Taxon: Prunella vulgaris L.

Summary

Genus:

Prunella

Family:

Lamiaceae (alt. Labiatae)

Subfamily:

Nepetoideae

Tribe:

Mentheae

Subtribe:

Menthinae

Nomen number:

29826

Place of publication:

Sp. pl. 2:600. 1753

Protologue link:

https://biodiversitylibrary.org/page/358621

Typification:

View in Linnean Typification Project

Verified:

11/08/2007 ARS Systematic Botanists.

Accessions:

59 (55 active, 53 available) in National Plant Germplasm System. (Map it)

Other conspecific taxa

Prunella vulgaris L. subsp. asiatica (Nakai) H. Hara (4 active accession[s])
Prunella vulgaris L. subsp. lanceolata (W. P. C. Barton) Piper & Beattie (9 active accession[s])
Prunella vulgaris L. subsp. vulgaris (0 active accession[s])

Autonyms (not in current use), synonyms and invalid designations

Images (1 total. Click on image for more.)

Links

Seeds or fruits – Seed ID Workshop image, from Department of Horticulture and Crop Science, Ohio State University

Conservation Status

Reference(s)

Aldén, B., S. Ryman, & M. Hjertson. 2012. Svensk Kulturväxtdatabas, SKUD (Swedish Cultivated and Utility Plants Database; online resource)
APHIS. Agricultural Risk Management System United States Department of Agriculture.
Arechavaleta, M. et al., coords. 2009. Lista de especies silvestres de Canarias: hongos, plantas y animales terrestres
CIBA-GEIGY, Basel, Switzerland. 1968-1974. The CIBA-GEIGY Weed Tables 82.1.
Darbyshire, S. J. 2003. Inventory of Canadian Agricultural Weeds Agriculture and Agri-Food Canada. 209.
Davis, P. H., ed. 1965-1988. Flora of Turkey and the east Aegean islands.
Demilly, D. et al. 1996. Liste alphabétique des principales espèces de plantes cultivées et de mauvaises herbes. Noms latins et noms français, ed. 6. Note: GEVES-SNES, Beaucouzé, France
Duke, J. A. et al. 2002. CRC Handbook of medicinal herbs
Dyer, R. A. et al., eds. 1963-. Flora of southern Africa.
Food and Agriculture Organization of the United Nations (FAO). 2010. Ecocrop (on-line resource).
Gleason, H. A. & A. Cronquist. 1991. Manual of vascular plants of northeastern United States and adjacent Canada, ed. 2
Greuter, W. et al., eds. 1984-. Med-Checklist.
Hara, H. et al. 1978-1982. An enumeration of the flowering plants of Nepal.
Hickman, J. C., ed. 1993. The Jepson manual: higher plants of California
Iwatsuki, K. et al. 1993-. Flora of Japan.
Komarov, V. L. et al., eds. 1934-1964. Flora SSSR.
Krasnoborov, I. M., ed. 2000-. Flora of Siberia (English translation)
Lee, Y. N. 1997. Flora of Korea.
Magee, D. W. & H. E. Ahles. 1999. Flora of the Northeast. A manual of the vascular flora of New England and adjacent New York.
Martin, W. C. & C. R. Hutchins. 1980. A flora of New Mexico.
McGregor, R. L. et al. (The Great Plains Flora Association). 1986. Flora of the Great Plains.
McGuffin, M., J. T. Kartesz, A. Y. Leung, & A. O. Tucker. 2000. Herbs of commerce, ed. 2 American Herbal Products Association, Silver Spring, Maryland.
Mouterde, P. 1966-. Nouvelle flore du Liban et de la Syrie.
Nasir, E. & S. I. Ali, eds. 1970-. Flora of [West] Pakistan.
Personal Care Products Council. International Nomenclature Cosmetic Ingredient
Rechinger, K. H., ed. 1963-. Flora iranica.
Silva, L. et al. 2005. Listagem da fauna e flora terrestres dos Açores. Lista des plantas vasculares (Pteridophyta e Spermatophyta).
Turker, A. U. et al. 2010. Effect of phytohormones on micropropagation of Self-Heal (Prunella vulgaris L.). J. Herbs Spices Med. Plants 15:293-302. DOI: 10.1080/10496470903507809.
Tutin, T. G. et al., eds. 1964-1980. Flora europaea.
Vibrans, H., ed. Malezas de México (on-line resource).
Wu Zheng-yi & P. H. Raven et al., eds. 1994-. Flora of China (English edition).
2018. Mid-Atlantic invasive plant species (on-line resource)

