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ACCESSIONPLANT NAMETAXONOMYORIGINGENEBANKIMAGEAVAILABILITYRECEIVEDSOURCE TYPESOURCE DATECOLLECTION SITECOORDINATESELEVATIONHABITATIMPROVEMENT LEVELNARRATIVE
0GSHO 3656eog1.eHordeum vulgare L. subsp. vulgare Washington, United StatesNSGCKERNELSNot Available2013DEVELOPEDGenetic materialIncreased size of the outer glumes. Fine smooth awn.1913964GSHO 3656
1GSHO 3657eog1.fHordeum vulgare L. subsp. vulgare Washington, United StatesNSGCKERNELSNot Available2013DEVELOPEDGenetic materialIncreased size of the outer glumes.1913965GSHO 3657
2GSHO 3658eog1.gHordeum vulgare L. subsp. vulgare Washington, United StatesNSGCKERNELSNot Available2013DEVELOPEDGenetic materialIncreased size of the outer glumes.1913966GSHO 3658
3GSHO 3659eog1.hHordeum vulgare L. subsp. vulgare Washington, United StatesNSGCKERNELSNot Available2013DEVELOPEDGenetic materialIncreased size of the outer glumes.1913967GSHO 3659
4GSHO 3660eog1.iHordeum vulgare L. subsp. vulgare Washington, United StatesNSGCKERNELS2013DEVELOPEDGenetic materialIncreased size of the outer glumes. This mutant the outer glume is short, spike near normal.1913968GSHO 3660
5GSHO 3661eog1.jHordeum vulgare L. subsp. vulgare Washington, United StatesNSGCKERNELS2013DEVELOPEDGenetic materialIncreased size of the outer glumes. This mutant outer glume is short, spike near normal.1913969GSHO 3661
6GSHO 3662eog1.kHordeum vulgare L. subsp. vulgare Washington, United StatesNSGCKERNELSNot Available2013DEVELOPEDGenetic materialIncreased size of the outer glumes.1913970GSHO 3662
7GSHO 3663eog1.lHordeum vulgare L. subsp. vulgare Washington, United StatesNSGCKERNELSNot Available2013DEVELOPEDGenetic materialIncreased size of the outer glumes.1913971GSHO 3663
8GSHO 3664eog1.mHordeum vulgare L. subsp. vulgare Washington, United StatesNSGCKERNELSNot Available2013DEVELOPEDGenetic materialIncreased size of the outer glumes.1913972GSHO 3664
9GSHO 3665eog1.nHordeum vulgare L. subsp. vulgare Washington, United StatesNSGCKERNELSNot Available2013DEVELOPEDGenetic materialIncreased size of the outer glumes.1913973GSHO 3665
10GSHO 3666eog1.oHordeum vulgare L. subsp. vulgare Washington, United StatesNSGCKERNELSNot Available2013DEVELOPEDGenetic materialIncreased size of the outer glumes.1913974GSHO 3666
11GSHO 3667eog1.pHordeum vulgare L. subsp. vulgare Washington, United StatesNSGCKERNELSNot Available2013DEVELOPEDGenetic materialIncreased size of the outer glumes.1913975GSHO 3667
12GSHO 3668eog1.qHordeum vulgare L. subsp. vulgare Washington, United StatesNSGCKERNELSNot Available2013DEVELOPEDGenetic materialIncreased size of the outer glumes.1913976GSHO 3668
13GSHO 3669eog1.rHordeum vulgare L. subsp. vulgare Washington, United StatesNSGCKERNELSNot Available2013DEVELOPEDGenetic materialIncreased size of the outer glumes.1913977GSHO 3669
14GSHO 3670eog1.sHordeum vulgare L. subsp. vulgare Washington, United StatesNSGCKERNELSNot Available2013DEVELOPEDGenetic materialIncreased size of the outer glumes.1913978GSHO 3670
15GSHO 1231mtt4.eHordeum vulgare L. subsp. vulgare JapanNSGCKERNELS1996DEVELOPEDGenetic materialWhen the plants are grown under cool conditions, white-yellowish blotches develop as horizontal bands in seedling leaves, 2 to 4 per leaf blade. The bands fade as the plant grows or under warmer environmental conditions. At later stages of development, the plants have a slightly yellow-green color. Inheritance: Monofactorial recessive. Located in chromosome 2HL, about 13.7 cM distal from the vrs1 (six-rowed spike 1) locus. Previous nomenclature and gene symbolization: Mottled leaf-e= mt,,e. Mottled leaf=mt. Mutational events: mtt4.e in Victorie (OUL076). Mutant used for description and seed stocks: mtt4.e in Victorie (GSHO 1231); mtt4.e in Bowman (PI 483237)*6 (GSHO 1914).1514087GSHO 1231
16GSHO 1233btr1.aHordeum vulgare L. subsp. vulgare North Dakota, United StatesNSGCKERNELS1996DONATED04/1996Genetic materialJoints at the rachis nodes are tough or non-brittle, which prevents segmentation or disarticulation of the rachis prior to maturity. Plants homozygous for the btr1.a allele do not shatter prior to harvest in contrast to wild barley (Hordeum vulgare subsp spontaneum). All cultivars with the btr1.a allele have a dominant allele at the Btr2 locus. When crossed to cultivars with btr2.b allele at the Btr2 locus (genotype Btr1Btr1btr2btr2), F1 plants have a brittle rachis. The segregation ratio in the F2 generation is 1 brittle: 1 tough rachis. The majority of the cultivars distributed in Occidental region of the Old World have the btr1.a allele. Inheritance: Monofactorial recessive in crosses to Hordeum vulgare subsp. spontaneum. Located in chromosome 3HS, very close or semi-allelic to the Btr2 (brittle rachis 2) locus; the Btr1Btr2 complex is about 15.4 cM distal from the alm (albino lemma) locus. Previous nomenclature and gene symbolization: Non-brittle rachis=r. Non-brittle rachis=b. Non-brittle rachis 1=bt1. Mutational events: btr1.a in an unknown race of Hordeum vulgare subsp spontaneum; the line A222 (CIho 11555) is recommended as the source line for the btr1.a gene. Mutant used for description and seed stocks: btr1.a in A 222 (GSHO 1233); Btr1Btr2 from Wolfe's Multiple Dominant Marker Stock in Bowman (PI 4833237)*10 (GSHO 1937).1513891GSHO 1233
17GSHO 1576lax.a1Hordeum vulgare L. subsp. vulgare North Dakota, United StatesNSGCKERNELS1996DONATED04/1996Genetic materialBoth the upper and lower surface of the leaf blade have widely scattered, short (1 to 2 mm) hairs. The hairs tend to be aligned along the smaller leaf veins. The hairs are more difficult to observe on older leaves or plants grown in the field. Some accessions of Hordeum vulgare subsp spontaneum have numerous short hairs on both surfaces of the leaf. Inheritance: Monofactorial dominant. Located in chromosome 3HL. Previous nomenclature and gene symbolization: None. Mutational events: Pub1.a in Wolfe's Multiple Dominant Marker Stock (GSHO 1580); Pub1.a in Steptoe. Mutant used for description and seed stocks: Pub1.a in Steptoe (GSHO 1576); Pub1.a from Wolfe's Multiple Dominant Marker Stock in Bowman (PI 483237)*6 (GSHO 1976).1000525GSHO 1576
18GSHO 1588Rph10.oHordeum vulgare L. subsp. vulgare North Dakota, United StatesNSGCKERNELS1996DONATED04/1996Genetic materialThe seedling reaction varies from a necrotic fleck to a moderate sized uredinium with heterozygotes having an intermediate reaction. Culture Aust 220 from Australia was required to detect a resistant reaction for allelism tests and backcrosses. Inheritance: Monofactorial dominant. Located in chromosome 3HL, about 15.0 cM from the Est2 (esterase 2) locus. Previous nomenclature and gene symbolization: None. Mutational events: Rph10.o from Hordeum vulgare subsp spontaneum in Clipper*4 as C8. Mutant used for description and seed stocks: Rph10.o in Clipper C8 (GSHO 1588); Rph10.o in Bowman (PI 483237)*2 (GSHO 1995).1001707GSHO 1588
19GSHO 1589Rph11.pHordeum vulgare L. subsp. vulgare North Dakota, United StatesNSGCKERNELS1996DONATED04/1996Genetic materialThe seedling reaction varies from a necrotic fleck to a moderate sized uredinium with heterozygotes having an intermediate reaction. Culture Aust 220 from Australia was required to detect a resistant reaction in allelism tests and backcrosses. Seedlings tend to show a moderate sized uredinium as a reaction to many isolates of Puccinia hordei. Inheritance: Monofactorial dominant. Located in chromosome 6HL, about 7.0 cM proximal from the Acp3 (acid phosphatase 3) locus, and about 11.0 cM proximal from the Dip2 (dipeptidase 2) locus. Previous nomenclature and gene symbolization: None. Mutational events: Rph11.p from Hordeum vulgare subsp spontaneum in Clipper*4 as C67. Mutant used for description and seed stocks: Rph11.p in Clipper C67 (GSHO 1589); Rph11.p in Bowman (PI 483237)*2 (GSHO 2092).1001709GSHO 1589
20GSHO 1590Rph12.zHordeum vulgare L. subsp. vulgare North Dakota, United StatesNSGCKERNELSNot Available1996DONATED04/1996Genetic materialThe seedling reaction type is 0; or a necrotic fleck. Incomplete dominance was demonstrated by the intermediate reaction of heterozygotes to pathogenic isolates of Puccinia hordei. Inheritance: Monofactorial dominant. Located in chromosome 5HL [7L], about 26.1 cM distal from the raw1 (smooth awn 1) locus. Previous nomenclature and gene symbolization: Resistance to Puccinia hordei Otth 9=Pa9. Mutational events: Rph12.z in Triumph. Mutant used for description and seed stocks: Rph12.z in Triumph (GSHO 1590; Rph12.z in Bowman (PI 483237)*4 (GSHO 2145).1001755GSHO 1590
21GSHO 1593Rph2.bHordeum vulgare L. subsp. vulgare North Dakota, United StatesNSGCKERNELS1996DONATED04/1996Genetic materialThe seedling reaction type is 0n - 1c with race 4 culture 57-19; heterozygotes have reaction types ranging from 1 to 3, depending on parents. Responses will vary for homozygotes and heterozygotes when different rust cultures are tested. Inheritance: Monofactorial incomplete dominant. Located in chromosome 5HS [7S], not in the long arm; distal from the secondary constriction and 3.5 cM proximal from RFLP marker CD0749. Previous nomenclature and gene symbolization: Resistance to Puccinia anomola Rostr.=Pa. Resistance to Puccinia hordei Otth 2=Pa2. Resistance to Puccinia hordei A=A. Mutational events: Rph2.b in Peruvian; Rph2.j in Batna (CIho 3391); Rph2.k in Weider (No 22, PI 39398); Rph2.l in Juliaca (PI 39151); Rph2.m in Kwan (PI 39367, GSHO 1392); Rph2.n in Chilean D (PI 48136); an allele at the Rph2 locus is present in Purple Nepal (CIho 1373), Modia (CIho 2483), Morocco (CIho 4975), Barley 305 (CIho 6015), Marco (PI 94877); Austral (CIho 6358); Marocaine 079 (CIho 8334); Q21861 (PI 584766), TR306; accessions with a second Rph gene besides the Rph2 allele include Carre 180 (CIho 3390), CIho 14077; Ricardo (PI 45492); Ariana (CIho 14081); Quinn (PI 39401); Bolivia (PI 36360); Reka 1 (CIho 5051); tentative Rph2 allele symbols are Rph2.q in Quinn, Rph2.r in Bolivia (GSHO 1598), Rph2.s in Ricardo, Rph2.t in Reka 1 (GSHO 1594) , and Rph2.u in Ariana based on differential reactions and different cultivar origins; Rph2.y from HJ198*3/HS2310 (PI 531841, GSHO 1595). Mutant used for description and seed stocks: Rph2.b in Peruvian (GSHO 1593); Rph2.b in Bowman (PI 483237)*2 (GSHO 2320); Rph2.t in Bowman*5 (GSHO 2321).1001807GSHO 1593
22GSHO 1597Rph5.eHordeum vulgare L. subsp. vulgare North Dakota, United StatesNSGCKERNELSNot Available1996DONATED04/1996Genetic materialThe seedling reaction type is 0-0-c, with race 4 culture 57-19. Incomplete dominance was observed in segregating progenies. Inheritance: Monofactorial incomplete dominant. Located in chromosome 5H [7], about 30.5 cM from the Rph2 (reaction to Puccinia hordei 2) locus. Previous nomenclature and gene symbolization: Resistance to Puccinia hordei Otth 5=Pa5. Resistance to Puccina hordei B=B. Mutational events: Rph5.d in Quinn along with Rph2.q. Mutant used for description and seed stocks: Rph5.e in Bowman (PI 483237)*4 (GSHO 1865).1001898GSHO 1597
23GSHO 1598Rph6.fHordeum vulgare L. subsp. vulgare North Dakota, United StatesNSGCKERNELS1996DONATED04/1996Genetic materialThe seedling reaction type is 0-n - 1c with race 4 culture 57-19. Heterozygotes may have a type 2 or 3 reaction under certain conditions. Inheritance: Monofactorial dominant, but present with an allele at the Rph (reaction to Puccinia hordei 2) locus. Location is unknown. Previous nomenclature and gene symbolization: Resistance to Puccinia hordei Otth=X. Resistance to Puccinia hordei Otth=Pax. Mutational events: Rph6.f in Bolivia. Mutant used for description and seed stocks: Rph6.f in Bolivia (GSHO 1598); Rph6.f in Bowman (PI 483237)*4(GSHO 2323).1001942GSHO 1598
24GSHO 1600Rph8.hHordeum vulgare L. subsp. vulgare North Dakota, United StatesNSGCKERNELS1996DONATED04/1996Genetic materialThe seedling reaction type is 1-2 to 2CN with race 8 culture ND8702, depending on environmental conditions. Egypt 4 is susceptible to most isolates of Puccinia hordei, but a resistant reaction to a few isolates has been reported. Resistance was expressed better at 15 degrees C than at 20 degrees C, and incomplete dominance was observed in segregating progenies. Inheritance: Monofactorial dominant. Location is unknown. Previous nomenclature and gene symoblization: Resistance to Puccinia hordei Otth 8=Pa8. Mutational events: Rph8.h in Egypt 4. Mutant used for description and seed stocks: Rph8.h in Egypt 4 (GSHO 1600); Rph8.h in Bowman (PI 483237)*2 (GSHO 2324).1001990GSHO 1600
25GSHO 1601Rph9.iHordeum vulgare L. subsp. vulgare North Dakota, United StatesNSGCKERNELSNot Available1996DONATED04/1996Genetic materialThe seedling reaction type ranges from 0; or necrotic fleck to 23- or intermediate. The reaction of heterozygotes may be intermediate. The resistant reaction of the Rph9.i allele is temperature sensitive and is inactivated above 20 degrees C. Inheritance: Monofactorial dominant. Location is unknown, but about 30.4 cM from the Rph13 (reaction to Puccinia hordei 13) locus. Previous nomenclature and gene symbolization: Resistance to Puccinia hordei Ottth 9=Pa9. Mutational events: Rph9.i in Abyssinian. Mutant used for description and seed stocks: Rph9.i in Abyssinian (GSHO 1601); Rph9.i in Bowman (PI 483237)*5 (GSHO 1866).1002036GSHO 1601
26GSHO 1604Sil1.aHordeum vulgare L. subsp. vulgare Nebraska, United StatesNSGCKERNELS1996DEVELOPEDGenetic materialPlants have relatively short subcrown internodes, 0.0 to 1.0 cm, when planted 9 cm deep in vermiculite. Planting depth and subcrown internode length determine the position of the crown in relation to the soil surface. Plants with a deeper crown placement show higher levels of winter survival. In the Bowman backcross-derived line, the Sil gene has a dominant inheritance pattern and shows a repulsion linkage with the sls gene. Inheritance: Monofactorial incomplete dominant. Located in chromosome 1HS [5S], based on linkage drag with the sls (small lateral spikelet) locus. Previous nomenclature and gene symbolization: None. Mutational events: Sil1.a in NE 62203. Mutant used for description and seed stocks: Sil1.a in NE 62203 (GSHO 1604); Sil1.a in Bowman (PI 483237)*5 (GSHO 2032).1002079GSHO 1604
27GSHO 1606Xnt1.aHordeum vulgare L. subsp. vulgare JapanNSGCKERNELS1996DEVELOPEDGenetic materialThe original M2 line segregated for xantha, chlorina, and green seedlings, but only the last two classes survived to maturity. Chlorina plants have a slightly yellow-green color until near maturity. Crosses to the chlorina plant produced chlorina and green F1 plants in 1:1 ratio, and in the F2 progeny of chlorina plants xantha, chlorina, and green plants appeared in a 1:2:1 ratio. Color differences between normal and heterozygous plants were observed only prior to heading for the Xnt1.a2 mutant. Mutants at the Xnt1 locus must be maintained as heterozygous stocks. Inheritance: Monofactorial incomplete dominant. Located in chromosome 7HL [1L], over 35.9 cM distal from the lks2 (short awn 2) locus. Previous nomenclature and gene symbolization: Xantha a = Xa. Mutational events: Xnt1.a in the hybrid Akashinriki X linkage tester 191 (OUM215); Xnt1.a2 in the translocation T d; Xnt1.a3 in MC20. Mutant used for description and seed stocks: Xnt1.a in the hybrid Akashinriki X linkage tester 191 (GSHO 1606); Xnt1.a in Bowman (PI 483237)*7(GSHO 1862).1489961GSHO 1606
28GSHO 1613Zeo1.aHordeum vulgare L. subsp. vulgare GermanyNSGCKERNELS1996DEVELOPEDGenetic materialPlants heterozygous for Zeo1 have short culms, compact spikes, and wide kernels. Homozygotes have shorter culms (short peduncle), very compact spikes, large outer glumes with long awns, and reduced fertility. Generally, the spike emerges from the side of the sheath in homozygotes. Although the name zeocriton is used for this gene, this gene is not from Spratt, the dense ear type described by Engledow. Inheritance: Monofactorial incomplete dominant. Located in chromosome 2HL, about 9.2 cM distal from the lig (liguleless) locus. Previous nomenclature and gene symbolization: "Kurz und dicht"=Knd. Mutational events: Zeo1.a in Donaria (Mut 2657, DWS1340); Zeo1.b, received as "Kurz und dicht" and placed in Wolfe's Multiple Dominant Marker Stock (DWS1163, GSHO 1614), was probably derived from Mut 2657. Mutant used for description and seed stocks: Zeo1.a in Donaria (GSHO 1613); Zeo1.a in Bowman (PI 483237)*6(GSHO 1931); Zeo1.b in Bowman*9 (GSHO 1932).1002884GSHO 1613
29GSHO 1620ant1.1Hordeum vulgare L. subsp. vulgare North Dakota, United StatesNSGCKERNELS1996DONATED04/1996Genetic materialWhen grown under favorable light conditions, red pigmentation of the stem does not occur. Anthocyanin pigments are not observed in the stem, auricles, awns, or lemma of ant1 mutants. However, slight pigmentation of these plant parts may be observed occasionally in plants homozygous for the normal allele Ant1.a at this locus. Expression of alleles at the ant1 locus is easier to observe at the stem base of seedlings. Inheritance: Monofactorial recessive. Located in chromosome 7HS [1S], near the centromere, near the ert-m (erectoides-m) locus, and about 15.0 cM distal from the nud (naked caryopsis) locus. Previous nomenclature and gene symbolization: Green stem=rs. Exrubrum-a=rub-a. Mutational events: ant1.b (rs1.b) in Manchurian introductions; ant1.1, ant1.2, ant1.4 in Bonus (PI 189763); ant1.56 in Bonus. Mutant used for description and seed stocks: ant1.1 in Bonus (GSHO 1620); Rst1.a in Goldfoil (GSHO 185, CIho 1834); ant1.b in Bowman (PI 483237) from six-rowed Manchurian type cultivars.1003000GSHO 1620
30GSHO 1632ant2.20Hordeum vulgare L. subsp. vulgare North Dakota, United StatesNSGCKERNELS1996DONATED04/1996Genetic materialAnthocyanin pigments are not observed in any vegetative plant parts, including the stem, auricles, lemma, and awn. The straw does not develop a purple pigmentation as it approaches maturity. The recommended symbol for the dominant allele is Ant2.c (formally Pr1.a). Inheritance: Monofactorial recessive. Located in chromosome 2HL, about 15.1 cM distal from the vrs1 (six-rowed spike 1) locus. Previous nomenclature and gene symbolization: Non-purple straw=pr or pr. Anthocyanin-less=ant-2. Exrubrum =rub. Colorless leaf tip=clt2, c2. Mutational events: ant2.d (pr1.b) in Alva (NSGC1866), ant2.e (pr1.c) in Balder (PI 195481), ant2.f (pr1.d) in Cambrinus (PI 321779), ant2.g (pr1.e) in Sultan (PI 339814); ant2.15, 2.20, 2,23, 2,25, 2,26, 2,27, in Foma (CIho 11333); ant2.41 in Mari (CIho 11334, 2.46 in Foma, 2.47, 2.48, 2.49, 2.50, 2.51, 2.54, 2.55 in Bonus (PI 189763), 2.112, 2.113, 2.114, 2.115, 2.116, 2.117, 2.118, 2.120, 2.122, 2.130 in Nordal; ant2.h (pr1.f) in Shyri (GSHO 1632). Mutant used for description and seed stocks: ant2.20 in Foma (GSHO 1632); ant2.h from Shyri in Bowman (PI 483237)* 5 (GSHO 1919); ant2.20 in Bowman*2 (GSHO 1920).1003670GSHO 1632
