | 41.99910000, -93.87880000 | 328 | North and northwest-facing slopes. | Wild material | The native range of Carpinus caroliniana is concentrated in the eastern half of the U.S. with a very small disjunct population in central Iowa (Little, 1971). This accession represents a collection of C. caroliniana germplasm from the extreme western edge of the species native range, specifically the central Iowa disjunct population at Ledges State Park. The next closest occurrence is in eastern Iowa (100-150 miles). As of 2025, one specimen at Ledges State Park (McCoy Wildlife Area) currently represents the state of Iowa's largest C. caroliniana specimen measuring 6.68" DBH, 41.67’ in height, and 23.75’ average crown spread for a total of 68.608 points (Rouw, M., personal communication, 4 May 2022). This accession does include genotypes that do tend to sucker/form clonal colonies. Fall color (red/orange) tends to peak in the last week of October (central Iowa) with peak fruiting occurring simultaneously. Bark smooth, grey. | 1914037 | Ames 32093 |
| 14 | Ames 31515 | Duplicate of Ames 36172 | Cornus rugosa Lam. | Iowa, United States | | | Historic | 2012 | COLLECTED | 07/12/2012 | Ledges State Park, NE 1/4 of NW 1/4 of Section 21, T83N, R26W, Madrid NW Quad, near Boone, Boone County. | 41.99377000, -93.88624000 | 328 | Mixed hardwood forest. Edges of bluffs. North-facing slopes. | Wild material | | 1903742 | Ames 31515 |
| 15 | Ames 31256 | APS/GD/2008/005/005 | Gymnocladus dioicus (L.) K. Koch | Iowa, United States | NC7 |  | | 2012 | COLLECTED | 01/10/2012 | Ledges State Park, SW 1/4 of SW 1/4 of Section 21, T83N, R26W, Madrid NW Quad, Boone County | 41.98110000, -93.89010000 | 328 | Upland woods. 30% slope with a western aspect. | Wild material | Largest specimen noted was 22" d.b.h. | 1897494 | Ames 31256 |
| 16 | Ames 29614 | JDC/FN/2008/017/087 | Fraxinus nigra Marshall | Iowa, United States | NC7 |  | Not Available | 2008 | COLLECTED | 09/22/2008 | Along Davis Creek (starting at Peas Creek and heading north towards park boundary), Ledges State Park, W 1/2 of Section 16, Boone County. | 42.00222222, -93.88694444 | 289 | Mostly shaded; rock and limestone substrate. Species restricted to banks of creek. | Wild material | Approximately 70+ feet in height and approximately 3 feet DBH. | 1786593 | Ames 29614 |
| 17 | Ames 29506 | JDC-CA-2008-003 | Cornus alternifolia L. f. | Iowa, United States | NC7 |  | | 2008 | COLLECTED | 07/20/2008 | Tower Rock Park, N edge of SE 1/4 of SW 1/4 of Section 28, T88N, R19W, Steamboat Roak Quad, Steamboat Rock, Hardin County. | 42.40111111, -93.07166667 | 292 | Eroding (burnt orange) sandstone cliff, south side of Iowa River; mostly shaded; near base of slope with northern aspect; sand and organic matter above cliff; well drained. | Wild material | | 1782028 | Ames 29506 |
| 18 | Ames 28350 | 12 | Hypericum majus (A. Gray) Britton | Iowa, United States | NC7 |  | Not Available | 2006 | COLLECTED | 09/15/2006 | East-southeast across a deep gulley from a primitive campground parking lot, Ledges State Park, NE 1/4 of SW 1/4 of SW 1/4 of Section 15, T83N, R26W, Worth Township, Boone County. | 41.99638889, -93.87055556 | 329 | Hilltop savanna. Full sun. Dry (possibly loess) soil. | Wild material | Plants 1-6 inches tall. | 1712920 | Ames 28350 |