Common names

Language	Name	note	seq	Citation
English	heal-all		1	McGuffin, M., J. T. Kartesz, A. Y. Leung, & A. O. Tucker. 2000. Herbs of commerce, ed. 2 American Herbal Products Association, Silver Spring, Maryland.
English	self-heal		1	McGregor, R. L. et al. (The Great Plains Flora Association). 1986. Flora of the Great Plains.
English (Canada)	heal-all	Agriculture & Agri-Food Canada official name	2	Darbyshire, S. J. 2003. Inventory of Canadian Agricultural Weeds Agriculture and Agri-Food Canada. 209.
French	brunelle vulgaire		2	Demilly, D. et al. 1996. Liste alphabétique des principales espèces de plantes cultivées et de mauvaises herbes. Noms latins et noms français, ed. 6. Note: GEVES-SNES, Beaucouzé, France
French (Canada)	brunelle commune	Agriculture & Agroalimentaire Canada nom officiel	2	Darbyshire, S. J. 2003. Inventory of Canadian Agricultural Weeds Agriculture and Agri-Food Canada. 209.
Swedish	brunört		2	Aldén, B., S. Ryman, & M. Hjertson. 2012. Svensk Kulturväxtdatabas, SKUD (Swedish Cultivated and Utility Plants Database; online resource)