31GSHO 1648ari-a.6Hordeum vulgare L. subsp. vulgare Skåne län, SwedenNSGCKERNELS1996DEVELOPED58.98333300, 13.50000000Genetic materialA common feature of mutants at this locus is instability in the expression of awn length, ranging from very short to 5/6 normal length in various environments. In some environments, secondary or tertiary tillers may show a drastic reduction in stem in stem internode, spike, and awn length. Awns are thin and brittle and may flare somewhat on later tillers. Kernels are thin with fine wrinkles on the lemma. Inheritance: Monofactorial recessive. Located in chromosome 3HS, probably less than 2.4 cM from the centromere. Previous nomenclature and gene symbolization: Breviaristatum-6=ari-6. Short awn 7=lk7. Mutational events: ari-a.6, -a.7 in Bonus; ari-a.8, -a.23 in Bonus; ari-a.26 in Bonus; ari-a.31 in Bonus; ari-a.37, -a.48 in Bonus, -a.118, -a.121, -a.124, -a.126 in Foma (CIho 11333); ari-a.122, -a.125, -a.148 in Foma; ari-a.152, -a.162, -a.174, -a.211 in Foma; ari-a.216 in Foma; ari-a.273, -a.277, -a.279, -a.287, -a.293 in Kristina; ari-a.296, -a.305 in Kristina; lks7.i (Kmut 114) in Asahi 5 (OUJ509). Mutant used for description and seed stocks: ari-a.6 in Bonus (GSHO 1648); ari-a.6 in Bowman (PI 483237(*7 (GSHO 1958).1003911GSHO 1648
32GSHO 1649ari-b.11Hordeum vulgare L. subsp. vulgare Skåne län, Sweden SPIKEHistoric1996DEVELOPED58.98333300, 13.50000000Genetic materialPlants are semidwarf with reduced viability and fertility; leaves are broad, but shortened and lighter green in color; and awns are about 3/4 normal length. Growth is markedly retarded during warm seasons and relatively good during cool seasons. Development of reproductive organs is delayed and sterility causes a stock maintenance problem. Inheritance: Monofactorial recessive. Location is unknown. Previous nomenclature and gene symbolization: Breviaristatum-11=ari-11. Mutational events: ari-b.11, -b.13, -b.17, -b.19, -b.29 in Bonus, -b.175, -b.226, -b.238 in Foma (CIho 11333). Mutant used for description and seed stocks: ari-b.11 in Bonus (GSHO 1649); ari-b.11 in Bowman (PI 483237)*3 (GSHO 2157).1003916GSHO 1649
33GSHO 1650gig1.aHordeum vulgare L. subsp. vulgare Colorado, United StatesNSGCKERNELS1996DEVELOPEDGenetic materialCulm length and diameter, leaves, spikes, and glumes are greater than the control; awns are shorter than the control; leaves are dark green; spikes are lax; spikes, culms, and leaf sheaths have a heavy wax coating. Anther development is good, but the stigma has few stigma hairs and seed set is poor. In the Bowman backcross-derived line, plant height is normal and plants appear normal except for the spikelets and a heavier wax coating. The lemma is large and long, awns are 1/2 normal, stigma hairs are few, and selfed seed set is 10 to 15%. Hand pollination can increase seed set. Inheritance: Monofactorial recessive. Location in chromosome 2H is possible, and a possible linkage drag association with Gth (toothed lemma) locus in chromosome 2H was reported. Previous nomenclature and gene symbolization: Female sterile=sf. Gigas=gig. Mutational events: gig1.a in Tochigi Golden Melon (Kmut 72); gig1.b in Montcalm//Galore/Daples. Mutant used for description and seed stock: gig1.a in Tochigi Golden Melon (GSHO 1650); gig1.a in Bowman (PI 483237)*7 (GSHO 1924).1003947GSHO 1650
34GSHO 1653ari-e.1Hordeum vulgare L. subsp. vulgare Skåne län, SwedenNSGCKERNELS1996DEVELOPED58.98333300, 13.50000000Genetic materialAwns are about 2/3 normal, plants are 3/4 to 5/6 normal height, and kernels are smaller. Awns and spikes tend to be more erect than those of normal sibs. Mutant allelism at the ari-e locus are associated with salt tolerance, lower accumulation of Na+ and relatively insensitive to GA3. Inheritance: Monofactorial recessive. Located in chromosome 5HL [7L], about 5.6 cM proximal from the cer-zj (eceriferumm-zj) locus. Previous nomenclature and gene symbolization: Breviaristatum-1=ari-1. Short awn 9=lk9. Golden Promise erectoides=GPert. Mutational events: ari-e.1 in Bonus; ari-e.30, -e.39 in Bonus; ari-e.119, -e.156 in Foma (CIho 11333); ari-e.166, -e.178, -e.222 in Foma; ari-e.228 in Foma; ari-e.GP (Golden Promise, PI 343079) in Maythorpe (PI 260893). Mutant used for description and seed stocks: ari-e.1 in Bonus (GSHO 1653); ari-e.GP in Maythorpe (GSHO 1733); ari-e.1 in Bowman (PI 483237)*7 (GSHO 2104); ari-e.GP in Bowman*7 (GSHO 2105).1490019GSHO 1653
35GSHO 1654ari-f.21Hordeum vulgare L. subsp. vulgare Skåne län, SwedenNSGCKERNELS1996DEVELOPED58.98333300, 13.50000000Genetic materialAwns are thin and often 1/2 to 3/4 normal length, and awn length decreases toward the top of the spike. Spikes are more lax and have fewer rachis internodes than those of parental cultivars. Deformation of the lemma apex causes the apical kernels to be naked. Inheritance: Monofactorial recessive. Location is unknown. Previous nomenclature and gene symbolization: Breviaristatum-21=ari-21. Mutational events: ari-f.21 in Bonus; ari-f.42, -f.47 in Bonus; ari-f.52, -f.54, -f.55 in Bonus; ari-f.113 in Foma (CIho 11333); ari-f.115 in Foma; ari-f.123, -f.136 in Foma; ari-f.155 in Foma; ari-f.168, -f.169, -f.170, -f.224, -f.225, -f.233, -f.235 in Foma, -f.253, -f.254, -f.260, -f.266 in Kristina; ari-f.268, -f.275, -f.294, -f.295, -f.300 in Kristina. Mutant used for description and seed stocks: ari-f.21 in Bonus (GSHO 1654); ari-f.21 in Bowman (PI 483237)*5 (GSHO 2158).1490020GSHO 1654
36GSHO 1655ari-g.18Hordeum vulgare L. subsp. vulgare Skåne län, SwedenNSGCKERNELS1996DEVELOPED58.98333300, 13.50000000Genetic materialPlants are semidwarf and very weak. Awns are 1/4 to 1/3 normal and stiff, spikes are short and small, and seed set is low. Leaf blades are very narrow and often involuted. When grown in the field at Fargo, North Dakota, USA, plant development is very slow, the upper portion of the sheath is white as new leaves emerge, and spikes are seldom produced. Inheritance: Monofactorial recessive. Located in chromosome 2H; the linkage drag association with the Gth (toothed lemma) locus may be caused by poor expression of the Gth1.a allele in the Bowman backcross-derived line. Previous nomenclature and gene symbolization: Breviaristatum-18=ari-18. Short awn 10-lk10. Mutational events: ari-g.18 in Bonus; ari-g.24 in Bonus; ari-g.134, -g.138, -g.142, -g.158, -g.221, -g.234, -g.246 in Foma (CIho 11333), -g.267 in Kristina; ari-g.303 in Kristina. Mutant used for description and seed stocks: ari-g.18 in Bonus (GSHO 1655); ari-g.18 in Bowman (PI 483237, GSHO 2503)*7.1490021GSHO 1655
37GSHO 1658ari-j.32Hordeum vulgare L. subsp. vulgare Skåne län, SwedenNSGCKERNELS1996DEVELOPED58.98333300, 13.50000000Genetic materialPlants are semidwarf (3/4 of normal culm length) and awns are about 5/6 normal length. The peduncle is short and coiled or kinked, and as a result the spike rarely emerges above the flag leaf. Inheritance: Monofactorial recessive. Location is unknown. Previous nomenclature and gene symbolization: Breviaristatum-32=ari-32. Mutational events: ari-j.32 in Bonus; ari-j.101 in Foma (CIho 11333); ari-j.167 in Foma. Mutant used for description and seed stocks: ari-j.32 in Bonus (GSHO 1658); ari-j.32 in Bowman (PI 483237)*5 (GSHO 2159).1004293GSHO 1658
38GSHO 1659ari-k.504Hordeum vulgare L. subsp. vulgare Skåne län, SwedenNSGCKERNELS1996DEVELOPED58.98333300, 13.50000000Genetic materialAwns are often normal length, but they are dehiscent (dearistatum) at maturity. Variability in awn length has been observed over environments and awns seem thinner than normal. Awns of plants grown in fall greenhouse nurseries at Fargo, North Dakota, USA are very short and thread-like. Inheritance: Monofactorial recessive. Location is unknown. Previous nomenclature and gene symbolization: Breviaristatum-504=ari-504. Dearistatu-k=ari-k. Mutational events: ari-k.501, -k.502, -k.503, -k.504 in Bonus. Mutant used for description and seed stocks: ar-k.504 in Bonus (GSHO 1659); ari-k.504 in Bowman (PI 483237)*3 (GSHO 2160).1004308GSHO 1659
39GSHO 1661ari-mHordeum vulgare L. subsp. vulgare Skåne län, SwedenNSGCKERNELS1996DEVELOPED58.98333300, 13.50000000Genetic materialPlants are semidwarf (2/3 normal height) and were placed in the brachytic class of semidwarfs. Kernels are rounded and awns are about 1/2 normal length. Spikes are slightly more lax than normal. Inheritance: Monofactorial recessive. Location is unknown. Previous nomenclature and gene symbolization: Breviaristatum-28=ari-28. Mutational events: ari-m.12, -m.28 in Bonus, -m.141, -m.177 in Foma (CIho 11333), -m.251 in Kristina; ari-m.269 in Kristina. Mutant used for description and seed stocks: ari-m.12 in Bonus (GSHO 1661); ari-m.28 in Bowman (PI 483237)*6 (GSHO 2161).1004326GSHO 1661
40GSHO 1662ari-n.45Hordeum vulgare L. subsp. vulgare Skåne län, SwedenNSGCKERNELS1996DEVELOPED58.98333300, 13.50000000Genetic materialAwns are 3/4 normal and arise at an oblique angle, and they are drastically shortened towards the top of the spike. Plants are normal height, but spikes are short due to a reduced number of rachis internodes and apical abortion. Inheritance: Monofactorial recessive. Location is unknown. Previous nomenclature and gene symbolization: Breviaristatum-45=ari-45. Mutational events: ari-n.45, -n.46 in Bonus, -n.147, -n.163, -n.164, -n.181, -n.219, -n.220, -n.231 in Foma (CIho 11333), -n.258, -n.292 in Kristina. Mutant used for description and seed stocks: ari-n.45 in Bonus (GSHO 1662).1004330GSHO 1662
41GSHO 1663ari-o.40Hordeum vulgare L. subsp. vulgare Skåne län, SwedenNSGCKERNELS1996DEVELOPED58.98333300, 13.50000000Genetic materialPlants are semidwarf (2/3 normal height) and were placed in the brachytic class of semidwarfs. The culm may have a short or extra internode and a leafy bract below the spike. The peduncle is often slightly coiled, and the basal rachis internode is elongated. Kernels are rounded, and awns are about 3/4 normal length. Inheritance: Monofactorial recessive. Location is unknown. Previous nomenclature and gene symbolization: Breviaristatum-40=ari-40. Mutational events: ari-o.40, -o.43 in Bonus; ari-o.143 in Foma (CIho 11333), -o.297, -o.301, -o.304, -o.306 in Kristina. Mutant used for description and seed stocks: ari-o.40 in Bonus (GSHO 1663); ari-o.40 in Bowman (PI 483237)*6 (GSHO 2162).1004349GSHO 1663
42GSHO 1664ari-p.27Hordeum vulgare L. subsp. vulgare Skåne län, SwedenNSGCKERNELS1996DEVELOPED58.98333300, 13.50000000Genetic materialAwns are about 3/4 normal, and sterile spikelets occur at random in the original stocks. Plants in the Bowman backcross-derived line head very late. Inheritance: Monofactorial recessive. Location is unknown. Previous nomenclature and gene symbolization: Breviaristatum-27=ari-27. Mutational events: ari-p.27 in Bonus; ari-p.108, -p.172, -p.208, -p.209, -p.223 in Foma (CIho 11333); ari-p.278 in Kristina. Mutant used for description and seed stocks: ari-p.27 in Bonus (GSHO 1664); ari-p.27 in Bowman (PI 483237)*3 (GSHO 2163).1004354GSHO 1664
43GSHO 1665ari-q.280Hordeum vulgare L. subsp. vulgare Skåne län, SwedenNSGCKERNELS1996DEVELOPED58.98333300, 13.50000000Genetic materialThe spike is short with irregular placement of spikelets. Awns are about 3/4 normal length. Lateral spikelets are missing from a few basal rachis nodes and two awns occur occasionally on the terminal spikelet. Plants are about 3/4 normal height. Inheritance: Monofactorial recessive. Location is unknown. Previous nomenclature and gene symbolization: Breviaristatum-271=ari-271. Mutational events ari-q.271, -q.280, -q.281 in Kristina. Mutant used for description and seed stocks: ari-q.280 in Kristina (GSHO 1665); ari-q.280 in Bowman (PI 483237)*4 (GSHO 2164).1004374GSHO 1665
44GSHO 1666ari-r.14Hordeum vulgare L. subsp. vulgare Skåne län, Sweden SPIKEHistoric1996DEVELOPED58.98333300, 13.50000000Genetic materialPlants are semidwarf (less than 1/2 normal height) with compact spikes and rounded kernels. Awns are stiff and less than 1/3 normal length. Plants are vigorous, but heading is delayed. This mutant was placed in brachytic class of semidwarfs based on growth pattern. Inheritance: Monofactorial recessive. Location is unknown. Previous nomenclature and gene symbolization: Breviaristatum-14=ari-14. Mutational events: ari-r.14 in Bonus, -r.202 in Foma (CIho 11333); ari-r.236 in Foma; ari-r.282 in Kristina; ari-r.314, -r.318 in Kristina. Mutant used for description and seed stocks: ari-r.14 in Bonus (GSHO 1666); ari-r.14 in Bowman (PI 483237)*6 (GSHO 2165).1004379GSHO 1666
45GSHO 1667blf2.bHordeum vulgare L. subsp. vulgare Arizona, United StatesNSGCKERNELS1996DEVELOPEDGenetic materialLeaf width of mutant plants is narrower (about 2/3 normal) than those of Hannchen. Other plant characteristics appear normal. Inheritance: Monofactorial recessive. Located in chromosome 5HL [7L], based on linkage drag with the raw1 (smooth awn 1) locus. Previous nomenclature and gene symbolization: Narrow leaf=nlh. Broad leaf 2=bb2. Narrow leaf-k=nl.k. Mutational events: blf2.b in Hannchen. Mutant used for description and seed stocks: blf2.b in Hannchen (GSHO 1667); blf2.b in Bowman (PI 483237)*6 (GSHO 2136).1004397GSHO 1667
46GSHO 1696bra-d.7Hordeum vulgare L. subsp. vulgare Skåne län, SwedenNSGCKERNELS1996DEVELOPED58.98333300, 13.50000000Genetic materialThe characteristic trait of this mutant is the presence of a bract (third outer glume) outside the two empty glumes of the central spikelet. The bract associated with the lowest spikelet is always the largest, embracing in some cases about one-half the spike. Bracts become progressively smaller toward the tip of the spike and are associated with elongated basal rachis internodes. Inheritance: Monofactorial recessive. Location is unknown. Previous nomenclature and gene symbolization: None. Mutational events: bra-d.7 in Foma. Mutant used for description and seed stocks: bra-d.7 in Foma (T 41-31); bra-d.7 in Bowman (PI 483237)*2 (GSHO 2185).1005198GSHO 1696
47GSHO 1702com1.aHordeum vulgare L. subsp. vulgare Skåne län, SwedenNSGCKERNELS1996DEVELOPED58.98333300, 13.50000000Genetic materialThe spike is branched from the rachilla of the bottom 1 to 10 spikelets. Awns, which vary from normal to thread-like, and protruding branches are bent in various directions because of "packing" problems in the boot. More spike branches develop under favorable environmental conditions. Inheritance: Monofactorial recessive. Located in chromosome 5HL [7L], between the ert-n (erectoide-n) and the lax-a (laxatum-a) loci, but widely different estimates of their distance from the srh (short rachilla hair) locus are reported. Previous nomenclature and gene symbolization: Laxatum-270=lax-270. Short-haired, inflorescent rachilla=si. Branching, inflorescent rachilla=bir1a. Compositum=com1a. Mutational events: com1.a (lax.270), com1.b (lax.271), com1.c (lax.272), com1.d (lax.206), com1.e (lax.213) in Foma. Mutant used for description and seed stocks: com1.a in Foma (GSHO 1702); com1.a in Bowman (PI 483237)*6 (GSHO 2098).1490028GSHO 1702
48GSHO 1703com2.gHordeum vulgare L. subsp. vulgare North Dakota, United StatesNSGCKERNELS1996DONATED04/1996Genetic materialThe spike is malformed because the rachilla fails to abort and extra spikelets and rachis branches may form at rachis nodes in the basal half of the spike. More spike branches may develop under favorable environmental conditions. Inheritance: Monofactorial recessive. Located in chromosome 2HS, about 19.7 cM distal from the eog (elongated outer glume) locus. Previous nomenclature and gene symbolization: Branching inflorescent, rachilla 2=bir2. Mutational events: com2.f in Donaria (Mut 2201); com2.g a freak from the CIMMYT collection (DWS1358); com2.g may be different from the original source because it carries the Eam1 (early maturity 1) gene as a linked marker. Mutant used for description and seed stocks: com2.g in CIMMYT freak (GSHO 1703); com2.g in Bowman (PI 483237)*8 (GSHO 1878).1490029GSHO 1703
49GSHO 1705cur1.aHordeum vulgare L. subsp. vulgare North Dakota, United StatesNSGCKERNELS1996DONATED04/1996Genetic materialAll plant parts appear curved or twisted. Lemmas and awns are extremely curly. The rachis is usually bent, and tillers are curved or wavy. Inheritance: Monofactorial recessive. Located in chromosome 6HL, over 31.5 cM distal from the rob (orange lemma) locus, not linked to rob. Previous nomenclature and gene symbolization: Curly1=cu1. Mutational events: cur1.a in 48-cr cr-17 (Alb Acc 23, G395). Mutant used for description and seed stocks: cur1.a in 48-cr cr-17 (GSHO 1705); cur1.a in Bowman (PI 483237)*4 (GSHO 2088).1490031GSHO 1705
50GSHO 1707cur3.dHordeum vulgare L. subsp. vulgare JapanNSGCKERNELS1996DEVELOPEDGenetic materialStem internodes are curved in a bow-like manner and awns are slightly coiled. Seedlings elongate rapidly and leaves are nearly twice as long as normal. Seedlings homozygous for the cur3.e allele do not show additional elongation in response to gibberellic acid treatment. Plants in the Bowman backcross-derived line are shorter (5/6 normal), have short spikes (2/3 normal), and yield much less. Inheritance: Monofactorial recessive. Located in chromosome 6Hl, about 22.9 cM distal from the cul2 (uniculm 2) locus. Previous nomenclature and gene symbolization: Curly 3=cu3. Mutational events: cur3.e in Akashinriki (OUM301). Mutant used for description and seed stocks: cur3.e in Akashinriki (GSHO 1707); cur3.e in Bowman (PI 483237)*6 (GSHO 2087).1005507GSHO 1707