Distribution

order_code	Status	Continent	Subcontinent	Country	State	Note	Reference
1	Native	Africa	Macaronesia	Portugal	Madeira Islands
1	Native	Africa	Macaronesia	Spain	Canarias		Arechavaleta, M. et al., coords. 2009. Lista de especies silvestres de Canarias: hongos, plantas y animales terrestres
1	Native	Africa	Northern Africa	Algeria			Greuter, W. et al., eds. 1984-. Med-Checklist.
1	Native	Africa	Northern Africa	Morocco			Greuter, W. et al., eds. 1984-. Med-Checklist.
1	Native	Africa	Northern Africa	Tunisia			Greuter, W. et al., eds. 1984-. Med-Checklist.
1	Native	Asia-Temperate	Caucasus	Armenia			Komarov, V. L. et al., eds. 1934-1964. Flora SSSR.
1	Native	Asia-Temperate	Caucasus	Azerbaijan			Komarov, V. L. et al., eds. 1934-1964. Flora SSSR.
1	Native	Asia-Temperate	Caucasus	Georgia			Komarov, V. L. et al., eds. 1934-1964. Flora SSSR.
1	Native	Asia-Temperate	Caucasus	Russian Federation	Dagestan		Komarov, V. L. et al., eds. 1934-1964. Flora SSSR.
1	Native	Asia-Temperate	Caucasus	Russian Federation-Ciscaucasia	Ciscaucasia		Komarov, V. L. et al., eds. 1934-1964. Flora SSSR.
1	Native	Asia-Temperate	China	China	Fujian Sheng		Wu Zheng-yi & P. H. Raven et al., eds. 1994-. Flora of China (English edition).
1	Native	Asia-Temperate	China	China	Gansu Sheng		Wu Zheng-yi & P. H. Raven et al., eds. 1994-. Flora of China (English edition).
1	Native	Asia-Temperate	China	China	Guangdong Sheng		Wu Zheng-yi & P. H. Raven et al., eds. 1994-. Flora of China (English edition).
1	Native	Asia-Temperate	China	China	Guangxi Zhuangzu Zizhiqu		Wu Zheng-yi & P. H. Raven et al., eds. 1994-. Flora of China (English edition).
1	Native	Asia-Temperate	China	China	Guizhou Sheng		Wu Zheng-yi & P. H. Raven et al., eds. 1994-. Flora of China (English edition).
1	Native	Asia-Temperate	China	China	Henan Sheng		Wu Zheng-yi & P. H. Raven et al., eds. 1994-. Flora of China (English edition).
1	Native	Asia-Temperate	China	China	Hubei Sheng		Wu Zheng-yi & P. H. Raven et al., eds. 1994-. Flora of China (English edition).
1	Native	Asia-Temperate	China	China	Hunan Sheng		Wu Zheng-yi & P. H. Raven et al., eds. 1994-. Flora of China (English edition).
1	Native	Asia-Temperate	China	China	Jiangxi Sheng		Wu Zheng-yi & P. H. Raven et al., eds. 1994-. Flora of China (English edition).
1	Native	Asia-Temperate	China	China	Shaanxi Sheng		Wu Zheng-yi & P. H. Raven et al., eds. 1994-. Flora of China (English edition).
1	Native	Asia-Temperate	China	China	Sichuan Sheng		Wu Zheng-yi & P. H. Raven et al., eds. 1994-. Flora of China (English edition).
1	Native	Asia-Temperate	China	China	Xinjiang Uygur Zizhiqu		Wu Zheng-yi & P. H. Raven et al., eds. 1994-. Flora of China (English edition).
1	Native	Asia-Temperate	China	China	Xizang Zizhiqu		Wu Zheng-yi & P. H. Raven et al., eds. 1994-. Flora of China (English edition).
1	Native	Asia-Temperate	China	China	Yunnan Sheng		Wu Zheng-yi & P. H. Raven et al., eds. 1994-. Flora of China (English edition).
1	Native	Asia-Temperate	China	China	Zhejiang Sheng		Wu Zheng-yi & P. H. Raven et al., eds. 1994-. Flora of China (English edition).
1	Native	Asia-Temperate	Eastern Asia	Japan	Hokkaidô		Iwatsuki, K. et al. 1993-. Flora of Japan.
1	Native	Asia-Temperate	Eastern Asia	Japan	Honshu		Iwatsuki, K. et al. 1993-. Flora of Japan.
1	Native	Asia-Temperate	Eastern Asia	Japan	Kyushu		Iwatsuki, K. et al. 1993-. Flora of Japan.
1	Native	Asia-Temperate	Eastern Asia	Japan	Shikoku		Iwatsuki, K. et al. 1993-. Flora of Japan.
1	Native	Asia-Temperate	Eastern Asia	Korea			Lee, Y. N. 1997. Flora of Korea.
1	Native	Asia-Temperate	Eastern Asia	Taiwan
1	Native	Asia-Temperate	Middle Asia	Kazakhstan			Komarov, V. L. et al., eds. 1934-1964. Flora SSSR.
1	Native	Asia-Temperate	Middle Asia	Kyrgyzstan			Komarov, V. L. et al., eds. 1934-1964. Flora SSSR.
1	Native	Asia-Temperate	Middle Asia	Tajikistan			Komarov, V. L. et al., eds. 1934-1964. Flora SSSR.
1	Native	Asia-Temperate	Middle Asia	Turkmenistan			Komarov, V. L. et al., eds. 1934-1964. Flora SSSR.
1	Native	Asia-Temperate	Middle Asia	Uzbekistan			Komarov, V. L. et al., eds. 1934-1964. Flora SSSR.
1	Native	Asia-Temperate	Mongolia	Mongolia
1	Native	Asia-Temperate	Russian Far East	Russian Federation-Far East	Far East		Komarov, V. L. et al., eds. 1934-1964. Flora SSSR.
1	Native	Asia-Temperate	Siberia	Russian Federation	Altay		Krasnoborov, I. M., ed. 2000-. Flora of Siberia (English translation)
1	Native	Asia-Temperate	Siberia	Russian Federation	Buryatia		Krasnoborov, I. M., ed. 2000-. Flora of Siberia (English translation)
1	Native	Asia-Temperate	Siberia	Russian Federation	Gorno-Altay		Krasnoborov, I. M., ed. 2000-. Flora of Siberia (English translation)
1	Native	Asia-Temperate	Siberia	Russian Federation	Irkutsk		Krasnoborov, I. M., ed. 2000-. Flora of Siberia (English translation)