51GSHO 1708cur4.fHordeum vulgare L. subsp. vulgare North Dakota, United StatesNSGCKERNELS1996DONATED04/1996Genetic materialDiagnostic characteristics can be observed from the seedling stage to maturity. Roots are curved compared to straight roots in normal plants. Leaves tend to coil or bend and may have wrinkles at the margins. Culms are bent slightly at the nodes and about 3/4 normal length, and awns are frequently slightly coiled. Inheritance: Monofactorial recessive. Located in chromosome 2HL, close to the Gth (toothed lemma) locus based on linkage drag with the glo-d.1006 allele at the cur4 locus. Previous nomenclature and gene symbolization: Curly 4=cu4. Spiral neck=spn. Globosum-d=glo-d. Mutational events: cur4.f in Asahi 5 (Kmut 118); glo-d.1006 (1114/66, GSHO 1754) in Donaria (PI 161974); glo-d.13 (glo-d.1009), glo-d.14 in Bonus (PI 189763); cur4.i (OUM163, GSHO 1709) in Akashinriki (OUJ659, PI 467400). Mutant used for description and seed stocks: cur4.f in Asahi 5 (GSHO 1708); cur4.f in Bowman (PI 483237)*7 (GSHO 1915); glo-d.1006 in Bowman*7 (GSHO 1917); cur4.i in Bowman*7 (GSHO 1916).1005508GSHO 1708
52GSHO 1710cur5.hHordeum vulgare L. subsp. vulgare North Dakota, United StatesNSGCKERNELS1996DONATED04/1996Genetic materialLeaves are short and partially coiled, and awns are slightly coiled. Plants are semidwarf (2/3 normal height), have relatively short spikes, and much reduced vigor. Inheritance: Monofactorial recessive. Located in chromosome 1HS [5S], based on l linkage drag with the sls (small lateral spikelet) locus. Previous nomenclature and gene symbolization: Curly 5=cu5. Mutational events: cur5.h in Glenn. Mutant used for description and seed stocks: cur5.h in Glenn (GSHO 1710); cur5.h in Bowman (PI 483237)*7 (GSHO 2045).1005528GSHO 1710
53GSHO 1711cud1.aHordeum vulgare L. subsp. vulgare JapanNSGCKERNELS1996DEVELOPEDGenetic materialMost leaves are short and more less twisted, and they often have a longitudinal crease. Culms are short (1/2 normal), the spike is compact, awns are short (1/3 normal), and kernels are small and round. Inheritance: Monofactorial recessive. Located in chromosome 5HL [7L], about 14.4 cM proximal from the fst1 (fragile stem 1) locus. Previous nomenclature and gene symbolization: Curly dwarf=cud. Mutational events: cud1.a in Akashinriki (OUM120). Mutant used for description and seed stocks: cud1.a in Akashinriki (GSHO 1711); cud1.a in Bowman (PI 483237)*6 (GSHO 2094).1005543GSHO 1711
54GSHO 1712cud2.bHordeum vulgare L. subsp. vulgare JapanNSGCKERNELS1996DEVELOPEDGenetic materialLemmas and awns are curly or strongly twisted, and stem internodes are curved. Culms are short (1/2 normal), spikes are semi-compact, awns are short (2/3 normal), and kernels are globe-shaped. Most leaves are short and slightly twisted. Inheritance: Monofactorial recessive. Located in chromosome 1HL [5L], about 14.6 cM proximal from the Blp (black lemma and pericarp) locus. Previous nomenclature and gene symbolization: None. Mutational events: cud2.b in Akashinriki (OUM112). Mutant used for description and seed stocks: cud2.b in Akashinriki (GSHO 1712); cud2.b in Bowman (PI 483237)*3 (GSHO 2062).1005544GSHO 1712
55GSHO 1714dsk1.aHordeum vulgare L. subsp. vulgare JapanNSGCKERNELS1996DEVELOPEDGenetic materialAfter heading, leaves begin to dehydrate and turn a dusky, olive-gray, color. Many, small dark spots occur scattered over the surface of the olive-gray plant parts. Even after maturity, spikes and culms remain faint green in color. Bowman backcross-derived plants grown in the greenhouse are about 2/3 normal height and moisture stress causes lower leaves to die rapidly without loss of green pigments. Anthers are pale green instead of yellow at anthesis. Inheritance: Monofactorial recessive. Located in chromosome 5HL [7L], about 17.6 cM distal from the srh (short rachilla hair) locus and 7.8 cM proximal from the raw1 (smooth awn1) locus. Previous nomenclature and gene symbolization: None. Mutational events: dsk1.a in Chikurin Ibaragi 1 (OUM299). Mutant used for description and seed stocks: dsk1.a in Chikurin Ibaragi 1 (GSHO 1714); dsk1.a in Bowman (PI 483237)*4 (GSHO 2120).1005562GSHO 1714
56GSHO 1738fch11.tHordeum vulgare L. subsp. vulgare Minnesota, United StatesNSGCKERNELS1996DEVELOPEDGenetic materialSeedlings are pale yellow-green and often have white blotches on the seedling leaves. Plants remain light green until maturity. Plant vigor is good, but plant developmental stages are delayed. Inheritance: Monofactorial recessive. Located in chromosome 6HL, about 6.1 cM distal from the rob (orange lemma) locus. Previous nomenclature and gene symbolization: Pale green=pg. Chlorina seedling 11=g11. Mutational events: fch11.t in Himalaya (GBC357 and GBC359). Mutant used for description and seed stocks: fch11.t in Himalaya (GSHO 1738); fch11.t in Bowman (PI 483237)*6 (GSHO 2082).1005775GSHO 1738
57GSHO 1739fch14.wHordeum vulgare L. subsp. vulgare North Dakota, United StatesNSGCKERNELS1996DEVELOPEDGenetic materialSeedlings have a pale yellow-green color. The leaves gradually become greener starting at the tip of the leaf blade, and mutant plants are indistinguishable in color from normal sibs at heading. When grown in the field, plants produce fewer tillers than normal plants and maturity is delayed. Inheritance: Monofactorial recessive. Located in chromosome 2HL, probably between the vrs1 (six-rowed spike 1) and the ant2 (anthocyanin-less 2) loci. Previous nomenclature and gene symbolization: Chlorina seedling 14=f14. Mutational events fch14.w in Shyri. Mutant used for description and seed stocks: fch14.w in Shyri (GSHO 1739); fch14.w in Bowman (PI 483237)*6 (GSHO 1911).1005794GSHO 1739
58GSHO 1742flo-b.3Hordeum vulgare L. subsp. vulgare Skåne län, SwedenNSGCKERNELS1996DEVELOPED58.98333300, 13.50000000Genetic materialExtra bracts develop occasionally at the base of the central spikelet on the abaxial side. Formation of the extra bracts is most common in the central portion of the spike, but rarely will the bracts from another spikelet. Inheritance: Monofactorial recessive. Located in chromosome 5HL [7L], near the raw1 (smooth awn 1) locus based on linkage drag. Previous nomenclature and gene symbolization: None. Mutational events: flo-b.3 in Foma. Mutant used for description and seed stocks: flo-b.3 in Foma (GSHO 1742); flo-b.3 in Bowman (PI 483237)*6 (GSHO 2128).1005815GSHO 1742
59GSHO 1743flo-c.5Hordeum vulgare L. subsp. vulgare Skåne län, SwedenNSGCKERNELS1996DEVELOPED58.98333300, 13.50000000Genetic materialExtra bracts develop occasionally at the base of the central spikelet on the abaxial side. Formation of the extra bracts is most common in the central portion of the spike, but rarely will the bracts form another spikelet. Inheritance: Monofactorial recessive. Located in chromosome 2HS, over 33.8 cM from the Gth (toothed lemma) locus. Previous nomenclature and gene symbolization: None. Mutational events: flo-c.5 in Foma. Mutant used for description and seed stocks: flo-c.5 in Foma (GHSO 1743); flo-c.5 in Bowman (PI 483237)*7 (GSHO 1877).1005838GSHO 1743
60GSHO 1744fol-a.1Hordeum vulgare L. subsp. vulgare Bayern, GermanyNSGCKERNELS1996DEVELOPED49.00000000, 11.50000000Genetic materialMost organs are reduced in size. Plants have narrow, dark green leaves and a stunted, grassy growth habit. Plant height is about 2/3 normal and spikes are 1/2 normal. Spikelets are about 2/3 normal size and the kernels are thin. Inheritance: Monofactorial recessive, but heterozygotes can be indentified based on leaf width. Located in chromosome 2HL, about 19.0 cM distal from the vrs1 (six-rowed spike 1) locus. Previous nomenclature and gene symbolization: None. Mutational events: fol-a.1 in Proctor (1135/64). Mutant used for description and seed stocks: fol-a.1 in Proctor (GSHO 1744); fol-a.1 in Bowman (PI 483237)*7 (GSHO 1927).1005839GSHO 1744
61GSHO 1746fst3.cHordeum vulgare L. subsp. vulgare Japan ImageHistoric1996DEVELOPEDGenetic materialThe leaf and stem characteristics of fst3 plants are similar to those of fst1 plants. Leaves and stems are very fragile and easily broken when slightly bent. Plants are about 2/3 normal height when protected in the greenhouse, but in the field plants are weak and are easily damaged by wind. Inheritance: Monofactorial recessive. Located in chromosome 7HS [1S], probably proximal from the dsp1 (dense spike 1) locus based on linkage drag. Previous nomenclature and gene symbolization: Fragile stem 3 =fs3. Mutational events: fst3.c in Kobinkatagi 4 (OUM382). Mutant used for description and seed stocks: fst3.c in Kobinkatagi 4 (GSHO 1746); fst3.c in Bowman (PI 483237) * 5 (GSHO 1842).1005855GSHO 1746
62GSHO 1755glo-e.15Hordeum vulgare L. subsp. vulgare Skåne län, SwedenNSGCKERNELS1996DEVELOPED58.98333300, 13.50000000Genetic materialPlants appear normal, but kernels are larger and more rounded than those of normal sibs. Inheritance: Monofactorial recessive. Located in chromosome 1HL [5L]. Previous nomenclature and gene symbolization: None. Mutational events: glo-e.15 (glo-e. 1010) in Foma. Mutant used for description and seed stocks: glo-e.15 in Foma (GSHO 1755); glo-e.15 in Bowman (PI 483237)*7 (GSHO 2050).1005959GSHO 1755
63GSHO 1757gra-a.1Hordeum vulgare L. subsp. vulgare Bayern, GermanyNSGCKERNELS1996DEVELOPED49.00000000, 11.50000000Genetic materialPlants have numerous, thin tillers with narrow leaves and short internodes (1/2 normal plant height). Spikes are less than half normal length and seeds are thin and small. Spikes emerge from the sheath of the flag leaf prematurely, 4 to 7 days before anthesis. Inheritance: Monofactorial recessive. Located in chromosome 3HL. Previous nomenclature and gene symbolization: Granum-a=gran-a. Mutational events: gra-a.1 (DWS1145) in Donaria; gra-a.2 (DWS1237, GSHO 1758) from a Tokak (PI 178421) mutant in OR-SS-2; gra-a.3 (DWS1176) in HE2816 (likely from HE-MN-C 292) from Milan Vasa. Mutant used for description and seed stocks: gra-a.1 (GSHO 1757); gra-a.1 in Bowman (PI 483237)*7 (GSHO 1980); gra-a.2 in Bowman*7 (GSHO 1981).1005980GSHO 1757
64GSHO 1759gsh7.aeHordeum vulgare L. subsp. vulgare Japan SPIKEHistoric1996DEVELOPEDGenetic materialThe spike, leaf sheath and stem of mutant plants are glossy and bright green in color (wax code --++). In early generation backcrosses to Bowman, the glossy sheath 7 segregates headed early and had poor vigor and small spikes. Also, spikes frequently abort after emasculation. Inheritance: Monofactorial recessive. Location is unknown. Previous nomenclature and gene symbolization: Glossy sheath 7=gs7. Mutational events: gsh7.ae in Akashinriki (OUM022). Mutant used for description and seed stocks: gsh7.ae in Akashinriki (GSHO 1759); gsh7.ae in Bowman (PI 483237)*5(GSHO 2269).1006000GSHO 1759
65GSHO 1764int-b.3Hordeum vulgare L. subsp. vulgare Skåne län, SwedenNSGCKERNELS1996DEVELOPED58.98333300, 13.50000000Genetic materialThe spike appears similar to the six-rowed spike, but developmental irregularities occur commonly in the lower portion of the spike. All lateral spikelets are reduced in size, and the lemma awn is short or reduced to a pointed tip. Commonly, only lateral spikelets in the middle of the spike set seed. Mutant plants have reduced vigor and tillering. Inheritance: Monofactorial recessive. Located in chromosome 5HL [7L], near the raw1 (smooth awn 1) locus based on linkage drag. Previous nomenclature and gene symbolization: None. Mutational events: int-b.3, -b.6 in Bonus; int-b.75 in Bonus. Mutant used for description and seed stocks: int-b.3 in Bonus (GSHO 1764); int-b.3 in Bowman (PI 483237)*7 (GSHO 2129).1006044GSHO 1764
66GSHO 1767int-f.19Hordeum vulgare L. subsp. vulgare Skåne län, SwedenNSGCKERNELS1996DEVELOPED58.98333300, 13.50000000Genetic materialThe spike appears to be six-rowed, but the lateral spikelets are much smaller, less than half the size of the central spikelets. Lateral spikelets are pointed and often have short awns. Seed set occurs in the lateral spikelets in the upper third of the spike. The base of the spike has shortened rachis internodes and appears erectoides-like. Inheritance: Monofactorial recessive. Location is unknown. Previous nomenclature and gene symbolization: None. Mutational events: int-f.19 in Foma. Mutant used for description and seed stocks: int-f.19 in Foma (GSHO 1767); int-f.19 in Bowman (PI 483237)*3 (GSHO 2270).1006080GSHO 1767
67GSHO 1768int-h.42Hordeum vulgare L. subsp. vulgare Skåne län, SwedenNSGCKERNELS1996DEVELOPED58.98333300, 13.50000000Genetic materialLateral spikelets are enlarged and have an inconspicuously pointed apex, but they do not set seed. Induced mutants show early heading and have an elongated basal internode. The spike appears lax with shortened rachis internodes at the base. The Bowman backcross-derived line is slightly shorter (5/6 normal) and produces extra spikelets (up to five fertile ones) at several rachis nodes in the lower half of the spike. Fusion of some spikelets results in double and occasionally triple kernels. Inheritance: Monofactorial recessive. Location is unknown. Previous nomenclature and gene symbolization: None. Mutational events: int-h.42, -h.43, -h.44 in Kristina; int-h.83 in Bonus (PI 189763). Mutant used for description and seed stocks: int-h.42 in Kristina (GSHO 1768); int-h.42 in Bowman (PI 483237)*5 (GSHO 2271).1006081GSHO 1768
68GSHO 1769int-i.39Hordeum vulgare L. subsp. vulgare Skåne län, SwedenNSGCKERNELS1996DEVELOPED58.98333300, 13.50000000Genetic materialLateral spikelets are enlarged and slightly pointed at the apex, but they fail to set seed. The tip of the spike may have very short rachis internodes and appears very dense or fasciated. The Bowman backcross-derived line exhitbits reduced plant height (5/6 normal), reduced spike length, and a distinct elongation of the basal rachis internode. Inheritance: Monofactorial recessive. Location is unknown. Previous nomenclature and gene symbolization: None. Mutational events: int-i.39 in Kristina. Mutant used for description and seed stocks: int-i.39 in Kristina (GSHO 1769); int-i.39 in Bowman (PI 483237)*4 (GSHO 2272).1006102GSHO 1769
69GSHO 1770int-k.47Hordeum vulgare L. subsp. vulgare Skåne län, SwedenNSGCKERNELSNot Available1996DEVELOPED58.98333300, 13.50000000Genetic materialThe spike is short and dense in the original mutant. Lateral spikelets are enlarged and the apex is pointed, and they occasionally have a short awn. Seed set does not occur in lateral spikelets and central spikelets are partially fertile. Plants of the original stock have a dense coating of surface waxes. The Bowman backcross-derived line has small, weak plants (about half normal height), which have short spikes (1/2 normal), reduced awn length (3/4 normal), and very poor seed set. Awns of plants in the derived line are semi-rough, but F1 hybrids with Bowman have semi-smooth awns. Inheritance: Monofactorial recessive. Location is unknown. Previous nomenclature and gene symbolization: None. Mutational events: int-k.47 in Kristina. Mutant used for description and seed stocks: int-k.47 in Krisitna (GSHO 1770); int-k.47 in Bowman (PI 483237)*5.1006103GSHO 1770
70GSHO 1772int-mHordeum vulgare L. subsp. vulgare Skåne län, SwedenNSGCKERNELS1996DEVELOPED58.98333300, 13.50000000Genetic materialThe spike is very short and has irregular rachis internode lengths. Lateral spikelets are enlarged and pointed, but they do not set seed. Spikelet density at the base of the spike is increased. Rachis internodes at the tip of the spike are very short, and the spike appears to have two or three fused or fasciated terminal spikelets. Tillering of int-m plants is increased and heading is slightly earlier. Inheritance: Monofactorial recessive. Location is unknown. Previous nomenclature and gene symbolization: None. Mutational events: int-m.85 in Bonus; int-m.la (GSHO 1772) in Lamont (PI 512036). Mutant used for description and seed stocks: int-m.85 in Bonus (GSHO 1771); int-m.85 in Bowman (PI 483237)*5 (GSHO 2273); int-m.la in Bowman (PI 483237)*4 (GSHO 2274).1006123GSHO 1772
71GSHO 1774dsp9.iHordeum vulgare L. subsp. vulgare JapanNSGCKERNELS1996DEVELOPEDGenetic materialExtremely dense spikes are easily distinguished from normal types when grown in the field or greenhouse. The spike often emerges from the side of the sheath. Yield and seed size are reduced somewhat in the Bowman backcross-derived line. Inheritance: Monofactorial recessive. Located in chromosome 6HL, about 27.2 cM distal from the xnt5 (xantha seedling 5) locus, and over 26.9 cM distal from the Aat2 (aspartate aminotransferase 2) locus. Previous nomenclature and gene symbolization: Dense spike la. Dense spike 9=l9. Mutational events: dsp9.i (OUM113) in Akashinriki, dsp9.j (OUM106), dsp9.k (OUM107), dsp9.l (OUM115), dsp9.m (OUM118) in Akashinriki. Mutant used for description and seed stocks: dsp9.i in Akashinriki (GSHO 1774); dsp9.i in Bowman (PI 483237)*7 (GSHO 2090).1006148GSHO 1774
72GSHO 1775lax-a.8Hordeum vulgare L. subsp. vulgare Skåne län, SwedenNSGCKERNELS1996DEVELOPED58.98333300, 13.50000000Genetic materialFlorets have five anthers with two developing from transformed lodicules. The grain is thin and angular with the caryopsis exposed between the lemma and palea. The awn has a very wide base, without a distinct notch in the lemma attachment region. Rachis internodes are 13% longer than normal. Tillers arise at oblique angles giving isolated plants an appearance of a tufty growth habit. Treatment of leaves after tillering with GA3 increased rachis internode length. Inheritance: Monofactorial recessive. Located in chromosome 5HL [7L], about 2.4 cM proximal from the ari-e (breviaristatum-e) locus, and about 3.1 cM from the ert-g (erectoides-g) locus. Previous nomenclature and gene symbolization: Laxatum-01=lax-01. Laxatum-a=lax-a01. Mutational events: lax-a.01, -a.4, -a.8, -a.20, -a.37, -a.39, -a.54 in Bonus; lax-a.92 in Bonus; lax-a.208, -a.218, -a.222, -a.229, -a.249, -a.256, -a.278, -a.286 in Foma (CIho 11333); a lax-a mutant (Mut 2100/61) in Proctor (PI 280420). Mutant used for description and seed stocks: lax-a.8 in Bonus (GSHO 1775); lax-a.8 in Bowman (PI 483237)*7 (GSHO 2103).1006167GSHO 1775
73GSHO 1776lax-b.1Hordeum vulgare L. subsp. vulgare Skåne län, SwedenNSGCKERNELS1996DEVELOPED58.98333300, 13.50000000Genetic materialPlants are weak and short (3/4 normal), spikes are lax and spikelet number is reduced. Seeds are small and thin and have a shrunken endosperm (xenia), which is detectable in the heterozygous condition. Treatment of leaves after tillering with GA3 increased rachis internode length. Seed set is reduced in the Bowman backcross-derived line. Inheritance: Monofactorial recessive. Located in chromosome 6HL, about 27.3 cM distal from the rob (orange lemma) locus, and about 18.8 cM proximal from the xnt5 (xantha seedling 5) locus. Previous nomenclature and gene symbolization: Laxatum-1=lax-1. Mutational events: lax-b.1 in Bonus. Mutant used for description and seed stocks: lax-b.1 in Bonus (GSHO 1776); lax-b.1 in Bowman (PI 483237)*7 (GSHO 2083).1006168GSHO 1776
74GSHO 1777lax-c.21Hordeum vulgare L. subsp. vulgare Skåne län, SwedenNSGCKERNELS1996DEVELOPED58.98333300, 13.50000000Genetic materialThe grain is thin and angular with the caryopsis exposed between the lemma and palea. The awn has a very wide base. Rachis internodes are about 7% longer than normal. The original stock in Bonus has reduced seed set in the tip of the spike, and the tiller number is reduced. Inheritance: Monofactorial recessive. Located in chromosome 6HL, over 27.3 cM distal from the lax-b (laxatum-b) locus. Previous nomenclature and gene symbolization: Laxatum-21=lax-21. Mutational events: lax-c.21, -c.23 in Bonus. Mutant used for description and seed stocks: lax-c.21 in Bonus (GSHO 1777); lax-c.21 in Bowman (PI 483237)*6 (GSHO 2086).1006186GSHO 1777
75GSHO 1784lys1.jHordeum vulgare L. subsp. vulgare North Dakota, United StatesNSGCKERNELS1996DONATED04/1996Genetic materialAfter the hard dough stage, kernels develop a depression in the center of the lemma which becomes progressively more distinct with maturity. The mature endosperm of the mutant appears to be harder than normal. The mutant expresses xenia permitting classification of kernels from heterozygous plants as normal or shrunken with an expected 3:1 ratio. Mutant plants have a higher lysine content in the endosperm than normal sibs. Field establishment and growth of mutant plants is normal, but grain yield is low. Grain size and hordein protein content are reduced by the lys1.j gene. Inheritance: Monofactorial recessive. Located in chromosome 5HL [7L], about 10.8 cM distal from the srh (short rachilla hair) locus. Previous nomenclature and gene symbolization: None. Mutational events: lys1.j in Hiproly. Mutant used for description and seed stocks: lys1.j in Hiproly (GSHO 1784); lys1.j in Bowman (PI 483237)*2 (GSHO 2115).1006248GSHO 1784
76GSHO 1785lys3.aHordeum vulgare L. subsp. vulgare DenmarkNSGCKERNELS1996DEVELOPEDGenetic materialAfter the hard dough stage, kernels develop a depression in the dorsal side or lemma which becomes progressively more distinct with maturity. The 1000-kernel weight of the mutant is about 75% of normal. The mutant expresses xenia permitting classification of kernels from heterozygous plants as normal or shrunken with an expected 3:1 ratio. Field establishment and growth of mutant plants is normal. Mutant kernels have a higher lysine content than normal kernels and the association with shrunken endosperm is pleiotropic. The endosperm is nearly devoid of all hordein polypeptides. A deficiency was caused by a linked factor for distorted male gamete transmission. The gamet lethal factor in Riso 1508 was named Gam3 (gametophyte factor 3) and is about 6.5 cM from the lys3 locus. Inheritance: Monofactorial recessive. Located in chromosome 5HL [7L], near the msg19 (male sterile genetic 19) locus, about 14.5 cM proximal from the fst1 (fragile stem 1) locus, and about 20.7 cM proximal from the ddt (reaction to DDT) locus. Previous nomenclature and gene symbolization: Shrunken endosperm xenia 3=sex3. Mutational events: lys3.a in Bomi (Riso 1508); lys3.b (Riso 18), lys3.c (Riso 19) in Bomi. Mutant used for description and seed stocks: lys3.a in Bomi (GSHO 1785).1006263GSHO 1785
77GSHO 1786lys6.iHordeum vulgare L. subsp. vulgare DenmarkNSGCKERNELS1996DEVELOPEDGenetic materialAt the hard dough stage, kernels homozygous for the lys6.i gene develop a depression near the center of the lemma, which becomes progressively more distinct as the kernels mature. Grain produced by mutant plants has a 12% increase in lysine content, and shrunken endosperm has a pleiotropic association with the high lysine gene lys6.i. Inheritance: Monofactorial recessive. Located in chromosome 6H. Previous nomenclature and gene symbolization: None. Mutational events: lys6.i in Bomi (Riso 527). Mutant used for description and seed stocks: lys6.i in Bomi (GSHO 1786); lys6.i in Bowman (PI 483237)*3(GSHO 2077).1006265GSHO 1786
78GSHO 1787lzd1.aHordeum vulgare L. subsp. vulgare JapanNSGCKERNELS1996DEVELOPEDGenetic materialPlants have a very procumbent growth habit prior to the boot stage. Plants are 3/4 normal height, maturity is delayed, and seedlings are very responsive to gibberellic acid. In the Bowman backcross-derived line, leaves are wider and shorter than in the recurrent parent. Inheritance: Monofactorial recessive. Located in chromosome 3HS, over 16.3 cM distal from the alm (albino lemma) locus. Previous nomenclature and gene symbolization: Dwarf 4=dw4. Mutational events: lzd1.a in Akashinriki (OUM005). Mutant used for description and seed stocks: lzd1.a in Akashinriki (GSHO 1787); lzd1.a in Bowman (PI 483237)* (GSHO 1938).1006278GSHO 1787
79GSHO 1788mat-b.7Hordeum vulgare L. subsp. vulgare Skåne län, SwedenNSGCKERNELS1996DEVELOPED58.98333300, 13.50000000Genetic materialEarly heading mutants at the mat-b locus show a rather drastic increase in earliness (heading 7 days earlier than their parental cultivars) under field cultivation in Sweden. When grown under controlled environmental conditions, mutants are photo- and thermoperiod sensitive and are classified as a pronounced long-day type. Early heading is also associated with decreased culm length, kernels per spike, and grain yield. They are generally more productive in field trials than mat-a (eam8) mutants, but less resistant to lodging, partially due to a modified culm structure. Compared with the original cultivars, they have one internode less per culm, a longer first internode, and a short basal internode. At an early stage of development, mat-b mutants are characterized by pale green color when grown at 15 to 20 degrees C, but they are bright green in color at lower temperatures. Inheritance: Monofactorial recessive. Location is unknown. Previous nomenclature and gene symbolization: Early 7=ea-b7. Early maturity-b=ea-b. Mutational events: mat-b.7, -b.10 (trans), -b.13 in Bonus; mat-b.34, -b.35, -b.38, -b.66 in Bonus, -b.106, -b.108, -b.134, -b.137, -b.145, -b.214, -b.216, -b.237, -b.244, -b.261, -b.279, -b.286, -b.313, -b.337, -b.403 in Foma (CIho 11333), -b.852, -b.853, -b.855, -b.859, -b.861, -b.867, -b.869, -b.870, -b.871, -b.904, -b.920, -b.922 in Bonus, -b.963, -b.977, -b.980, -b.986, -b.991, -b.992, -b.1008 in Sv 79353, -b.1082 in Sv Vg74233, -b.1103 in Sv Semira (NSL 206731), -b.1110, -b.1112, -b.1116 in Sv Frida; mat-b.1401 in Bonus; mat-b.1404, -b.1405 in Bonus. Mutant used for description and seed stocks: mat-b.7 in Bonus (GSHO 1788); mat-b.7 in Bowman (PI 483237)*3 (GSHO 2282).1006279GSHO 1788
80GSHO 1789mat-c.19Hordeum vulgare L. subsp. vulgare Skåne län, SwedenNSGCKERNELS1996DEVELOPED58.98333300, 13.50000000Genetic materialEarly heading mutants at the mat-c locus show a rather drastic increase in earliness (heading 7 days earlier than their parental cultivars) under field cultivation in Sweden. When grown under controlled environmental conditions, mutants are photo- and thermoperiod sensitive and are classified as a pronounced long-day type. Slightly later heading of some mat-c alleles has been reported. Early heading is also associated with decreased culm length, kernels per spike, and grain yield. Mutants have reduced culm length, due to a changed internode number, and the length of the first internode is increased. Spike length and spikelet number are markedly reduced. Inheritance: Monofactorial recessive. Location is unknown. Previous nomenclature and gene symbolization: Early 16=ea-c16. Early maturity-c=ea-c. Mutational events: mat-c.16 in Bonus; mat-c.19, -c.32, -c.93, -c.94 in Bonus, -c.101, -c.122, -c.400 in Foma (CIho 11333), -c.745 in Kristina, -c.758, -c.760, -c.770, -c.865, -c.881, -c.907, -c.910, -c.913, -c.926, -c.943 in Bonus, -c.966 in Sv 79353, -c.1091 in Sv Vg74233, -c.1096 in Sv Frida, -c.1102 in Sv Semira (NSL 206731), -c.1107, -c.1108, -c.1109, -c.1111, -c.1114, -c.1115 in Foma, -c.1118, -c.1120 in Sv Semira. Mutant used for description and seed stocks: mat-c.19 in Bonus (GSHO 1789); mat-c.19 in Bowman (PI 483237)*2 (GSHO 2283).1006294GSHO 1789
81GSHO 1790mat-d.14Hordeum vulgare L. subsp. vulgare Skåne län, SwedenNSGCKERNELS1996DEVELOPED58.98333300, 13.50000000Genetic materialEarly heading mutants at the mat-d locus have a moderate increase in earliness (heading 3 days earlier than their parental culitvars) under field cultivation in Sweden. When grown under controlled environmental conditions, mutants are photo- and thermoperiod sensitive and are classified as a pronounced long-day type. Early heading is also associated with increased culm length and grain yield. The length of the upper internode is increased and spike length is increased. Compared with the drastic maturity mutants at eam8 (mat-a), mat-b, and mat-c loci, mutants at the mat-d locus have an unchanged number of the culm internodes. Inheritance: Monofactorial incomplete dominant. Location is unknown. Previous nomenclature and gene symbolization: Early 14=ea-d14. Early maturity-d=ea-d. Mutational events: mat-d.14 in Bonus; mat-d.124 in Foma (CIho 11333). Mutant used for description and seed stocks: mat-d.14 in Bonus (GSHO 1790).1006296GSHO 1790
82GSHO 1791mat-e.18Hordeum vulgare L. subsp. vulgare Skåne län, SwedenNSGCKERNELS1996DEVELOPED58.98333300, 13.50000000Genetic materialEarly heading mutants at the mat-e locus show a rather drastic increase in earliness (heading 6 days earlier than their parental cultivars) under field cultivation in Sweden. When grown under controlled environmental conditions, the mutant mat-e.18 is photo- and thermoperiod insensitive. It has a pronounced short-day type of photoperiod response. A slight delay in heading and less short-day neutrality is associated with other alleles at this locus. Inheritance: Monofactorial recessive. Location is unknown. Previous nomenclature and gene symbolization: Praematurum-18=mat-18. Mutational events: mat-e.18 in Bonus; mat-e.55, -e.69, -e.98 in Bonus, -e.655 in Kristina, -e.854, -e.883, -e.890 in Bonus, -e.1093 in Sv Og74233. Mutant used for description and seed stocks: mat-e.18 in Bonus (GSHO 1791).1006309GSHO 1791
83GSHO 1792mat-f.23Hordeum vulgare L. subsp. vulgare Skåne län, SwedenNSGCKERNELS1996DEVELOPED58.98333300, 13.50000000Genetic materialEarly heading mutants at the mat-f locus have a moderate increase in earliness (heading 3 to 4 days earlier than their parental cultivars) under field cultivation in Sweden. When grown under controlled environmental conditions, mutants are photo- and thermoperiod sensitive and have a pronounced long-day response. Inheritance: Monofactorial recessive. Location is unknown. Previous nomenclature and gene symbolization: Praematurum-23=mat-23. Mutational events: mat-f.23, -f.780, -f.875, -f.891, -f.908, -f.932 in Bonus, -f.983 in Sv 79353. Mutant used for description and seed stocks: mat-f.23 in Bonus (GSHO 1792); mat-f.23 in Bowman (PI 483237)*2 (GSHO 2284).1006312GSHO 1792
84GSHO 1793mat-g.30Hordeum vulgare L. subsp. vulgare Skåne län, SwedenNSGCKERNELS1996DEVELOPED58.98333300, 13.50000000Genetic materialEarly heading mutants at the mat-g locus have a moderate increase in earliness (heading 3 to 4 days earlier than their parental cultivars) under field cultivation in Swede. When grown under controlled environmental conditions, mutants are photo- and thermoperiod sensitive and have a pronounced long-day response. Inheritance: Monofactorial recessive. Location is unknown. Previous nomenclature and gene symoblization: Praematurum-30=mat-30. Mutational events: mat-g.30, -g.873, -g.877 in Bonus, -g.985 in Sv 79353. Mutant used for description and seed stocks: mat-g.30 in Bonus (GSHO 1793); mat-g.30 in Bowman (PI 483237)*2 (GSHO 2285).1006326GSHO 1793
85GSHO 1794mat-h.36Hordeum vulgare L. subsp. vulgare Skåne län, SwedenNSGCKERNELS1996DEVELOPED58.98333300, 13.50000000Genetic materialEarly heading mutants at the mat-h locus show a drastic increase in earliness (heading 6 days earlier than their parental cultivars) under field cultivation in Sweden. When grown under controlled environmental conditions, mutants are photo- and thermoperiod sensitive and have a pronounced long-day response. The original mat-h.36 mutant stock also contains a narrow phenotype that is inherited in a recessive manner. Inheritance: Monofactorial recessive. Location is unknown. Previous nomenclature and gene symbolization: Praematurum-36=mat-36. Mutational events: mat-h.36, -h.935 in Bonus. Mutant used for description and seed stocks: mat-h.36 in Bonus (GSHO 1794); mat-h.36 in Bowman (PI 483237)*2 (GSHO 2286).1006327GSHO 1794
86GSHO 1795mat-i.37Hordeum vulgare L. subsp. vulgare Skåne län, SwedenNSGCKERNELS1996DEVELOPED58.98333300, 13.50000000Genetic materialEarly heading mutants at the mat-i locus have a drastic increase in earliness (heading 6 days earlier than their parental cultivars) under field cultivation in Sweden. When grown under controlled environmental conditions, mutants are photo- and thermoperiod sensitve and have a pronounced long-day response. Spikes of mutants are relatively short compared to their parental cultivars. Inheritance: Monofactorial recessive. Location is unknown. Previous nomenclature and gene symbolization: Praematurum-37=mat-37. Mutational events: mat-i.37, -i.76 in Bonus, -i.303 in Foma (CIho 11333), -i.742 in Kristina, -i.901, -i.914 in Bonus. Mutant used for description and seed stocks: mat-i.37 in Bonus (GSHO 1795); mat-i.37 in Bowman (PI 483237)*3 (GSHO 2287).1006340GSHO 1795
87GSHO 1810msg1.caHordeum vulgare L. subsp. vulgare California, United StatesNSGCKERNELSNot Available1996DEVELOPEDGenetic materialSelfing-none. Outcrossing-complete female fertility. Stamens-anthers smaller than fertile sib, no stomium or filament elongation. Pollen-microspores degenerate at or before the free microspore stage; non-staining, shrunken, and devoid of cytoplasm. Cytology-normal development and differentiation of anthers until completion of meiosis. Inheritance: Monofactorial recessive. Located in chromosome 1HL [5L], near the centromere, and about 10.0 cM proximal from the nec1 (necrotic leaf spot 1) locus. Previous nomenclature and gene symbolization: Male sterile=ms. Male sterile 1=ms1. Mutational events: msg1.ca in CIho 5368 (MSS005); msg1.i (MSS077) in 80TT25 (CIho 13638), msg1.t (MSS042) in Trophy (CIho 10647), msg1.ai (MSS100) in Betzes (PI 129430); msg1.ar (MSS310) in Glossy Brachytic (CIho 15246), msg1.bp (MSS330) in Betzes; msg1.cz (MSS348) in Betzes; msg1.gb (MSS429) in Maris Mink; msg1.jv (MSS527) in a Harrington outcross. Mutant used for description and seed stocks: msg1.ca in Betzes*11(GSHO 1810); msg1.ca in Bowman (PI 483237)*8 (GSHO 2042).1006455GSHO 1810
88GSHO 1811msg10.ayHordeum vulgare L. subsp. vulgare Montana, United StatesNSGCKERNELS1996DEVELOPEDGenetic materialSelfing-none. Outcrossing-complete female fertility. Stamens-anthers smaller than fertile sib, no stomium or filament elongation. Pollen-no staining with 2,3,5-triphenyltetrazolium, but staining with acetocarmine, some normal-appearing grains. Inheritance: Monofactorial recessive. Located in chromosome 7HS [1S], about 7.2 cM distal from the nud (naked caryopsis) locus, and about 14.1 cM from the lks2 (short awn 2) locus. Previous nomenclature and gene symbolization: Male sterile 3=ms3. Male sterile 10=ms10. Mutational events: msg10.ay in Compana (MSS058); msg10.cy (MSS059) in Manchuria*9 (CIho 2330). Mutant used for description and seed stocks: msg10.ay in Compana (GSHO 1811); msg10.ay in Bowman (PI 483237)*7 (GSHO 1835).1490036GSHO 1811
89GSHO 1812msg11.ckHordeum vulgare L. subsp. vulgare Manitoba, CanadaNSGCKERNELSNot Available1996DEVELOPEDGenetic materialSelfing-none. Outcrossing-complete female fertility. Stamens-anthers smaller than fertile sib, no stomium or filament elongation. Pollen-no staining in msg11.ck plants, but very low staining in msg11.az plants, no normal-appearing grains. Inheritance: Monofactorial recessive. Location is unknown, but a weak linkage to the raw1 (smooth awn 1) locus was reported. Previous nomenclature and gene symbolization: Male sterile 10=ms10. Male sterile 11=ms11. Male sterile 12=msg12. Mutational events: msg11.ck in Gateway (MSS060); msg11.az (MSS061) in Svalof 50-109 (CIho 10524). Mutant used for description and seed stocks: msg11.ck in Gateway (GSHO 1812); msg11.az in Bowman (PI 483237)*7 (GSHO 2299).1490037GSHO 1812
90GSHO 1813msg13.clHordeum vulgare L. subsp. vulgare Arizona, United StatesNSGCKERNELSNot Available1996DEVELOPEDGenetic materialSelfing-none. Outcrossing-complete female fertility. Stamens-anthers smaller than those of fertile sibs, no stomium or filament elongation. Pollen-no staining, shrunken, no normal-appearing grains. Inheritance: Monofactorial recessive. Location is unknown. Previous nomenclature and gene symbolization: Male sterile 13=ms13. Mutational events: msg13.cl in Haisa II (MSS062). Mutant used for description and seed stocks: msg13.cl in Haisa II (GSHO 1813); msg13.cl in Bowman (PI 483237)*7 (GSHO 2300).1490038GSHO 1813
91GSHO 1814msg14.cmHordeum vulgare L. subsp. vulgare Montana, United StatesNSGCKERNELSNot Available1996DEVELOPEDGenetic materialSelfing-none. Outcrossing-complete female fertility. Stamens-anthers smaller than fertile sib, no stomium or filament elongation. Pollen-less than 1% staining with 2,3,5-triphenyltetrazolium, about 9% with acetocarmine, some normal-appearing grains. Inheritance: Monofactorial recessive. Located in chromosome 7HS [1S], about 2.0 cM from the msg10 (male sterile genetic 10) locus. Previous nomenclature and gene symbolization: Male sterile 14=ms14. Mutational events: msg14.cm in Unitan (MSS063); msg14.da (MSS349) in Betzes (PI 129430); msg14.dl (MSS360) in Hector (CIho 15514). Mutant used for description and seed stocks: msg14.cm in Unitan (GSHO 1814); msg14.cm in Bowman (PI 483237)*7 (GSHO 1836).1490039GSHO 1814