1	Native	Asia-Temperate	Siberia	Russian Federation	Kemerovskaja oblast'		Krasnoborov, I. M., ed. 2000-. Flora of Siberia (English translation)
1	Native	Asia-Temperate	Siberia	Russian Federation	Krasnoyarsk		Krasnoborov, I. M., ed. 2000-. Flora of Siberia (English translation)
1	Native	Asia-Temperate	Siberia	Russian Federation	Kurganskaja oblast'		Krasnoborov, I. M., ed. 2000-. Flora of Siberia (English translation)
1	Native	Asia-Temperate	Siberia	Russian Federation	Novosibirsk		Krasnoborov, I. M., ed. 2000-. Flora of Siberia (English translation)
1	Native	Asia-Temperate	Siberia	Russian Federation	Omsk		Krasnoborov, I. M., ed. 2000-. Flora of Siberia (English translation)
1	Native	Asia-Temperate	Siberia	Russian Federation	Tomsk		Krasnoborov, I. M., ed. 2000-. Flora of Siberia (English translation)
1	Native	Asia-Temperate	Siberia	Russian Federation	Tyumen		Krasnoborov, I. M., ed. 2000-. Flora of Siberia (English translation)
1	Native	Asia-Temperate	Siberia	Russian Federation	Tyva, Respublika		Krasnoborov, I. M., ed. 2000-. Flora of Siberia (English translation)
1	Native	Asia-Temperate	Siberia	Russian Federation	Yakutia-Sakha		Krasnoborov, I. M., ed. 2000-. Flora of Siberia (English translation)
1	Native	Asia-Temperate	Western Asia	Afghanistan			Rechinger, K. H., ed. 1963-. Flora iranica.
1	Native	Asia-Temperate	Western Asia	Cyprus
1	Native	Asia-Temperate	Western Asia	Iran			Rechinger, K. H., ed. 1963-. Flora iranica.
1	Native	Asia-Temperate	Western Asia	Iraq			Rechinger, K. H., ed. 1963-. Flora iranica.
1	Native	Asia-Temperate	Western Asia	Lebanon			Mouterde, P. 1966-. Nouvelle flore du Liban et de la Syrie.
1	Native	Asia-Temperate	Western Asia	Syria			Mouterde, P. 1966-. Nouvelle flore du Liban et de la Syrie.
1	Native	Asia-Temperate	Western Asia	Turkey			Davis, P. H., ed. 1965-1988. Flora of Turkey and the east Aegean islands.
1	Native	Asia-Tropical	Indian Subcontinent	Bhutan
1	Native	Asia-Tropical	Indian Subcontinent	India
1	Native	Asia-Tropical	Indian Subcontinent	Nepal			Hara, H. et al. 1978-1982. An enumeration of the flowering plants of Nepal.
1	Native	Asia-Tropical	Indian Subcontinent	Pakistan			Nasir, E. & S. I. Ali, eds. 1970-. Flora of [West] Pakistan.
1	Native	Europe	Eastern Europe	Belarus
1	Native	Europe	Eastern Europe	Estonia
1	Native	Europe	Eastern Europe	Latvia
1	Native	Europe	Eastern Europe	Lithuania
1	Native	Europe	Eastern Europe	Moldova
1	Native	Europe	Eastern Europe	Russian Federation-European part	European part		Komarov, V. L. et al., eds. 1934-1964. Flora SSSR.
1	Native	Europe	Eastern Europe	Ukraine		incl. Krym
1	Native	Europe	Middle Europe	Austria			Tutin, T. G. et al., eds. 1964-1980. Flora europaea.
1	Native	Europe	Middle Europe	Belgium			Tutin, T. G. et al., eds. 1964-1980. Flora europaea.
1	Native	Europe	Middle Europe	Czechoslovakia			Tutin, T. G. et al., eds. 1964-1980. Flora europaea.
1	Native	Europe	Middle Europe	Germany			Tutin, T. G. et al., eds. 1964-1980. Flora europaea.
1	Native	Europe	Middle Europe	Hungary			Tutin, T. G. et al., eds. 1964-1980. Flora europaea.
1	Native	Europe	Middle Europe	Netherlands			Tutin, T. G. et al., eds. 1964-1980. Flora europaea.
1	Native	Europe	Middle Europe	Poland			Tutin, T. G. et al., eds. 1964-1980. Flora europaea.
1	Native	Europe	Middle Europe	Switzerland			Tutin, T. G. et al., eds. 1964-1980. Flora europaea.
1	Native	Europe	Northern Europe	Denmark			Tutin, T. G. et al., eds. 1964-1980. Flora europaea.
1	Native	Europe	Northern Europe	Finland			Tutin, T. G. et al., eds. 1964-1980. Flora europaea.
1	Native	Europe	Northern Europe	Ireland			Tutin, T. G. et al., eds. 1964-1980. Flora europaea.
1	Native	Europe	Northern Europe	Norway			Tutin, T. G. et al., eds. 1964-1980. Flora europaea.
1	Native	Europe	Northern Europe	Sweden			Tutin, T. G. et al., eds. 1964-1980. Flora europaea.
1	Native	Europe	Northern Europe	United Kingdom			Tutin, T. G. et al., eds. 1964-1980. Flora europaea.
1	Native	Europe	Southeastern Europe	Albania			Greuter, W. et al., eds. 1984-. Med-Checklist.
1	Native	Europe	Southeastern Europe	Bulgaria			Greuter, W. et al., eds. 1984-. Med-Checklist.
1	Native	Europe	Southeastern Europe	Former Yugoslavia			Greuter, W. et al., eds. 1984-. Med-Checklist.
1	Native	Europe	Southeastern Europe	Greece		incl. Crete	Greuter, W. et al., eds. 1984-. Med-Checklist.
1	Native	Europe	Southeastern Europe	Italy		incl. Sardinia, Sicily	Greuter, W. et al., eds. 1984-. Med-Checklist.
1	Native	Europe	Southeastern Europe	Romania			Tutin, T. G. et al., eds. 1964-1980. Flora europaea.
1	Native	Europe	Southwestern Europe	France		incl. Corsica	Tutin, T. G. et al., eds. 1964-1980. Flora europaea.
1	Native	Europe	Southwestern Europe	Portugal			Tutin, T. G. et al., eds. 1964-1980. Flora europaea.
1	Native	Europe	Southwestern Europe	Spain		incl. Baleares	Tutin, T. G. et al., eds. 1964-1980. Flora europaea.
1	Native	Northern America	Eastern Canada	Canada	New Brunswick
1	Native	Northern America	Eastern Canada	Canada	Newfoundland and Labrador
1	Native	Northern America	Eastern Canada	Canada	Nova Scotia
1	Native	Northern America	Eastern Canada	Canada	Ontario
1	Native	Northern America	Eastern Canada	Canada	Prince Edward Island
1	Native	Northern America	Eastern Canada	Canada	Québec
1	Native	Northern America	North-Central U.S.A.	United States	Illinois		Gleason, H. A. & A. Cronquist. 1991. Manual of vascular plants of northeastern United States and adjacent Canada, ed. 2
1	Native	Northern America	North-Central U.S.A.	United States	Iowa		Gleason, H. A. & A. Cronquist. 1991. Manual of vascular plants of northeastern United States and adjacent Canada, ed. 2
1	Native	Northern America	North-Central U.S.A.	United States	Kansas		McGregor, R. L. et al. (The Great Plains Flora Association). 1986. Flora of the Great Plains.
1	Native	Northern America	North-Central U.S.A.	United States	Minnesota		Gleason, H. A. & A. Cronquist. 1991. Manual of vascular plants of northeastern United States and adjacent Canada, ed. 2
1	Native	Northern America	North-Central U.S.A.	United States	Missouri		Gleason, H. A. & A. Cronquist. 1991. Manual of vascular plants of northeastern United States and adjacent Canada, ed. 2
1	Native	Northern America	North-Central U.S.A.	United States	Nebraska		McGregor, R. L. et al. (The Great Plains Flora Association). 1986. Flora of the Great Plains.
1	Native	Northern America	North-Central U.S.A.	United States	North Dakota		McGregor, R. L. et al. (The Great Plains Flora Association). 1986. Flora of the Great Plains.
1	Native	Northern America	North-Central U.S.A.	United States	Oklahoma		McGregor, R. L. et al. (The Great Plains Flora Association). 1986. Flora of the Great Plains.
1	Native	Northern America	North-Central U.S.A.	United States	South Dakota		McGregor, R. L. et al. (The Great Plains Flora Association). 1986. Flora of the Great Plains.
1	Native	Northern America	North-Central U.S.A.	United States	Wisconsin		Gleason, H. A. & A. Cronquist. 1991. Manual of vascular plants of northeastern United States and adjacent Canada, ed. 2
1	Native	Northern America	Northeastern U.S.A.	United States	Connecticut		Magee, D. W. & H. E. Ahles. 1999. Flora of the Northeast. A manual of the vascular flora of New England and adjacent New York.
1	Native	Northern America	Northeastern U.S.A.	United States	Indiana		Gleason, H. A. & A. Cronquist. 1991. Manual of vascular plants of northeastern United States and adjacent Canada, ed. 2
1	Native	Northern America	Northeastern U.S.A.	United States	Maine		Magee, D. W. & H. E. Ahles. 1999. Flora of the Northeast. A manual of the vascular flora of New England and adjacent New York.
1	Native	Northern America	Northeastern U.S.A.	United States	Massachusetts		Magee, D. W. & H. E. Ahles. 1999. Flora of the Northeast. A manual of the vascular flora of New England and adjacent New York.
1	Native	Northern America	Northeastern U.S.A.	United States	Michigan		Gleason, H. A. & A. Cronquist. 1991. Manual of vascular plants of northeastern United States and adjacent Canada, ed. 2
1	Native	Northern America	Northeastern U.S.A.	United States	New Hampshire		Magee, D. W. & H. E. Ahles. 1999. Flora of the Northeast. A manual of the vascular flora of New England and adjacent New York.
1	Native	Northern America	Northeastern U.S.A.	United States	New Jersey		Gleason, H. A. & A. Cronquist. 1991. Manual of vascular plants of northeastern United States and adjacent Canada, ed. 2
1	Native	Northern America	Northeastern U.S.A.	United States	New York		Gleason, H. A. & A. Cronquist. 1991. Manual of vascular plants of northeastern United States and adjacent Canada, ed. 2
1	Native	Northern America	Northeastern U.S.A.	United States	Ohio		Gleason, H. A. & A. Cronquist. 1991. Manual of vascular plants of northeastern United States and adjacent Canada, ed. 2
1	Native	Northern America	Northeastern U.S.A.	United States	Pennsylvania		Gleason, H. A. & A. Cronquist. 1991. Manual of vascular plants of northeastern United States and adjacent Canada, ed. 2
1	Native	Northern America	Northeastern U.S.A.	United States	Rhode Island		Magee, D. W. & H. E. Ahles. 1999. Flora of the Northeast. A manual of the vascular flora of New England and adjacent New York.
1	Native	Northern America	Northeastern U.S.A.	United States	Vermont		Magee, D. W. & H. E. Ahles. 1999. Flora of the Northeast. A manual of the vascular flora of New England and adjacent New York.
1	Native	Northern America	Northeastern U.S.A.	United States	West Virginia		Gleason, H. A. & A. Cronquist. 1991. Manual of vascular plants of northeastern United States and adjacent Canada, ed. 2