92GSHO 1815msg15.cnHordeum vulgare L. subsp. vulgare California, United StatesNSGCKERNELSNot Available1996DEVELOPEDGenetic materialSelfing-early tillers have 7.7% and late tillers have 46.2% at Bozeman, Montana, USA; only fully fertile plants were recovered from the attempt to develop a Bowman backcross-derived line. Outcrossing-complete female fertility. Stamens-anthers smaller than fertile sibs. The phenotype of msg15.cn plants differ from other male sterile genetic mutants because late tillers are fertile. Anthesis seems to be delayed in relationship to spike emergence from the flag leaf. Inheritance: Monofactorial recessive. Location is unknown. Previous nomenclature and gene symbolization: Male sterile 15=ms15. Mutational events: msg15.cn in Atlas/2*Kindred (MSS064). Mutant used for description and seed stocks: msg15.cn in Atlas/2*Kindred (GSHO 1815).1490040GSHO 1815
93GSHO 1816msg16.coHordeum vulgare L. subsp. vulgare Montana, United StatesNSGCKERNELS1996DEVELOPEDGenetic materialSelfing-none. Outcrossing-complete female fertility. Stamens-anthers are almost equal in size to fertile sibs, stomium present, and filament elongation. Pollen-partial staining (15 to 35%) from both field and greenhouse grown plants, normal-appearing grains occur in some samples. Pollen appears immature based on staining with Fast Blue B, grains lack a pore or aperture, the exine is thinner, not distinctly two-layered, and bears fewer spicules than normal pollen. Tapetal tissue degeneration was observed at the free microspore stage. Inheritance: Monofactorial recessive. Located in chromosome 5HS [7S], about 8.0 to 15.0 cM distal from the T2-7a breakpoint. Previous nomenclature and gene symbolization: Male sterile 16=ms16. Mutational events: msg16.co in Betzes (MSS065); msg16.bi in Betzes (MSS323). Mutant used for description and seed stocks: msg16.co in Betzes (GSHO 1816); msg16.co in Bowman (PI 483237)*7 (GSHO 2116).1490041GSHO 1816
94GSHO 1817msg17.cpHordeum vulgare L. subsp. vulgare Montana, United StatesNSGCKERNELS1996DEVELOPEDGenetic materialSelfing-1% at Bozeman, Montana and 3% at Tucson, Arizona, USA. Outcrossing-complete female fertility. Stamens-anthers smaller than fertile sib, no stomium or filament elongation. Pollen-no staining with 2,3,5-triphenyltetrazolium, but staining with acetocarmine, some normal-appearing grains. Inheritance: Monofactorial recessive. Location is unknown. Previous nomenclature and gene symbolization: Male sterile 17=ms17. Mutational events: msg17.cp in Compana (MSS066). Mutant used for description and seed stocks: msg17.cp in Compana (GSHO 1817); msg17.cp in Bowman (PI 483237)*7 (GSHO 2301).1490042GSHO 1817
95GSHO 1818msg18.cqHordeum vulgare L. subsp. vulgare Montana, United StatesNSGCKERNELSNot Available1996DEVELOPEDGenetic materialSelfing-none. Outcrossing-complete female fertility. Stamens-anthers smaller than fertile sib, no stomium or filament elongation. Pollen-no staining with 2,3,5-triphenyltetrazolium, but staining with acetocarmine, some normal-appearing grains. Inheritance: Monofactorial recessive. Located in chromosome 5HL [7L], about 37.0 cM distal from the centromere. Previous nomenclature and gene symbolization: Male sterile 18=ms18. Mutational events: msg18.cq in Compana (MSS067); msg18.z (MSS093) in Betzes (PI 129430); msg18.am (MSS304) in Betzes. Mutant used for description and seed stocks: msg18.cq in Compana (GSHO 1818); msg18.cq in Bowman (PI 483237)*7 (GSHO 2117).1490045GSHO 1818
96GSHO 1819msg19.crHordeum vulgare L. subsp. vulgare Montana, United StatesNSGCKERNELS1996DEVELOPEDGenetic materialSelfing-none. Outcrossing-complete female fertility. Stamens-anthers smaller than fertile sib, no stomium or filament elongation. Pollen-no staining with 2,3,5-triphenyltetrazolium, but staining with acetocarmine, some normal-appearing grains. Inheritance: Monofactorial recessive. Located in chromosome 5HS [7S], between the centromere and the T2-7a breakpoint. Previous nomenclature and gene symbolization: Male sterile 19=ms19. Mutational events: msg19.cr in an introduction from Russia (MSS068). Mutant used for description and seed stocks: msg19.cr in an introduction from Russia (GSHO 1819); msg19.cr in Bowman (PI 483237)*7 (GSHO 2118).1490047GSHO 1819
97GSHO 2371msg2.cbHordeum vulgare L. subsp. vulgare Idaho, United StatesNSGCKERNELSNot Available1996DEVELOPEDGenetic materialSelfing-none. Outcrossing-complete female fertility. Stamens-anthers smaller than fertile sib, no stomium or filament elongation. Pollen-non-staining, shrunken, and no normal grains. Inheritance: Monofactorial recessive. Located in chromosome 2HL, about 2.4 cM proximal from the eog (elongated outer glume) locus. Previous nomenclature and gene symbolization: Male sterile 2=ms2. Male sterile 2=ms2. Mutational events: msg2.cb (MSS046) in F2 of Machuria/CIho 4363; msg2.ax (MSS045) in Compana (PI 539111); msg2.ed (MSS379) in Ingrid (CIho 10083). Mutant used for description and seed stocks: msg2.cb in Herta*10 (GSHO 2371), msg2.cb in Manchuria*19 (MSS047), msg2.cb in Ogalitsu*14 (MSS048), msg2.cb in Trebi*19 (MSS050); msg2.cb from Herta*10 in Bowman (PI 483237)*7 (GSHO 1890).1545167GSHO 2371
98GSHO 2372msg20.adHordeum vulgare L. subsp. vulgare Arizona, United StatesNSGCKERNELS1996DEVELOPEDGenetic materialSelfing-none. Outcrossing-complete female fertility. Stamens-anthers smaller than fertile sib. The attempt to develop a Bowman backcross-derived line was partially successful at best. Both the original stock and the derived line have short awns (3/4 normal), but the selfed seed set in the Bowman-backcross line is 50 to 75%. Inheritance: Monofactorial recessive. Located in chromosome 1H [5]. Previous nomenclature and gene symbolization: Male sterile ad=msg,,ad. Male sterile 20=ms20. Mutational events: msg20.ad in Hannchen (MSS096). Mutant used for description and seed stocks: msg20.ad in Hannchen (GSHO 2372); msg20.ad in Bowman (PI 483237)*4 (GSHO 2059).1545052GSHO 2372
99GSHO 2373msg21.qHordeum vulgare L. subsp. vulgare Idaho, United StatesNSGCKERNELS1996DEVELOPEDGenetic materialSelfing-3.8% at Bozeman, Montana and 2.9% at Tucson, Arizona, USA. Outcrossing-complete female fertility. Stamens-anthers almost equal in size to fertile sibs. The attempt to develop a Bowman backcross-derived line was partially successful. The presumed Bowman backcross-derived line has 70 to 85% selfed seed set. Inheritance: Monofactorial recessive. Location is unknown. Previous nomenclature and gene symbolization: Male sterile q=msg,,q. Male sterile 21=ms21. Mutational events: msg21.q in CIho 13640 (MSS085). Mutant used for description and seed stocks: msg21.q in CIho 13640 (GSHO 2373); msg21.q in Bowman (PI 483237)*7 (GSHO 2302).1545046GSHO 2373
100GSHO 2375msg23.bHordeum vulgare L. subsp. vulgare Montana, United StatesNSGCKERNELS1996DEVELOPEDGenetic materialSelfing-none in msg23.b and msg23.bg, but 1.9 to 13.6% in msg23.y. Outcrossing-complete female fertility. Stamens-anthers smaller than fertile sibs, no stomium or filament elongation. Inheritance: Monofactorial recessive. Located in chromosome 7HL [1L], not near the centromere. Previous nomenclature and gene symbolization: Male sterile b=msg,,b. Male sterile 23=ms23. Mutational events: msg23.b in Betzes (MSS071); msg23.y in Betzes (MSS092); msg23.bg in Betzes (MSS321). Mutant used for description and seed stocks: msg23.b in Betzes (GSHO 2375); msg23.b in Bowman (PI 483237)*7 (GSHO 1867).1545172GSHO 2375
101GSHO 2376msg24.vHordeum vulgare L. subsp. vulgare Montana, United StatesNSGCKERNELSNot Available1996DEVELOPEDGenetic materialSelfing-none. Outcrossing-complete female fertility. Stamens-anthers smaller than fertile sibs, no stomium or filament elongation. Inheritance: Monofactorial recessive. Located in chromosome 4HL, over 11.0 cM proximal from the Blx1 (non-blue aleurone xenia 1) locus. Previous nomenclature and gene symbolization: Male sterile v=msg,,v. Male sterile 24=ms24. Mutational events: msg24.j (MSS078) in Betzes/Domen (CIho 13639); msg24.v in Betzes (MSS089); msg24.ak in Betzes (MSS302); msg24.an in Betzes (MSS305); msg24.at (MSS312) in OAC21 (CIho 1470), msg24.bc in Betzes (MSS317); msg24.hg in an unknown cultivar (MSS460). Mutant used for description and seed stocks: msg24.v in Betzes (GSHO 2376); msg24.v in Bowman (PI 483237)*7 (GSHO 2018).1545047GSHO 2376
102GSHO 2379msg27.aeHordeum vulgare L. subsp. vulgare Colorado, United StatesNSGCKERNELS1996DEVELOPEDGenetic materialSelfing-none. Outcrossing-complete female fertility. Stamens-anthers rudimentary, no stomium or filament elongation. Inheritance: Monofactorial recessive. Located in chromosome 2HL, about 20.5 cM distal from the vrs1 (six-rowed spike 1) locus. Previous nomenclature and gene symbolization: Male sterile ae=msg,,ae. Mutational events: msg27.ae in Firlbecks III (MSS097); msg27.jr (MSS523) in Mona (PI 466726). Mutant used for description and seed stocks: msg27.ae in Firlbecks III (GSHO 2379); msg27.ae in Bowman (PI 483237)*7 (GSHO 1921).1545140GSHO 2379
103GSHO 2380msg28.asHordeum vulgare L. subsp. vulgare Ontario, CanadaNSGCKERNELSNot Available1996DEVELOPEDGenetic materialSelfing-none. Outcrossing-complete female fertility. Stamens-anthers rudimentary, with no stomium or filament elongation. Pollen-non-stainig, shrunken, and no normal-appearing grains. Cytology-normal meiosis. Inheritance: Monofactorial recessive. Located in chromosome 6H, near the rob (orange lemma) locus. Previous nomenclature and gene symbolization: Male sterile as=msg,,as. Mutational events: msg28.as in York (MSS311). Mutant used for description and seed stocks: msg28.as in York (GSHO 2380); msg28.as in Bowman (PI 483237)*7 (GSHO 2079).1545044GSHO 2380
104GSHO 2381msg29.aHordeum vulgare L. subsp. vulgare Montana, United StatesNSGCKERNELS1996DEVELOPEDGenetic materialSelfing-none. Outcrossing-complete female fertility. Stamens-anthers smaller than fertile sibs, with no stomium or filament elongation. Inheritance: Monofactorial recessive. Located in chromosome 5HL [7L], about 22.7 cM from the raw1 (smooth awn 1) locus. Previous nomenclature and gene symbolization: Male sterile a=msg,,a. Male sterile=63msx1 and 63msx2 (ms-aa). Mutational events: msg29.a in Ackermans MGZ (MSS069); msg29.aa in Ackermans MGZ (MSS070). Mutant used for description and seed stocks: msg29.a in Ackermans MGZ (GSHO 2381); msg29.a in Bowman (PI 483237)*7 (GSHO 2140).1545127GSHO 2381
105GSHO 2382msg30.cHordeum vulgare L. subsp. vulgare Montana, United StatesNSGCKERNELSNot Available1996DEVELOPEDGenetic materialSelfing-0.7% at Bozeman, Montana, USA. Outcrossing-complete female fertility, open pollinated seed set of 15% at Bozeman, MT and 2% at Elimaki, Finland. Stamens-anthers smaller than fertile sibs, stomium present and the filament elongates. Inheritance: Monofactorial recessive. Located in chromosome 7HL [1L], about 11.4 cM from the lks2 (short awn 2) locus. Previous nomenclature and gene symbolization: Male sterile c=msg,,c. Male sterile=msx2 (msa). Mutational events: msg30.c in Compana (MSS072). Mutant used for description and seed stocks: msg30.c in Compana (GSHO 2382); msg30.c in Bowman (PI 483237)*7 (GSHO 1859).1545139GSHO 2382
106GSHO 2383msg31.dHordeum vulgare L. subsp. vulgare Idaho, United StatesNSGCKERNELSNot Available1996DEVELOPEDGenetic materialSelfing-none. Outcrossing-complete female fertility. Stamens-anthers rudimentary, no stomium or filament elongation. In addition to male sterility, mutant plants are weak and short (about 1/2 normal height), show delayed development, and appear to l lack surface wax on the spike (wax code - ++ ++). Inheritance: Monofactorial recessive. Located in chromosome 1HS [5S], probably proximal from the Cer-yy (eceriferum-yy) locus and distal from the sls (small lateral spikelet) locus. Previous nomenclature and gene symbolization: Male sterile d=msg,,d. Male sterile dwarf=msdwf. Mutational events: msg31.d in 51Ab4934 (MSS306). Mutant used for description and seed stocks: msg31.d in 51Ab4934 (GSHO 2383); msg31.d in Bowman (PI 483237)*7 (GSHO 2031).1545159GSHO 2383
107GSHO 2384msg32.wHordeum vulgare L. subsp. vulgare Montana, United StatesNSGCKERNELSNot Available1996DEVELOPEDGenetic materialSelfing-none. Outcrossing-complete female fertility. Stamens-anthers smaller than fertile sibs, no stomium or filament elongation. Inheritance: Monofactorial recessive. Located in chromosome 7H [1], linked to the nud (naked caryopsis) locus. Previous nomenclature and gene symbolization: Male sterile w=msg,,w. Male sterile=63msx7. Mutational events: msg32.w in Betzes (MSS090). Mutant used for description and seed stocks: msg32.w in Betzes (GSHO 2384); msg32.w in Bowman (PI 483237)*7 (GSHO 1860).1545149GSHO 2384
108GSHO 2385msg33.xHordeum vulgare L. subsp. vulgare Montana, United StatesNSGCKERNELSNot Available1996DEVELOPEDGenetic materialSelfing-20% at Bozeman, Montana, 17% at Tucson, Arizona, USA, and 0% at Elimaki, Finland. Outcrossing-complete female fertility. Stamens-anthers smaller than fertile sibs, stomium present, and filament elongation. Inheritance: Monofactorial recessive. Located in chromosome 2HS, about 5.2 cM distal from the centromere, and 25.6 cM proximal from the vrs1 (six-rowed spike 1) locus. Previous nomenclature and gene symbolization: Male sterile x=msg,,x. Male sterile=63msx8. Mutational events: msg33.x in Betzes (MSS091). Mutant used for seed description and seed stocks: msg33.x in Betzes (GSHO 2385); msg33.x in Bowman (PI 483237)*7 (GSHO 1884).1545098GSHO 2385
109GSHO 2386msg34.avHordeum vulgare L. subsp. vulgare Montana, United StatesNSGCKERNELS1996DEVELOPEDGenetic materialSelfing-none. Outcrossing-complete female fertility. Stamens-anthers rudimentary, with no stomium or filament elongation. Heterozygotes show reduced fertility (75 to 85% seed set), and male sterile plants always have a long glume awn (Lga) trait from Paragon. Thus, the msg34.av allele probably involves a translocation between chromosome 6HS and an unknown chromosome. Inheritance: Monofactorial recessive. Located in chromosome 6H, recombination with the rob (orange lemma) locus was not observed. Previous nomenclature and gene symbolization: Male sterile av=msg,,av. Male sterile=63msy1. Mutational events: msg34.av in Paragon (MSS314). Mutant used for description and seed stocks: msg34.av in Paragon (GSHO 2386); msg34.av in Bowman (PI 483237)* 7 (GSHO 2070).1545045GSHO 2386
110GSHO 2387msg35.drHordeum vulgare L. subsp. vulgare Montana, United StatesNSGCKERNELSNot Available1996DEVELOPEDGenetic materialSelfing-none. Outcrossing-complete female fertility. Stamens-anthers smaller than fertile sib, with no stomium of filament elongation. Inheritance: Monofactorial recessive. Located in chromosome 2HL, about 23.0 cM from the wst7 (white streak 7) locus. Previous nomenclature and gene symbolization: Male sterile genetic,,dr=msg,,dr. Male sterile=76Y17. Mutational events: msg35.dr in Karl (MSS366). Mutant used for description and seed stocks: msg35.dr in Karl (GSHO 2387); msg35.dr in Bowman (PI 483237)*7 (GSHO 1933).1545116GSHO 2387
111GSHO 2388msg36.bkHordeum vulgare L. subsp. vulgare Montana, United StatesNSGCKERNELSNot Available1996DEVELOPEDGenetic materialSelfing-about 1.1%. Outcrossing-complete female fertility. Stamens-anthers nearly normal sized with stomium and filament elongation. Inheritance: Monofactorial recessive. Located in chromosome 6HS, about 10.8 cM distal from the rob (orange lemma) locus. Previous nomenclature and gene symbolization: Male sterile bk=msg,,bk. Male sterile=867N-89. Mutational events: msg36.bk in Betzes (MSS325); msg36.eg in Pavo P57 (MSS382). Mutant used for description and seed stocks: msg36.bk in Betzes (GSHO 2388); msg36.bk in Bowman (PI 483237)*7 (GSHO 2067).1545114GSHO 2388
112GSHO 2389msg37.hlHordeum vulgare L. subsp. vulgare Ceredigion, United KingdomNSGCKERNELS1996DEVELOPED52.25000000, -4.00000000Genetic materialSelfing-none to 6% or more, the degree of selfing seems sensitive to environmental conditions, favored by cool conditions. Outcrossing-complete female fertility. Stamens-anthers shrunken, no stomium or filament elongation to nearly normal. Inheritance: Monofactorial recessive. Location is unknown, not associated with chromosome 2H. Previous nomenclature and gene symbolization: Male sterile hl=msg,,hl. Mutational events: msg37.hl in Clermont (MSS465); msg37.jx (MSS529) in a dwarf mutant 17:17:2 (DWS1009) selected from Birgitta (NSGC 1870). Mutant used for description and seed stocks: msg37.hl in Clermont (GSHO 2389); msg37.jx in Bowman (PI 483237)*7 (GSHO 2303).1545148GSHO 2389
113GSHO 2390msg38.jlHordeum vulgare L. subsp. vulgare Skåne län, SwedenNSGCKERNELSNot Available1996DEVELOPED58.98333300, 13.50000000Genetic materialSelfing-none. Outcrossing-complete female fertility. Stamens-anthers nearly normal in size, but without stomium, filament elongates. Inheritance: Monofactorial recessive. Location is unknown. Previous nomenclature and gene symbolization: Male sterile jl=msg,,jl. Mutational events: msg38.jl in Ingrid (MSS517). Mutant used for description and seed stocks: msg38.jl in Ingrid (GSHO 2390); msg38.jl in Bowman (PI 483237)*7 (GSHO 2304).1545039GSHO 2390
114GSHO 2391msg39.dmHordeum vulgare L. subsp. vulgare FinlandNSGCKERNELSNot Available1996DEVELOPEDGenetic materialSelfing-about 1% in Bozeman, Montana, USA and Elimaki, Finland. Outcrossing-complete female fertility. Stamens-anthers are the same size as fertile sibs, stomium present, filament elongates. Inheritance: Monofactorial recessive. Located in chromosome 6H, over 33.9 cM from the rob (orange lemma) locus. Previous nomenclature and gene symbolization: Male sterile dm=msg,,dm. Mutational events: msg39.dm in P11 (MSS361); msg39.dn (MSS362) in a Finnish six-rowed barley (H31, CIho15837); msg39.ff (MSS407) in Sabarlis (CIho 15484). Mutant used for description and seed stocks: msg39.dm in P11 (GSHO 2391); msg39.dm in Bowman (PI 483237)*7 (GSHO 2080).1545040GSHO 2391