1	Native	Northern America	Northwestern U.S.A.	United States	Colorado		McGregor, R. L. et al. (The Great Plains Flora Association). 1986. Flora of the Great Plains.
1	Native	Northern America	Northwestern U.S.A.	United States	Idaho		McGregor, R. L. et al. (The Great Plains Flora Association). 1986. Flora of the Great Plains.
1	Native	Northern America	Northwestern U.S.A.	United States	Montana		McGregor, R. L. et al. (The Great Plains Flora Association). 1986. Flora of the Great Plains.
1	Native	Northern America	Northwestern U.S.A.	United States	Oregon		Hickman, J. C., ed. 1993. The Jepson manual: higher plants of California
1	Native	Northern America	Northwestern U.S.A.	United States	Washington		Hickman, J. C., ed. 1993. The Jepson manual: higher plants of California
1	Native	Northern America	Northwestern U.S.A.	United States	Wyoming		McGregor, R. L. et al. (The Great Plains Flora Association). 1986. Flora of the Great Plains.
1	Native	Northern America	South-Central U.S.A.	United States	New Mexico		Martin, W. C. & C. R. Hutchins. 1980. A flora of New Mexico.
1	Native	Northern America	South-Central U.S.A.	United States	Texas		Gleason, H. A. & A. Cronquist. 1991. Manual of vascular plants of northeastern United States and adjacent Canada, ed. 2
1	Native	Northern America	Southeastern U.S.A.	United States	Alabama		Gleason, H. A. & A. Cronquist. 1991. Manual of vascular plants of northeastern United States and adjacent Canada, ed. 2
1	Native	Northern America	Southeastern U.S.A.	United States	Arkansas		Gleason, H. A. & A. Cronquist. 1991. Manual of vascular plants of northeastern United States and adjacent Canada, ed. 2
1	Native	Northern America	Southeastern U.S.A.	United States	Delaware		Gleason, H. A. & A. Cronquist. 1991. Manual of vascular plants of northeastern United States and adjacent Canada, ed. 2
1	Native	Northern America	Southeastern U.S.A.	United States	Florida		Gleason, H. A. & A. Cronquist. 1991. Manual of vascular plants of northeastern United States and adjacent Canada, ed. 2
1	Native	Northern America	Southeastern U.S.A.	United States	Georgia		Gleason, H. A. & A. Cronquist. 1991. Manual of vascular plants of northeastern United States and adjacent Canada, ed. 2
1	Native	Northern America	Southeastern U.S.A.	United States	Kentucky		Gleason, H. A. & A. Cronquist. 1991. Manual of vascular plants of northeastern United States and adjacent Canada, ed. 2
1	Native	Northern America	Southeastern U.S.A.	United States	Louisiana		Gleason, H. A. & A. Cronquist. 1991. Manual of vascular plants of northeastern United States and adjacent Canada, ed. 2
1	Native	Northern America	Southeastern U.S.A.	United States	Maryland		Gleason, H. A. & A. Cronquist. 1991. Manual of vascular plants of northeastern United States and adjacent Canada, ed. 2
1	Native	Northern America	Southeastern U.S.A.	United States	Mississippi		Gleason, H. A. & A. Cronquist. 1991. Manual of vascular plants of northeastern United States and adjacent Canada, ed. 2
1	Native	Northern America	Southeastern U.S.A.	United States	North Carolina		Gleason, H. A. & A. Cronquist. 1991. Manual of vascular plants of northeastern United States and adjacent Canada, ed. 2
1	Native	Northern America	Southeastern U.S.A.	United States	South Carolina		Gleason, H. A. & A. Cronquist. 1991. Manual of vascular plants of northeastern United States and adjacent Canada, ed. 2
1	Native	Northern America	Southeastern U.S.A.	United States	Tennessee		Gleason, H. A. & A. Cronquist. 1991. Manual of vascular plants of northeastern United States and adjacent Canada, ed. 2
1	Native	Northern America	Southeastern U.S.A.	United States	Virginia		Gleason, H. A. & A. Cronquist. 1991. Manual of vascular plants of northeastern United States and adjacent Canada, ed. 2
1	Native	Northern America	Southwestern U.S.A.	United States	Arizona
1	Native	Northern America	Southwestern U.S.A.	United States	California		Hickman, J. C., ed. 1993. The Jepson manual: higher plants of California
1	Native	Northern America	Southwestern U.S.A.	United States	Nevada		Hickman, J. C., ed. 1993. The Jepson manual: higher plants of California
1	Native	Northern America	Southwestern U.S.A.	United States	Utah		Hickman, J. C., ed. 1993. The Jepson manual: higher plants of California
1	Native	Northern America	Subarctic America	United States	Alaska
1	Native	Northern America	Western Canada	Canada	Alberta
1	Native	Northern America	Western Canada	Canada	British Columbia
1	Native	Northern America	Western Canada	Canada	Manitoba
4	Naturalized					widely natzd.
5	Uncertain	Africa	Macaronesia	Portugal	Azores		Silva, L. et al. 2005. Listagem da fauna e flora terrestres dos Açores. Lista des plantas vasculares (Pteridophyta e Spermatophyta).
6	Other					exact native range obscure

Native
Africa

MACARONESIA: Spain [Canarias], Portugal [Madeira Islands]
NORTHERN AFRICA: Algeria, Morocco, Tunisia

Asia-Temperate

WESTERN ASIA: Afghanistan, Cyprus, Iran, Iraq, Lebanon, Syria, Turkey
CAUCASUS: Russian Federation-Ciscaucasia [Ciscaucasia], Armenia, Azerbaijan, Georgia, Russian Federation [Dagestan]
SIBERIA: Russian Federation [Altay, Buryatia, Gorno-Altay, Irkutsk, Kemerovskaja oblast', Krasnoyarsk, Kurganskaja oblast', Novosibirsk, Omsk, Tomsk, Tyumen, Tyva, Respublika, Yakutia-Sakha]
MIDDLE ASIA: Kazakhstan, Kyrgyzstan, Tajikistan, Turkmenistan, Uzbekistan
MONGOLIA: Mongolia
RUSSIAN FAR EAST: Russian Federation-Far East [Far East]
CHINA: China [Fujian Sheng, Gansu Sheng, Guangdong Sheng, Guangxi Zhuangzu Zizhiqu, Guizhou Sheng, Henan Sheng, Hubei Sheng, Hunan Sheng, Jiangxi Sheng, Shaanxi Sheng, Sichuan Sheng, Xinjiang Uygur Zizhiqu, Xizang Zizhiqu, Yunnan Sheng, Zhejiang Sheng]
EASTERN ASIA: Korea, Japan [Hokkaidô, Honshu, Kyushu, Shikoku], Taiwan

Asia-Tropical

INDIAN SUBCONTINENT: Bhutan, India, Nepal, Pakistan

Europe

NORTHERN EUROPE: Denmark, Finland, United Kingdom, Ireland, Norway, Sweden
MIDDLE EUROPE: Czechoslovakia, Austria, Belgium, Switzerland, Germany, Hungary, Netherlands, Poland
EASTERN EUROPE: Russian Federation-European part [European part], Belarus, Estonia, Lithuania, Latvia, Moldova, Ukraine (incl. Krym)
SOUTHEASTERN EUROPE: Former Yugoslavia, Albania, Bulgaria, Greece (incl. Crete), Italy (incl. Sardinia, Sicily), Romania
SOUTHWESTERN EUROPE: Spain (incl. Baleares), France (incl. Corsica), Portugal