115GSHO 2392msg4.cdHordeum vulgare L. subsp. vulgare Montana, United StatesNSGCKERNELSNot Available1996DEVELOPEDGenetic materialSelfing-none. Outcrossing-complete female fertility. Stamens-anthers smaller than fertile sib, no stomium or filament elongation. Pollen-non-staining, shrunken, and no normal grains. Inheritance: Monofactorial recessive. Located in chromosome 1H, near the centromere. Previous nomenclature and gene symbolization: Male sterile 4=ms4. Mutational events: msg4.cd in Freja (MSS052). Mutant used for description and seed stocks: msg4.cd in Freja (GSHO 2392); msg4.cd in Bowman (PI 483237)*7 (GSHO 2043).1545117GSHO 2392
116GSHO 2393msg40.acHordeum vulgare L. subsp. vulgare Manitoba, CanadaNSGCKERNELSNot Available1996DEVELOPEDGenetic materialSelfing-none. Outcrossing-less than 30% in crosses and in the Bowman backcross-derived line. Stamens-anthers are smaller than those of fertile sibs, no stomium or filament elongation. Inheritance: Monofactorial recessive. Located in chromosome 6H, over 27.8 cM from the rob (orange lemma) locus. Previous nomenclature and gene symbolization: Male sterile ac=msg,,ac. Male sterile=ms-C. Mutational events: msg40.ac in Conquest (MSS095). Mutant used for description and seed stocks: msg40.ac in Bowman (PI 483237)*6 (GSHO 2081).1545041GSHO 2393
117GSHO 2394msg41.ajHordeum vulgare L. subsp. vulgare Montana, United StatesNSGCKERNELSNot Available1996DEVELOPEDGenetic materialSelfing-none. Outcrossing-complete female fertility, the Bowman backcross-derived line frequently has less than 20% seed set as a female parent in crosses. Stamens-anthers rudimentary, no stomium or filament elongation. Inheritance: Monofactorial recessive. Location is unknown. Previous nomenclature and gene symbolization: Male sterile aj=msg,,aj. Male sterile=65msx166. Mutational events: msg41.aj in Betzes (MSS101); msg41.dk in Betzes (MSS359); msg41.do (MSS363) in Maris Baldric (PI 294512); msg41.ef (MSS381) in Hector (CIho 15514), msg41.el (MSS387) in Sabarlis (CIho 15484), msg41.eq in Sel 12387Co (MSS392), msg41.fa (MSS402) in Midas (PI 343078), msg41.gl (MSS439) in Maris Mink (PI 467824), msg41.ij (MSS489) in Mazurka (PI 410868). Mutant used for description and seed stocks: msg41.aj in Betzes (GSHO 2394); msg41.dk in Bowman (PI 483237)*6 (GSHO 2305).1545042GSHO 2394
118GSHO 2395msg42.dbHordeum vulgare L. subsp. vulgare Montana, United StatesNSGCKERNELS1996DEVELOPEDGenetic materialSelfing-none. Outcrossing-complete female fertility. Stamens-anthers rudimentary, no stomium or filament elongation. Inheritance: Monofactorial recessive. Located in chromosome 3H, over 18.9 cM from the alm (albino lemma) locus. Previous nomenclature and gene symbolization: Male sterile db=msg,,db. Male sterile=B68-N-109. Mutational events: msg42.db in Betzes (MSS350); msg42.gt (MSS447) in Berac (PI 355136), msg42.hw in Sel 15025Co (MSS476), msg42.iy in Sv73608 (MSS504). Mutant used for description and seed stocks: msg42.db in Betzes (GSHO 2395); msg42.db in Bowman (PI 483237)*6 (GSHO 1948).1545043GSHO 2395
119GSHO 2396msg43.brHordeum vulgare L. subsp. vulgare Montana, United StatesNSGCKERNELSNot Available1996DEVELOPEDGenetic materialSelfing-none. Outcrossing-complete female fertility, the Bowman backcross-derived line has less than 30% seed set in crosses. Stamens-anthers rudimentary, no stomium or filament elongation. Inheritance: Monofactorial recessive. Location is unknown. Previous nomenclature and gene symbolization: Male sterile br=msg,,br. Male sterile=B65msx38. Mutational events: msg43.br in Betzes (MSS332). Mutant used for description and seed stocks: msg43.br in Betzes (GSHO 2396); msg43.br in Bowman (PI 483237)* 7 (GSHO 2306).1545097GSHO 2396
120GSHO 2397msg44.cxHordeum vulgare L. subsp. vulgare FinlandNSGCKERNELS1996DEVELOPEDGenetic materialSelfing-none. Outcrossing-complete female fertility. Stamens-anthers rudimentary, no stomium or filament elongation. Inheritance: Monofactorial recessive. Location is unknown. Previous nomenclature and gene symbolization: Male sterile cx=msg,,cx. Mutational events: msg44.cx in HA6-33-02 (MSS346). Mutant used for description and seed stocks: msg44.cx in HA6-33-02 (GSHO 2397); msg44.cx in Bowman (PI 483237)*7 (GSHO 2307).1545115GSHO 2397
121GSHO 2398msg45.dpHordeum vulgare L. subsp. vulgare Montana, United StatesNSGCKERNELSNot Available1996DEVELOPEDGenetic materialSelfing-none. Outcrossing-complete female fertility. Stamens-anthers rudimentary, no stomium or filament elongation. Inheritance: Monofactorial recessive. Location is unknown. Previous nomenclature and gene symbolization: Male sterile dp=msg,,dp. Male sterile=76Y2. Mutational events: msg45.dp in RPB439-71 (MSS364). Mutant used for description and seed stocks: msg45.dp in RPB439-71 (GSHO 2398); msg45.dp in Bowman (PI 483237)*4 (GSHO 2308).1545113GSHO 2398
122GSHO 2399msg46.ecHordeum vulgare L. subsp. vulgare Montana, United StatesNSGCKERNELS1996DEVELOPEDGenetic materialSelfing-none. Outcrossing-complete female fertility. Stamens-anthers smaller than those of fertile sibs, no stomium or filament elongation. Inheritance: Monofactorial recessive. Location is unknown. Previous nomenclature and gene symbolization: Male sterile ec=msg,,ec. Mutational events: msg46.ec in Hector (MSS378). Mutant used for description and seed stocks: msg46.ec in Hector (GSHO 2399); msg46.ec in Bowman (PI 483237)*4 (GSHO 2309).1545096GSHO 2399
123GSHO 2400msg47.epHordeum vulgare L. subsp. vulgare Ceredigion, United KingdomNSGCKERNELS1996DEVELOPED52.25000000, -4.00000000Genetic materialSelfing-none. Outcrossing-complete female fertility. Stamens-anthers rudimentary, no stomium or filament elongation. Inheritance: Monofactorial recessive. Location is unknown. Previous nomenclature and gene symbolization: Male sterile ep=msg,,ep. Mutational events: msg47.ep in Sel 12384Co (MSS391). Mutant used for description and seed stocks: msg47.ep in Sel 12384Co (GSHO 2400); msg47.ep in Bowman (PI 483237)*5 (GSHO 2310).1545181GSHO 2400
124GSHO 2401msg48.jtHordeum vulgare L. subsp. vulgare Skåne län, SwedenNSGCKERNELSNot Available1996DEVELOPED58.98333300, 13.50000000Genetic materialSelfing-5%, but it may be near 50% in certain environments. Outcrossing-Complete female fertility. Stamens-anthers slightly smaller than fertile sib with filament elongation, but no stomium. Inheritance: Monofactorial recessive. Located in chromosome 2H, more than 31.9 cM from the vrs1 (six-rowed spike 1) locus. Previous nomenclature and gene symbolization: Male sterile genetic jt=msg,,jt and Msg76-5. Mutational events: msg48.jt in Simba (MSS525). Mutant used for description and seed stocks: msg48.jt in Simba (GSHO 2401); msg48.jt in Bowman (PI 483237)*4 (GSHO 1925).1545138GSHO 2401
125GSHO 2402msg49.jwHordeum vulgare L. subsp. vulgare North Dakota, United StatesNSGCKERNELS1996DEVELOPEDGenetic materialSelfing-none. Outcrossing-Complete female fertility. Stamens-anthers rudimentary, no stomium or filament elongation. Inheritance: Monofactorial recessive. Located in chromosome 5HL [7L], about 10.4 cM from the raw1 (smooth awn 1) locus. Previous nomenclature and gene symbolization: Male sterile genetic jw=msg,,jw. Mutational events: msg49.jw in ND7369 (MSS528). Mutant used for description and seed stocks: msg49.jw in ND7369 (GSHO 2402); msg49.jw in Bowman (PI 483237)*8 (GSHO 2141).1545173GSHO 2402
126GSHO 2403msg5.ceHordeum vulgare L. subsp. vulgare Montana, United StatesNSGCKERNELSNot Available1996DEVELOPEDGenetic materialSelfing-none. Outcrossing-complete female fertility. Stamens-anthers smaller than fertile sib, no stomium or filament elongation. Pollen-non-staining, some normal-appearing grains. Inheritance: Monofactorial recessive. Located in chromosome 3HS, about 13.1 cM proximal from the uzu (uzu) locus, and about 6.8 cM proximal from the alm (albino lemma) locus.Previous nomenclature and gene symbolization: Male sterile 5=ms5. Mutational events: msg5.ce in Carlsberg II (MSS053); msg5.s (MSS087) in Schweigers Erika (CIho 11501); msg5.fr (MSS419) in Midas (PI 343078). Mutant used for description and seed stocks: msg5.ce in Carlsberg II (GSHO 2403); msg5.ce in Bowman (PI 483237)*7 (GSHO 1954).1545050GSHO 2403
127GSHO 2404msg50.hmHordeum vulgare L. subsp. vulgare Ceredigion, United KingdomNSGCKERNELSNot Available1996DEVELOPED52.25000000, -4.00000000Genetic materialSelfing-0% reported, but occasionally 5 to 10% selfed seed set is observed. Outcrossing-complete female fertility. Stamens-anthers slightly smaller than fertile sib with filament elongation and stomium. Inheritance: Monofactorial recessive. Located in chromosome 7HL [1L], about 13.6 cM from the lks2 (short awn 2) locus. Previous nomenclature and gene symbolization: Male sterile genetic hm=msg,,hm. Mutational events: msg50.gh in Maris Mink (MSS435); msg50.hm (MSS466) in Berac (PI 355136). Mutant used for description and seed stocks: msg50.hm in Berac (GSHO 2404); msg50.hm in Bowman (PI 483237)*7 (GSHO 1861).1545165GSHO 2404
128GSHO 2405msg6.cfHordeum vulgare L. subsp. vulgare Montana, United StatesNSGCKERNELSNot Available1996DEVELOPEDGenetic materialSelfing-none, but occasionally a few selfed seeds occur. Outcrossing-complete female fertility. Stamens-equal in size to normal anthers; stomium present, filament elongation. Pollen-nearly normal stained in both field and greenhouse plants, most grains appear normal; grains are non-functional because aperture development is abnormal. Cytology-normal development and differentiation of anthers. Inheritance: Monofactorial recessive. Located in chromosome 6HS, near the centromere, and about 1.0 cM distal from the rob (orange lemma) locus. Previous nomenclature and gene symbolization: Male sterile 6=ms6. Mutational events: msg6.cf in Heines Hanna (MSS054). Mutant used for description and seed stocks: msg6.cf in Heines Hanna (GSHO 2405); msg6.cf in Bowman (PI 483237)*7 (GSHO 2078).1545150GSHO 2405
129GSHO 2406msg7.cgHordeum vulgare L. subsp. vulgare Montana, United StatesNSGCKERNELS1996DEVELOPEDGenetic materialSelfing-none. Outcrossing-complete female fertility. Stamens-anthers much smaller than fertile sibs, no stomium or filament elongation. Pollen-microspores degenerate before the free microspore stage, non-staining, no free grains. Cytology-normal development and differentiation of anthers until completion of meiosis, but degeneration of tapetal tissue at the tetrad stage. Inheritance: Monofactorial recessive. Located in chromosome 5HL [7L], about 5.5 cM from the raw1 (smooth awn 1) locus. Previous nomenclature and gene symbolization: Male sterile 7=ms7. Mutational events: msg7.cg in Dekap (MSS055); msg7.ah (MSS099) in HB 421/78 (CIho 13641); msg7.fx (MSS425) in Proctor. Mutant used for description and seed stocks: msg7.cg in Dekap (GSHO 2406); msg7.cg in Bowman (PI 483237)*7 (GSHO 2109).1545128GSHO 2406
130GSHO 2407msg8.chHordeum vulgare L. subsp. vulgare Montana, United StatesNSGCKERNELS1996DEVELOPEDGenetic materialSelfing-none. Outcrossing-complete female fertility. Stamens-anthers smaller than fertile sibs, stomium present, and filament elongation. Pollen-reduced staining (6.9% stains with 2,3,5-triphenyltetrazolium, 20.7% stains with acetocarmine), some grains appear normal. Cytology-normal development and differentiation of microspores up to the free microspore stage. Grains are undeveloped, have a stainable ring at the apertural annulus without any actual pore; the exine staining with Fast Blue B is darker than in normal grains. Inheritance: Monofactorial recessive. Located in chromosome 5HL [7L], about 8.3 cM from the raw1 (smooth awn 1) locus. Previous nomenclature and gene symbolization: Male sterile 8=ms8. Mutational events: msg8.ch in Betzes (MSS056); msg8.au (MSS313) in OAC 21 (CIho 1470). Mutant used for description and seed stocks: msg8.ch in Betzes (GSHO 2407); msg8.ch in Bowman (PI 483237)*6 (GSHO 2110).1545099GSHO 2407
131GSHO 2408msg9.ciHordeum vulgare L. subsp. vulgare Montana, United StatesNSGCKERNELSNot Available1996DEVELOPEDGenetic materialSelfing-10% at Bozeman, Montana and 24% at Tucson, Arizona, USA. Outcrossing-complete female fertility. Stamens-anthers smaller than fertile sib, stomium present and filament elongation occurs. Pollen-stained in plants grown in the field and in the greenhouse, in some samples the grains appear normal. Inheritance: Monofactorial recessive. Located in chromosome 2HS, over 15.1 cM distal from the eog (elongated outer glume) locus. Previous nomenclature and gene symbolization: Male sterile 9=ms9. Mutational events: msg9.ci in Vantage (MSS057). Mutant used for description and seed stocks: msg9.ci in Vantage (GSHO 2408); msg9.ci in Bowman (PI 483237)*7 (GSHO 1883).1545051GSHO 2408
132GSHO 2410mtt5.fHordeum vulgare L. subsp. vulgare Japan SEEDHistoric1996DEVELOPEDGenetic materialMottled bands occur on the leaf blade and sheath and are retained until heading. The first few seedling leaves often do not show the mottled trait. Plants are much less vigorous than normal sibs. Inheritance: Monofactorial recessive. Located in chromosome 6HL, about 6.4 cM distal from the rob (orange lemma) locus. Previous nomenclature and gene symbolization: Mottled f=mtt,,f. Mutational events: mtt5.f in Akashinriki (OUM022). Mutant used for description and seed stocks: mtt5.f in Akashinriki (GSHO 2410); mtt5.f in Bowman (PI 483237)*6 (GSHO 2084).1545160GSHO 2410
133GSHO 2437raw6.fHordeum vulgare L. subsp. vulgare North Dakota, United StatesNSGCKERNELS1996DEVELOPEDGenetic materialPlants have very fewer stigma hairs and seed set is greatly reduced. In the Bowman backcross-derived line, which contains the raw6.f and raw1.a genes, plants have very smooth awns and very few stigma hairs, and seed set is commonly less than 5%. Phenotypically, raw6.f plants closely resemble those produced by the raw5.e (smooth awn 5) mutant in chromosome 6H. A recombinant between the raw6.f and raw1.a alleles was not observed until several backcrosses to Bowman were completed. Inheritance: Monofactorial recessive. Located in chromosome 5HL, near the raw1 (smooth awn 1) locus based on linkage drag. Previous nomenclature and gene symbolization: Smooth awn 6=r6. Mutational events: raw6.f in Glenn. Mutant used for description and seed stocks: raw6.f in Glenn (GSHO 2437); raw6.f in Bowman (PI 483237)*8 (GSHO 2134).1545100GSHO 2437
134GSHO 2438rpg4.dHordeum vulgare L. subsp. vulgare North Dakota, United StatesNSGCKERNELS1996DONATED04/1996Genetic materialThe rpg4.d gene confers low infection types to pathotypes QCC and MCC of the wheat stem rust pathogen (Puccinia graminis f sp tritici) at the seedling stage under low incubation temperatures 18 to 22 degrees C. This gene is ineffective (exhibits compatible infection types) against some pathotypes at incubation temperatures greater than 27 degrees C. At the adult plant stage, the rpg4.d allele confers a moderate to high level of resistance to pathotype QCC. The rpg4.d gene confers a dominant resistant reaction to Puccinia graminis f sp secalis. Inheritance: Monofactorial recessive. Located in chromosome 5HL [7L], in central part of 5HL about 4.9 cM distal from RFLP marker MWG740. Previous nomenclature and gene symbolization: None. Mutational events: rpg4.d in Q21861. Mutant used for description and seed stocks: rpg4.d in Q21861 (GSHO 2438).1545054GSHO 2438
135GSHO 2439sca1.aHordeum vulgare L. subsp. vulgare JapanNSGCKERNELS1996DEVELOPEDGenetic materialIn the original stock, awns are reduced to a length of only 2 cm and are curved outward at the tip. Awn tips may have a stigma-like appearance in the original stock. In the Bowman backcross-derived line, awns are short (less than 1/4 normal), but awn tips are not curved or stigma-like. Inheritance: Monofactorial recessive. Located in chromosome 3HS, about 1.4 cM distal from the alm (albino lemma) locus. Previous nomenclature and gene symbolization: None. Mutational events: sca1.a in Akashinriki (OUM206). Mutant used for description and seed stocks: sca1.a in Akashinriki (GSHO 2439), sca1.a in Bowman (PI 483237)*6 (GSHO 1951).1545059GSHO 2439
136GSHO 2466sdw2.bHordeum vulgare L. subsp. vulgare PolandNSGCKERNELS1996DEVELOPEDGenetic materialPlants are about 3/4 normal height; cullms are thin with narrow, short, erect leaves. The flag leaf is narrow and short. The peduncle is short, the collar has a small leaf-like bract, and the basal rachis internode is elongated. Inheritance: Monofactorial recessive. Located in chromosome 3HL, over 34.5 cM distal from the sld1 (slender dwarf 1) locus. Previous nomenclature and gene symbolization: Semidwarf-b=sdw-b. Mutational events: sdw2.b in Mg4170 (267MK, later called 437MK). Mutant used for description and seed stocks: sdw2.b in Mg4170 (GSHO 2466); sdw2.b in Bowman (PI 483237)*7 (GSHO 1965).1545061GSHO 2466
137GSHO 2467seg6.fHordeum vulgare L. subsp. vulgare Arizona, United StatesNSGCKERNELSNot Available1996DEVELOPEDGenetic materialSeed size is reduced, but the degree reduction is affected by environment. Seed weights of 25, 50, and 75% of normal were obtained from plants grown in the field in Arizona, in the field in Montana, and in the greenhouse in Arizona, USA, respectively. Pollen mother cell meiosis and pollen fertility are normal. Seed from seg6.g plants can be used to establish stands under field conditions. Inheritance: Monofactorial recessive. Located in chromosome 3HL. Previous nomenclature and gene symbolization: Shrunken endosperm=se6. Mutational events: seg6.f in an unknown hybrid, seg6.g in Ingrid. Mutant used for description and seed stocks: seg6.g in Ingrid (GSHO 2467); seg6.g in Bowman (PI 483237)*3 (GSHO 1975).1545048GSHO 2467
138GSHO 2468seg7.hHordeum vulgare L. subsp. vulgare Arizona, United StatesNSGCKERNELS1996DEVELOPEDGenetic materialSeed size is reduced less than other seg mutants, but the degree reduction is affected by environment. Seed weights of 40, 75, and 90% of normal were obtained from plants grown in the field of Arizona, in the field in Montana, and in the greenhouse in Arizona, USA, respectively. Pollen mother cell meiosis and pollen fertility are normal. Seed from seg7.h plants can be used to establish stands under field conditions. Inheritance: Monofactorial recessive. Location is unknown. Previous nomenclature and gene symbolization: Shrunken endosperm=se7. Mutational events: seg7.h in Ingrid. Mutant used for description and seed stocks: seg7.h in Ingrid (GSHO 2468); seg7.h in Bowman (PI 483237)*2 (GSHO 2352).1545178GSHO 2468