Northern America

SUBARCTIC AMERICA: United States [Alaska]
EASTERN CANADA: Canada [Québec, Nova Scotia, Ontario, Prince Edward Island, New Brunswick, Newfoundland and Labrador]
WESTERN CANADA: Canada [Alberta, Manitoba, British Columbia]
NORTHEASTERN U.S.A.: United States [Connecticut, Indiana, Maine, Massachusetts, Michigan, New Hampshire, New Jersey, New York, Ohio, Pennsylvania, Rhode Island, Vermont, West Virginia]
NORTH-CENTRAL U.S.A.: United States [Illinois, Iowa, Kansas, Minnesota, Missouri, Nebraska, North Dakota, Oklahoma, South Dakota, Wisconsin]
NORTHWESTERN U.S.A.: United States [Colorado, Idaho, Montana, Oregon, Washington, Wyoming]
SOUTHEASTERN U.S.A.: United States [Alabama, Arkansas, Delaware, Florida, Georgia, Kentucky, Louisiana, Maryland, Mississippi, North Carolina, South Carolina, Tennessee, Virginia]
SOUTH-CENTRAL U.S.A.: United States [New Mexico, Texas]
SOUTHWESTERN U.S.A.: United States [Arizona, California, Nevada, Utah]

Naturalized (widely natzd.)
Uncertain
Africa

MACARONESIA: Portugal [Azores]

Other (exact native range obscure)

Reference(s)

Arechavaleta, M. et al., coords. 2009. Lista de especies silvestres de Canarias: hongos, plantas y animales terrestres
Davis, P. H., ed. 1965-1988. Flora of Turkey and the east Aegean islands.
Gleason, H. A. & A. Cronquist. 1991. Manual of vascular plants of northeastern United States and adjacent Canada, ed. 2
Greuter, W. et al., eds. 1984-. Med-Checklist.
Hara, H. et al. 1978-1982. An enumeration of the flowering plants of Nepal.
Hickman, J. C., ed. 1993. The Jepson manual: higher plants of California
Iwatsuki, K. et al. 1993-. Flora of Japan.
Komarov, V. L. et al., eds. 1934-1964. Flora SSSR.
Krasnoborov, I. M., ed. 2000-. Flora of Siberia (English translation)
Lee, Y. N. 1997. Flora of Korea.
Magee, D. W. & H. E. Ahles. 1999. Flora of the Northeast. A manual of the vascular flora of New England and adjacent New York.
Martin, W. C. & C. R. Hutchins. 1980. A flora of New Mexico.
McGregor, R. L. et al. (The Great Plains Flora Association). 1986. Flora of the Great Plains.
Mouterde, P. 1966-. Nouvelle flore du Liban et de la Syrie.
Nasir, E. & S. I. Ali, eds. 1970-. Flora of [West] Pakistan.
Rechinger, K. H., ed. 1963-. Flora iranica.
Silva, L. et al. 2005. Listagem da fauna e flora terrestres dos Açores. Lista des plantas vasculares (Pteridophyta e Spermatophyta).
Tutin, T. G. et al., eds. 1964-1980. Flora europaea.
Wu Zheng-yi & P. H. Raven et al., eds. 1994-. Flora of China (English edition).

Economic Uses

Usage	Type	Note	Reference
Medicines	folklore		Duke, J. A. et al. 2002. CRC Handbook of medicinal herbs
Medicines	folklore		McGuffin, M., J. T. Kartesz, A. Y. Leung, & A. O. Tucker. 2000. Herbs of commerce, ed. 2 American Herbal Products Association, Silver Spring, Maryland.
Weed	seed contaminant	potential	CIBA-GEIGY, Basel, Switzerland. 1968-1974. The CIBA-GEIGY Weed Tables 82.1.

Cite as: USDA, Agricultural Research Service, National Plant Germplasm System. 2026. Germplasm Resources Information Network (GRIN Taxonomy). National Germplasm Resources Laboratory, Beltsville, Maryland.
URL: https://npgstest.ars-grin.gov/gringlobal/taxon/taxonomydetail?id=29826. Accessed 12 May 2026.