139GSHO 2469seg8.kHordeum vulgare L. subsp. vulgare Arizona, United StatesNSGCKERNELSNot Available1996DEVELOPEDGenetic materialSeed size is reduced and maturity is delayed. Seed weights of 24, 23, and 27% of normal were obtained from plants grown in the field in Arizona, in the field in Montana, and in the greenhouse in Arizona, USA, respectively. Pollen mother cell meiosis and pollen fertility are normal. Seed from seg8.k plants can be used to establish stands under field conditions. Inheritance: Monofactorial recessive. Located in chromosome 7H [1]. Previous nomenclature and gene symbolization: None. Mutational events: seg8.k in 60Ab1810-53. Mutant used for description and seed stocks: seg8.k in 60Ab1810-53 (GSHO 2469); seg8.k in Bowman (PI 483237)*3 (GSHO 1854).1545133GSHO 2469
140GSHO 2475Lys4.dHordeum vulgare L. subsp. vulgare DenmarkNSGCKERNELS1996DEVELOPEDGenetic materialAt the hard dough stage, kernels homozygous for the Lys4.d gene develop a depression near the center of the lemma, which becomes progressively more distinct as the kernels mature. Grain produced by mutant plants has a 9% increase in lysine content. Shrunken endosperm has a pleiotropic association with the high lysine gene Lys4.d. Kernels heterozygous at the Lys4 locus may develop a slightly less distinct depression on the dorsal side. Maintenance of Lys4.d stocks may be difficultu because outcrosses are not recognized easily. Inheritance: Monofactorial incomplete dominant. Located in chromosome 1HS [5S], over 35.1 cM distal from the ert-b (erectoides-b) locus. Previous nomenclature and gene symbolizatioin:Shrunken endosperm xenia 5= sex5g. Mutational events: Lys4.d in Bomi (Riso 8). Mutant used for description and seed stocks: Lys4.d in Bomi (GSHO 2475).1545180GSHO 2475
141GSHO 2476sex6.hHordeum vulgare L. subsp. vulgare North Dakota, United StatesNSGCKERNELS1996DONATED04/1996Genetic materialAfter the hard dough stage, kernels develop a depression in the center of the lemma which becomes progressively more distinct with maturity. The depression is similar in size to that produced by sex1 (shrunken endosperm xenia 1) mutants. The mutant expresses xenia permitting classification of kernels from heterozygous plants as normal or shrunken with an expected 3:1 ratio. Field establishment and growth of mutant plants is normal. Inheritance: Monofactorial recessive. Located in chromosome 7HS [1S], about 2.8 cM distal from the seg2 (shrunken endosperm 2) locus, about 3.1 cM from the centromere, and over 45.8 cM proximal from the Est5 (esterase 5) locus. Previous nomenclature and gene symbolization: None. Mutational events: sex6.h in K6827 (MK6827). Mutant used for description and seed stocks: sex6.h in K6827 (GSHO 2476).1545151GSHO 2476
142GSHO 2477sid1.aHordeum vulgare L. subsp. vulgare JapanNSGCKERNELSNot Available1996DEVELOPEDGenetic materialThe mature plant has several culms, each having only a single elongated internode, the peduncle. Some of the elongated peduncles are much longer than normal. Plants are relatively weak and partially sterile, and have very lax spikes. The expression of mutant traits is less extreme in sid1.b plants, tillers often have two elongated internodes, fertility is better, and the spike is not as lax. Inheritance: Monofactorial recessive. Located in chromosome 4HL, over 13.2 cM distal from the glf3 (glossy leaf 3) locus, and over 29.5 cM proximal from the Kap (hooded lemma) locus. Previous nomenclature and gene symbolization: Node-less dwarf=nls. Mutational events: sid1.a in Akashinriki (R101, OUX052); sid1.b in (GSHO 2478) in Birgitta (NSGC 1870). Mutant used for description and seed stocks: sid1.a in Akashinriki (GSHO 2477); sid1.a in Bowman (PI 483237)*7 (GSHO 2024); sid1.b in Bowman*8 (GSHO 2025).1545053GSHO 2477
143GSHO 2488sld1.aHordeum vulgare L. subsp. vulgare JapanNSGCKERNELS1996DEVELOPEDGenetic materialPlant height is reduced to about 1/2 to 2/3 of normal, plant development is delayed, and plants are slightly yellow green in color. The normal distribution of leaves is altered because the peduncle is relatively long in proportion to the other i internodes. Plants homozygous for the sld1.c allele are slightly weaker and shorter than sld1.a plants in their respective Bowman backcross-derived lines. Inheritance: Monofactorial recessive. Located in chromosome 3HL, over 6.3 cM distal from the uzu (uzu) locus, and about 20.5 cM distal from the uzu locus. Previous nomenclature and gene symbolization: Dwarf 1=dw1. Light green dwarf=dw-1. Mutational events: sld1.a in Akashinriki (OUM148); sld1.c (862PK, GSHO 2489) in Plena (PI 321822). Mutant used for description and seed stocks: sld1.a in Akashinriki (GSHO 2488); sld1.a in Bowman (PI 483237)*7 (GSHO 1970); sld1.c in Bowman*7 (GSHO 1972).1545058GSHO 2488
144GSHO 2491sld2.bHordeum vulgare L. subsp. vulgare JapanNSGCKERNELS1996DEVELOPEDGenetic materialMutant plants have short and slender culms, narrow leaves, short and lax spikes, and thin kernels. When grown at Langdon, North Dakota, USA, dwarf plants are difficult to identify in progenies from crosses between OUM142 and two-rowed spring barley. The frequency of dwarf, late maturing plants is low because sld2.b is linked to an Eam1 (early maturity 1) allele in the repulsion phase. Inheritance: Monofactorial recessive. Located in chromosome 2HS, over 18.2 cM distal from the eog (elongated outer glume) locus. Previous nomenclature and gene symbolization: None. Mutational events: sld2.b in Akashinriki (OUM142). Mutant used for description and seed stocks: sld2.b in Akashinriki (GSHO 2491); sld2.b in Bowman (PI 483237)*4 (GSHO 1876).1545101GSHO 2491
145GSHO 2492sls1.aHordeum vulgare L. subsp. vulgare North Dakota, United StatesNSGCKERNELS1996DONATED04/1996Genetic materialSterile lateral spikelets become progressively smaller toward the tip of two-rowed spikes in plants, which express the Vrs1.b and int-c.b alleles at the six-rowed spike 1 and intermedium spike-c loci, respectively. The terminal rachis nodes generally have rudimentary lateral spikelets. Expression of the sls is affected by genetic background, but often is stronger in late tillers. The trait can be observed only in some two-rowed segregates from crosses between six- and two-rowed cultivars. The sls phenotype can be observed in plants of the Bowman backcross-derived line for the msg1.ca gene. Two-rowed segregates in the Harrington/Morex doubled haploid population show the typical range of expressions associated various combinations of sls and int-c alleles.1545056GSHO 2492
146GSHO 2499wnd1.aHordeum vulgare L. subsp. vulgare JapanNSGCKERNELSNot Available1996DEVELOPEDGenetic materialPlants are semidwarf (15 cm shorter than the original cultivar) and have a pronounced coiling or winding of the upper portion of peduncle. In the Bowman backcross-derived line, which lacks the gene for dense spike (dsp1.a), very little reduction in plant height is observed. However, plants have a distinct coiling of the upper portion of the peduncle. Inheritance: Monofactorial recessive. Located in chromosome 7HS [1S], over 25.4 cM distal from the dsp1 (desnse spike 1) locus. Previous nomenclature and gene symbolization: None. Mutational events: wnd1.a in Kogen-mugi (G754). Mutant used for description and seed stocks: wnd1.a in Kogen-mugi (OUM309, GSHO 2499); wnd1.a in Bowman (PI 483237)*7 (GSHO 1832).1545055GSHO 2499
147GSHO 2500wst6.jHordeum vulgare L. subsp. vulgare JapanNSGCKERNELS1996DEVELOPEDGenetic materialWhen grown under low temperature conditions, the leaf sheath and leaf blade become whitish or white streaked. Only the leaf tip and the central portion of the blade remain green. White streaks are not expressed above 20 degrees Celsius, but the white streak trait is expressed when seedlings are grown in the greenhouse. Inheritance: Monofactorial recessive. Located in chromosome 3HL, about 10.6 cM distal from the uzu (uzu) locus. Previous nomenclature and gene symbolization: White stripe j=wst,,j. Mutational events: wst6.j in Akashinriki (OUM231). Mutant used for description and seed stocks: wst6.j in Akashinriki (GSHO 2500); wst6.j plus uzu1.a in Bowman (PI 483237)*3 (GSHO 1968).1545129GSHO 2500
148GSHO 2504eam10.mHordeum vulgare L. subsp. vulgare North Dakota, United StatesNSGCKERNELS1996DONATED04/1996Genetic materialIn winter nurseries at Cuidad Obregon, Sonora, Mexico and Davis, California, USA, plants of Super Precoz 2H head about 11 days earlier than lines with the genes eam7.g or eam8.k for photoperiod insensitivity from Atsel and Sv Mari, respectively. The eam10.m gene appears to suppress expression of the eam7.g and eam8.k genes. Plants expressing eam10.m become chlorotic (yellow green) under photothermal stress. Zeaxanthin increases at the expense of chlorophyll and other pigments. The chlorotic appearance is similar to that observed in plants homozygous for other recessive genes for early maturity (eam7, eam8, and eam9). Inheritance: Monofactorial recessive. Located in chromosome 3HL, about 2.0 + 5.8 cM from the Est1-Est4 (esterase 1, esterase 4) locus. Previous nomenclature and gene symbolization: Early maturity sp=easp. Mutational events: eam10.m in Super Precoz 2H plus a dominant maturity enhancer; eam10.m in Amber Nude without the enhancer. Mutant used for description and seed stocks: eam10.m in Super Precoz 2H (GSHO 2504); eam10.m in Amber Nude (GSHO 2505).1545060GSHO 2504
149GSHO 2506blx3.cHordeum vulgare L. subsp. vulgare Idaho, United States Historic1996DEVELOPEDGenetic materialBlue aleurone color is due to anthocyanin pigments which occur as lumps inside many aleurone granules in some or all aleurone cells. Variation in color expression from dark blue to an off-whit is caused by environmental factors and modifying genes. Aleurone color is best observed in well filled grain that is magnified to show individual aleurone cells, after more superficial tissues have been peeled off. Inheritance: Monofactorial recessive when complementary dominant alleles are present at the Blx1, Blx2, Blx4, and Blx5 loci. Located in chromosome 4HL, close to the blx1 (non-blue aleurone xenia 1) locus, and over 25.3 cM distal from the Kap (hooded lemma) locus. Previous nomenclature and gene symbolization: Non-blue aleurone 3=bl3. Complementary factors for blue vs white aleurone=Bly and bly. Mutational events: blx3.c in line Blx selected from Composite Cross V. Mutant used for description and seed stocks: blx3.c in line Blx selected from Composite Cross V.1545118GSHO 2506
150GSHO 2507blx4.dHordeum vulgare L. subsp. vulgare North Dakota, United StatesNSGCKERNELS1996DONATED04/1996Genetic materialBlue or pink aleurone color is due to anthocyanin pigments which occur as lumps inside many aleurone granules in some or all aleurone cells. Variation in color expression from pink to an off-white is caused by environmental factors and modifying genes. The pink and red color aleurone colors are easier to observe in well filled grain that is magnified to show individual aleurone cells, after more superficial tissues have been peeled off. A brick red aleurone color results from the interaction of the ibl (intense blue aleurone) gene with the blx4.d gene in homozyotes. Inheritance: Monofactorial recessive when complementary dominant alleles are present at the Blx1, Blx2, Blx3, and Blx5 loci. Located in chromosome 4HL, very close to the blx1 (non-blue aleurone xenia 1) locus, and over 29.5 cM distal from the Kap (hooded lemma) locus. Previous nomenclature and gene symbolization: Non-blue (pink) aleurone 4=bl4. Mutational events: blx4.d plus ibl in Ethiopian 637; blx4.d only in Ab 6, EP79, Grannenlose Zweizeilige. Mutant used for description and seed stocks: blx4.d in Ab 6 (PI ??).1545102GSHO 2507
151GSHO 2509blx5.eHordeum vulgare L. subsp. vulgare North Dakota, United StatesNSGCKERNELS1996DONATED04/1996Genetic materialBlue aleurone color is due to anthocyanin pigments which occur as lumps inside many aleurone granules in some or all aleurone cells. Variation in color expression from dark blue to an off-white is caused by environmental factors and modifying genes. Aleurone color is best observed in well filled grain that is magnified to show individual aleurone cells, after more superficial tissues have been peeled off. Inheritance: Monofactorial recessive when complementary dominant alleles are present at the Blx1, Blx2, Blx3, and Blx4 loci. Located in chromosome 7HL [1L], associated with the red stem trait controlled by the dominant allele at the ant1 (anthocyanin-less 1) locus; linked to the blx2 (non-blue aleurone xenia 2) locus, but not to the fch12 (chlorina seedling 12) locus. Previous nomenclature and gene symbolization: Non-blue aleurone 5=bl5. Mutational events: blx5.e in BGM 122. Mutant used for description and seed stocks: blx5.e in BGM 122.1545049GSHO 2509
152GSHO 2510Rsp1.aHordeum vulgare L. subsp. vulgare North Dakota, United States Historic1996DONATED04/1996Genetic materialThe Rsp1.a gene conditions a high level of resistance to a single spore culture of Septoria passerinii isolated in Minnesota, USA. Infection occurred on F1 seedlings, but was limited to a few lesions. Inheritance: Monofactorial incomplete dominant. Location is unknown, but not linked to the Rsp2 (reaction to Septoria passerinii 2) locus or the Rsp3 locus. Previous nomenclature and gene symbolization: Resistance to Septoria passerinii Sacc. = Sep. Mutational events: Rsp1.a in CIho 14300. Mutant used for description and seed stocks: Rsp1.a in CIho 14300 (GSHO 2510).1545037GSHO 2510
153GSHO 2511Rsp2.bHordeum vulgare L. subsp. vulgare North Dakota, United States Historic1996DONATED04/1996Genetic materialThe Rsp2.b gene conditions a high level of resistance to a single spore culture of Septoria passerinii isolated in Minnesota, USA. Pycnidia were observed in some, but not all lesions, on all F1 plants. Inheritance: Monofactorial incomplete dominant. Location is unknown, but linked to the Rsp3 (reaction to Septoria passerinii 3) locus and not linked to the Rsp1 locus. Previous nomenclature and gene symbolization: Resistance to Septoria passerinii Sacc.= Sep2. Mutational events: Rsp2.b in PI 70837. Mutant used for description and seed stocks: Rsp2.b in PI 70837 (GSHO 2511).1545038GSHO 2511
154GSHO 2512Rsp3.cHordeum vulgare L. subsp. vulgare North Dakota, United States Historic1996DONATED04/1996Genetic materialThe Rsp3.c gene conditions a high level of resistance to a single spore culture of Septoria passerinii isolated in Minnesota, USA. Infection occurred on F1 seedlings, but was limited to a few lesions. Inheritance: Monofactorial incomplete dominant. Location is unknown, but linked to the Rsp2 (reaction to Septoria passerinii 2) locus and not linked to Rsp1 locus. Previous nomenclature and gene symbolization: Resistance to Septoria passerinni Sacc. = Sep3. Mutational events: Rsp3.c in CIho 10644. Mutant used for description and seed stocks: Rsp3.c in CIho 10644 (GSHO 2512).1545155GSHO 2512
155GSHO 2513sdw1.aHordeum vulgare L. subsp. vulgare North Dakota, United States Historic1996DONATED04/1996Genetic materialResistant seedlings infested with greenbugs (aphids) are not killed or severely stunted by a buildup of the greenbug population, but susceptible seedlings are killed or severely stunted. Inheritance: Monofactorial dominant. Location is unknown. Previous nomenclature and gene symbolization: None. Mutational events: Rsg2.b in PI 426756. Mutant used for description and seed stocks: Rsg2.b in PI 426756.1545171GSHO 2513
156GSHO 984Rym2.bHordeum vulgare L. subsp. vulgare JapanNSGCKERNELS1996DEVELOPEDGenetic materialPlants with Rym2.b allele have much less yellowing and stunting than susceptible plants when grown if fields that are infested with BaYMV. Also, Mihori Hadaka 3 is resistant to BaYMV, BaYMV-2, and BaMMV in Germany. Inheritance: Monofactorial incomplete dominant. Located in chromosome 7HL [1L], approximately 31.4 cM distal from the nud (naked caryopsis) locus. Previous nomenclature and gene symbolization: Resistance to BaYMV 2=Ym2. Mutational events: Rym2.b in Mihori Hadaka 3. Mutant used for description and seed stocks: Rym2.b in Mihori Hadaka 3 (OUJ373, GSHO 984).1514465GSHO 984
157GSHO 987min1.aHordeum vulgare L. subsp. vulgare JapanNSGCKERNELS1996DEVELOPEDGenetic materialPlants are 1/2 as tall as the original cultivar, spikes and awns are slightly shorter, and the number of tillers, spikelets per spike, and numbers of kernels are slightly less than in the original cultivar. In meristematic tissues of root tips and young shoots, polyploid nuclei consisting of 4x, 8x, and sometimes many more chromosomes are gound together with normal diploid cells. Epridermal cells of the leaves are sometimes irregular and variable in shape and size. The mixoploid condition of tissues is believed to result from incomplete cytokinesis following nuclear division. Plant vigor in the Bowman backcross-derived line is very sensitive to environmental stress. When grown in the greenhouse, the plants are 2/3 normal height, spike size and length are reduced, and leaves are narrow. Under field conditions, plants are diminutive (less than 1/4 normal height and size) and seed set is low. Inheritance: Monofactorial recessive, but interaction with the recessive allele at the en- min (enhancer of minute) locus is observed in some cultivars. Located in chromosome 4HL, about 6.7 cM distal from the glf3 (glossy leaf 3) locus. Previous nomenclature and gene symbolization: Dwarf=d. Mutational events: min1.a in Taisho-mugi with En- min gene in the genetic background. Mutant used for description and seed stocks: min1.a in Taisho-mugi (GSHO 987); min1.a in Bowman (PI 483237)*7 (GSHO 2023).1513789GSHO 987
158GSHO 989nec1.aHordeum vulgare L. subsp. vulgare DenmarkNSGCKERNELS1996DEVELOPEDGenetic materialSmall black-brown spots develop on all light-exposed parts of the plant starting near the leaf tip at the three-leaf stage. The spots are oval (the longest dimension is parallel to the leaf veins) and generally less than 1 to 2 mm in size. The spots are concentrated in awn and the most distal parts of the leaf blade, but may occur on all plant parts. Inheritance: Monofactorial recessive. Located in chromosome 1HL [5L], near the centromere and about 34.5 cM proximal from the wst5 (white streak 5) locus, and about 10.0 cM distal from the msg1 (male sterile genetic 1) locus. Previous nomenclature and gene symbolization: Mutant no 10. Parkland spot=sp,,b. Mutational events: nec1.a in Carlsberg II (Mutant no 10); sp,,b (GSHO 1284) in Parkland (CIho 10001); a mutant in Morex (CIho 15773). Mutant used for description and seed stocks: nec1.a in Carlsberg II (GSHO 989); nec1.a from Wolfe's Multiple Recessive Marker Stock for Chromosome 5 in Bowman (PI 483237)*7 (GSHO 2052).1514322GSHO 989
159GSHO 1518cer-e.8Hordeum vulgare L. subsp. vulgare Skåne län, SwedenNSGCKERNELS1995DEVELOPED58.98333300, 13.50000000Genetic materialSurface wax coating on the spike appears to be absent or greatly reduced (wax code -/+ ++ ++). A translocation is present in the allele cer-e.18, but it is not linked to the cer-e locus. The Bowman backcross-derived line has less wax on the spike and upper leaf sheath (wax code - + ++). Inheritance: Monofactorial recessive. Located in chromosome 1HL [5L], close to the centromere and near the ert-b (erectoides-b) locus. Previous nomenclature and gene symbolization: None. Mutational events: cer-e.8, -e.18 (trans), -e.34 in Bonus; cer-e.90, -e.140 in Bonus, -e.272, -e.294, -e.388, -e.551, -e.560, -e.586, -e.607 in Foma (CIho 11333); cer-e.273, -e.277, -e.296, -e.317, -e.321, -e.473, -e.480 in Foma; cer-e.663, e.692, -e.716, -e.717 in Bonus; cer.161 in Bonus, -e.577 in Foma, -e.790 in Bonus, -e.1089, -e.1106 in Kristina; cer-e.908, -e.962 in Bonus, -e.1162 in Kristina; cer-e.1335 in Bonus; cer-e.1366, -e.1404, -e.1425, -e.1426 in Bonus; cer-e.1403, -e.1709, -e.1715 in Bonus; cer-e.1519 in Nordal, e.1760 in Bonus, -e.1873, -e.1878, -e.1880, -e.1884 in Sv 79353. Mutant used for description and seed stocks: cer-e.8 in Bonus (GSHO 1518); cer-e.8 in Bowman (PI 483237)*7 (GSHO 2048).1521258GSHO 1518
160GSHO 1519cer-w.38Hordeum vulgare L. subsp. vulgare Skåne län, SwedenNSGCKERNELS1995DEVELOPED58.98333300, 13.50000000Genetic materialSurface wax coating on the spike appears to be greatly reduced or absent (wax code +/- ++ ++). Reduced surface wax coating occurs on the leaf sheath and the stem in the Bowman backcross-derived line (wax code - + ++). Nodes lack surface wax and have a glossy appearance. Inheritance: Monofactorial recessive. Located in chromosome 5HL [7L], about 2.6 cM proximal from the raw1 (smooth awn 1) locus, but longer distances were reported. Previous nomenclature and gene symbolization: None. Mutational events: cer-w.48 in Bonus; cer-w.280, -w.347 (444), -w.351, -w.359, -w.375, -w.402 in Foma (CIho 11333); cer-w.380, -w.484, -w.495, -w.559, -w.569, -w.603, -w.604 in Foma, -w.691 in Bonus; cer-w.771 in Bonus; cer-w.858 in Bonus, -w.1192 in Kristina; cer-w.1445 in Bonus. Mutant used for description and seed stocks: cer-w.48 in Bonus (GSHO 1519); cer-w.48 in Bowman (PI 483237)*3 (GSHO 2122).1521259GSHO 1519
161GSHO 1521cer-za.227Hordeum vulgare L. subsp. vulgare Skåne län, SwedenNSGCKERNELS1995DEVELOPED58.98333300, 13.50000000Genetic materialSurface wax coating on the leaf blade appears to be absent (wax code ++ ++ -). Inheritance: Monofactorial recessive. Located in chromosome 5HL [7L], close to the raw1 (smooth awn 1) locus based on linkage drag. Previous nomenclature and gene symbolization: None. Mutational events: cer-za.126, -za.173, -za.185, -za.186 in Bonus (PI 189763); cer-za.227, -za.232, -za.318, -za.323, -za.328, -za.441, -za.462, -za.482, -za.490, -za.504 in Foma; cer-za.378, -za.465, -za.530, -za.541, -za.552, -za.572, -za.575, -za.583, -za.587, -za.588 in Foma, -za.672, -za.676, -za.685, -za.735, -za.739, -za.741 in Bonus; cer-za.681, -za.768, -za.782, -za.812, -za.831, -za.833, -za.853, -za.854, -za.869 in Bonus, -za.1122, -za.1126 in Kristina; cer-za.628, -za.637, -za.639, -za.643, -za.917 in Bonus, -za.1159, -za.1186, -za.1190, -za.1191 in Kristina; cer-za.478 in Foma, -za.1157, -za.1178, -za.1189, -za.1203, -za.1224, -za.1228, -za.1252 in Kristina; cer-za.1286 in Kristina, -za.1324, -za.1346 in Bonus; cer-za.1284 in Kristina, -za.1357, -za.1399, -za.1407, -za.1408, -za.1431 in Bonus; cer-za.1546 in Nordal, -za.1713, -za.1738, -za.1745, -za.1747, -za.1800, -za.1805, -za.1806, -za.1807, -za.1812, -za.1813 in Bonus. Mutant used for description and seed stocks: cer-za.227 in Foma (GSHO 1521); cer-za. 227 in Bowman (PI 483237)*6 (GSHO 2126).1521261GSHO 1521
162GSHO 1522cer-zb.38Hordeum vulgare L. subsp. vulgare Skåne län, Sweden SPIKEHistoric1995DEVELOPED58.98333300, 13.50000000Genetic materialSurface wax coating on the spike appears to be absent (wax code - ++ ++). All cer-zb mutants have pointed lateral spikelets, reduced fertility and narrow leaves. The Bowman backcross-derived line has reduced vigor, small spikes (about 1/2 normal), and narrow leaves; also tan necrotic blotches may develop prior to maturity. Inheritance: Monofactorial recessive. Location is unknown. Previous nomenclature and gene symbolization: None. Mutational events: cer-zb.38 in Bonus; cer-zb.105 in Bonus, -zb.279 in Foma (CIho 11333); cer-zb.1164 in Kristina; cer-zb.1279 in Kristina; cer-zb.1273 in Kristina. Mutant used for description and seed stocks: cer-zb.38 in Bonus (GSHO 1522); cer-zb.38 in Bowman (PI 483237)*5 (GSHO 2196).1521262GSHO 1522
163GSHO 1523cer-zn.244Hordeum vulgare L. subsp. vulgare Skåne län, SwedenNSGCKERNELS1995DEVELOPED58.98333300, 13.50000000Genetic materialSurface wax coating on the spike appears to be greatly reduced or absent (wax code +/- ++ ++). Inheritance: Monofactorial recessive. Location in chromosome 3HL, over 12.0 cM distal from the uzu (uzu) locus. Previous nomenclature and gene symbolization: None. Mutational events: cer-zn.244, -zn.362 in Foma; cer-zn.348, -zn.385, -zn.540 in Foma; cer-zn.1270 in Kristina. Mutant used for description and seed stocks: cer-zn.244 in Foma (GSHO 1523); cer-zn.244 in Bowman (PI 483237)*5 (GSHO 1983).1521263GSHO 1523
164GSHO 1524cer-zq.248Hordeum vulgare L. subsp. vulgare Skåne län, SwedenNSGCKERNELS1995DEVELOPED58.98333300, 13.50000000Genetic materialSurface wax coating on the leaf blade appears to be absent, but only on the three upper leaf blades (wax code ++ ++ -). The plant growth remains for a long time vegetative tuft before heading. The Bowman backcross-derived line is a semidwarf, about 3/4 normal height. Inheritance: Monofactorial recessive. Location is unknown. Previous nomenclature and gene symbolization: None. Mutational events: cer-zq.248 in Foma; cer-zq.761 in Bonus (PI 189763); cer-zq.1343 in Bonus; cer-zq.1378 in Bonus. Mutant used for description and seed stocks: cer-zq.248 in Foma (GSHO 1524); cer-zq.248 in Bowman (PI 483237)*5 (GSHO 2202).1521264GSHO 1524
165GSHO 1525cer-zr.260Hordeum vulgare L. subsp. vulgare Skåne län, SwedenNSGCKERNELS1995DEVELOPED58.98333300, 13.50000000Genetic materialSurface wax coating on the spike appears to be reduced or absent (wax code +/- ++ ++). The wax coating appears to be absent on the spike and nodes and reduced on stem and leaf sheath (wax code -+/++ ++) in the Bowman backcross-derived line. Inheritance: Monofactorial recessive. Location is unknown. Previous nomenclature and gene symbolization: None. Mutational events: cer-zr.260 in Foma; cer-zr.528 in Foma. Mutant used for description and seed stocks: cer-zr.260 in Foma (GSHO 1525); cer-zr.260 in Bowman (PI 483237)*3 (GSHO 2203).1521265GSHO 1525
166GSHO 1526cer-zs.467Hordeum vulgare L. subsp. vulgare Skåne län, SwedenNSGCKERNELS1995DEVELOPED58.98333300, 13.50000000Genetic materialSurface wax coating on the spike is reduced (wax code + ++ ++), maturity is delayed, and vigor is reduced somewhat. Inheritance: Monofactorial recessive. Location is unknown. Previous nomenclature and gene symbolization: None. Mutational events: cer-zs.467, -zs.474, -zs.475 in Foma; cer-zs.594 in Foma. Mutant used for description and seed stocks: cer-zs.467 in Foma (GSHO 1526).1521266GSHO 1526
167GSHO 1527cer-zt.389Hordeum vulgare L. subsp. vulgare Skåne län, SwedenNSGCKERNELS1995DEVELOPED58.98333300, 13.50000000Genetic materialSurface wax coating on the spike only appears to be slightly reduced (wax code + ++ ++). The reduction in the surface wax coating was greater in plants selected from the first backcross to Bowman (wax code -/+ + ++). Inheritance: Monofactorial recessive. Location is unknown. Previous nomenclature and gene symbolization: None. Mutational events: cer-zt.389, -zt.479 in Foma. Mutant used for description and seed stocks: cer-zt.389 in Foma (GSHO 1527); cer-zt.389 in Bowman (PI 483237)*2 (GSHO 2205).1521267GSHO 1527
168GSHO 1528cer-zu.122Hordeum vulgare L. subsp. vulgare Skåne län, SwedenNSGCKERNELS1995DEVELOPED58.98333300, 13.50000000Genetic materialSurface wax coating on the spike appears to be absent, the wax coating on the leaf sheath and stem is reduced (wax code - + ++). Leaf blades become golden yellow at or shortly after heading and later necrotic spots develop on the upper surface of the blade. Inheritance: Monofactorial recessive. Location is unknown. Previous nomenclature and gene symbolization: None. Mutational events: cer-zu.122 in Bonus; cer-zu.973 in Bonus; cer-zu.1163 in Kristina, -zu.1419, -zu.1428, -zu.1447 in Bonus; cer-zu.1494, -zu.1570, -zu.1571, -zu.1762, -zu.1784 in Bonus, -zu.`570 in Nordal. Mutant used for description and seed stocks: cer-zu.122 in Bonus (GSHO 1528); cer-zu.122 in Bowman (PI 483237)*5 (GSHO 2206).1521268GSHO 1528
169GSHO 1529cer-zv.268Hordeum vulgare L. subsp. vulgare Skåne län, SwedenNSGCKERNELS1995DEVELOPED58.98333300, 13.50000000Genetic materialSurface wax coating on the spike, the leaf sheath and stem, and the leaf blade appears to be absent (wax code - - -). Plants are semidwarf, relatively weak, and head late, and the hull is poorly attached to the seed. The Bowman backcross-derived line is a semidwarf (2/3 normal height) with a kinked upper peduncle and a strong wax coating on the nodes. Inheritance: Monofactorial recessive. Location is unknown. Previous nomenclature and gene symbolization: None. Mutational events: cer-zv.268, -zv.342, -zv.576 in Foma; cer-zv.1246 in Kristina. Mutant used for description and seed stocks: cer-zv.268 in Foma (GSHO 1529); cer-zv.268 in Bowman (PI 483237)*5 (GSHO 2207).1521269GSHO 1529
170GSHO 1530cer-zw.286Hordeum vulgare L. subsp. vulgare Skåne län, SwedenNSGCKERNELS1995DEVELOPED58.98333300, 13.50000000Genetic materialSurface wax coating on the spike, leaf sheath, and stem appears to be reduced (+ + ++) with leaf sheath and stem wax alternating in broad horizontal bands of waxless and wax covered zones (zebra-like stripes). Inheritance: Monofactorial recessive. Location is unknown. Previous nomenclature and gene symbolization: None. Mutational events: cer-zw.286 in Foma. Mutant used for description and seed stocks: cer-zw.286 in Foma (GSHO 1530); cer-zw.286 in Bowman (PI 483237)*2 (GSHO 2208).1521270GSHO 1530
171GSHO 1531cer-zx.100Hordeum vulgare L. subsp. vulgare Skåne län, SwedenNSGCKERNELS1995DEVELOPED58.98333300, 13.50000000Genetic materialSurface wax coating on the spike, leaf sheath, and stem appears to be reduced (wax code + + ++). Inheritance: Monofactorial recessive. Location is unknown. Previous nomenclature and gene symbolization: None. Mutational events: cer-zx.100 in Bonus. Mutant used for description and seed stocks: cer-zx.100 in Bonus (GSHO 153`); cer-zx.100 in Bowman (PI 483237)*2 (GSHO 1531).1521271GSHO 1531
172GSHO 1532cer-zy.118Hordeum vulgare L. subsp. vulgare Skåne län, SwedenNSGCKERNELS1995DEVELOPED58.98333300, 13.50000000Genetic materialSurface wax coating on the leaf blade appears to be slightly reduced (wax code ++ ++ +). Seedling leaves have a slightly yellow green color similar to that found in some chlorina mutants. Inheritance: Monofactorial recessive. Location is unknown. Previous nomenclature and gene symbolization: None. Mutational events: cer-zy.118 in Bonus; cer-zy.136 in Bonus. Mutant used for description and seed stocks: cer-zy.118 in Bonus (GSHO 1532); cer-zy.118 in Bowman (Pi 483237)*3.1521272GSHO 1532
173GSHO 1533cer-zz.615Hordeum vulgare L. subsp. vulgare Skåne län, SwedenNSGCKERNELS1995DEVELOPED58.98333300, 13.50000000Genetic materialSurface wax coating on the leaf blade appears to be absent (wax code ++ ++ -). Inheritance: Monofactorial recessive. Location is unknown. Previous nomenclature and gene symbolization: None. Mutational events: cer-zz.615 in Foma. Mutant used for description and seed stocks: cer-zz.615 in Foma (GSHO 1533); cer-zz.615 in Bowman (PI 483237)*2 (GSHO 2210).1521273GSHO 1533
174GSHO 1534cer-ya.180Hordeum vulgare L. subsp. vulgare Skåne län, SwedenNSGCKERNELS1995DEVELOPED58.98333300, 13.50000000Genetic materialSurface wax coating on the leaf blade appears to be absent (wax code ++ ++ -). Leaves have a light green color that persists from the seedling stage until maturity. Heading and maturity are delayed in plants homozygous for the cer-ya.180 gene. Inheritance: Monofactorial recessive. Located in chromosome 3HS, probably linked to the Btr (brittle rachis) locus based on linkage drag. Previous nomenclature and gene symbolization: None. Mutational events: cer-ya.180 in Bonus. Mutant used for description and seed stocks: cer-ya.180 in Bonus (GSHO 1534); cer-ya.180 in Bowman (PI 483237)*3.1521274GSHO 1534
175GSHO 1535cer-yb.200Hordeum vulgare L. subsp. vulgare Skåne län, SwedenNSGCKERNELS1995DEVELOPED58.98333300, 13.50000000Genetic materialSurface wax coating on the leaf blade appears to be absent (wax code ++ ++ -). Plants expressing the cer-yb gene are relatively late. Inheritance: Monofactorial recessive. Located in chromosome 2HL, near the Gth (toothed lemma) locus based on linkage drag. Previous nomenclature and gene symbolization: None. Mutational events: cer-yb.200 in Bonus, -yb.316, -yb.453 in Foma (CIho 11333); cer-yb.1174 in Kristina; cer-yb.1211 in Kristina. Mutant used for description and seed stocks: cer-yb.200 in Bonus (GSHO 1535); cer-yb.200 in Bowman (PI 483237)*6 (GSHO 1896).1521275GSHO 1535
176GSHO 1536cer-yc.135Hordeum vulgare L. subsp. vulgare Skåne län, SwedenNSGCKERNELS1995DEVELOPED58.98333300, 13.50000000Genetic materialSurface wax coating on the spike appears to be absent (wax code - ++ ++). Inheritance: Monofactorial recessive. Location is unknown. Previous nomenclature and gene symbolization: None. Mutational events: cer-yc.135 in Bonus; cer-yc.523 in Foma (CIho 11333); cer-yc.438, -yc.439 in Foma, -yc.783 in Bonus. Mutant used for description and seed stocks: cer-yc.135 in Bonus (GSHO 1536); cer-yc.135 in Bowman (PI 483237)*3 (GSHO 2211).1521276GSHO 1536
177GSHO 1537cer-yd.139Hordeum vulgare L. subsp. vulgare Skåne län, SwedenNSGCKERNELS1995DEVELOPED58.98333300, 13.50000000Genetic materialSurface wax coating on the spike appears to be absent (wax code - ++ ++). The original cer-yd.139 stock has narrow leaf blades, pointed lateral spikelets, and semi-sterility. In the Bowman backcross-derived line, surface wax coating appears to be reduced (wax code + ++ ++). The Bowman backcross-derived line has very few stigma hairs and slightly reduced to poor seed set depending upon environmental conditions, but male fertility is normal. Lodicules may be pistiloid in appearance. Inheritance: Monofactorial recessive. Located in chromosome 3HS, based on linkage drag with the Btr (brittle rachis) locus. Previous nomenclature and gene symbolization: None. Mutational events: cer-yd.139 in Bonus. Mutant used for description and seed stocks: cer-yd.139 in Bonus (GSHO 1537); cer-yd.139 in Bowman (PI 483237)*4 (GSHO 1946).1521277GSHO 1537
178GSHO 1538cer-ye.267Hordeum vulgare L. subsp. vulgare Skåne län, SwedenNSGCKERNELS1995DEVELOPED58.98333300, 13.50000000Genetic materialSurface wax coating on the leaf blade appears to be absent (wax code ++ ++-). Inheritance: Monofactorial recessive. Located in chromosome 5HL [7L], based on linkage drag with the raw1 (smooth awn 1) locus. Previous nomenclature and gene symbolization: None. Mutational events: cer-ye.267, -ye.582 in Foma, -ye.792 in Bonus (PI 189763); cer-ye.1103 in Kristina; cer-ye.1395 in Bonus. Mutant used for description and seed stocks: cer-ye.267 in Foma (GSHO 1538); cer-ye.267 in Bowman (PI 483237)*6 (GSHO 2127).1521278GSHO 1538
179GSHO 1539cer-yf.652Hordeum vulgare L. subsp. vulgare Skåne län, SwedenNSGCKERNELS1995DEVELOPED58.98333300, 13.50000000Genetic materialSurface wax coating on the leaf blade appears to be reduced (wax code ++ ++ +). In the Bowman backcross-derived line, mutant plants have pale geen leaves and heading is delayed. Inheritance: Monofactorial recessive. Location is unknown. Previous nomenclature and gene symbolization: None. Mutational events: cer-yf.652 in Foma, -yf.804 in Bonus (PI 189763). Mutant used for description and seed stocks: cer-yf.652 in Foma (GSHO 1539); cer-yf.652 in Bowman (PI 483237)*3 (GSHO 2212).1521279GSHO 1539
180GSHO 1540cer-yg.1014Hordeum vulgare L. subsp. vulgare Skåne län, SwedenNSGCKERNELS1995DEVELOPED58.98333300, 13.50000000Genetic materialSurface wax coating on the spike, leaf sheath, stem, and leaf blade appears to be absent (wax code - - -). The Bowman backcross-derived line is semidwarf (3/4 normal), and surface wax is present on the leaf blades (wax code - - +). Inheritance: Monofactorial recessive. Located in chromosome 7HS [1S], based on linkage drag with the ant1 (anthocyanin-less 1) locus. Previous nomenclature and gene symbolization: None. Mutational events: cer-yg.1014 in Carlsberg II. Mutant used for description and seed stocks: cer-yg.1014 in Carlsberg II (GSHO 1540); cer-yg.1014 in Bowman (PI 483237)*4 (GSHO 1839).1521280GSHO 1540
181GSHO 1541cer-yh.116Hordeum vulgare L. subsp. vulgare Skåne län, SwedenNSGCKERNELS1995DEVELOPED58.98333300, 13.50000000Genetic materialSurface wax coating on the spike appears to be absent (wax code - ++ ++). The mutant is semi-sterile, is late in heading, and has pointed lateral spikelets. The Bowman backcross-derived line is partially to highly female sterile because few hairs develop on the stigma. Although the anthers are slightly malformed, male fertility is good. Inheritance: Monofactorial recessive. Located in chromosome 3HS, based on linkage drag with the Btr (brittle rachis) locus. Previous nomenclature and gene symbolization: None. Mutational events: cer-yh.116, -yh.654 in Bonus. Mutant used for description and seed stocks: cer-yh.116 in Bonus (GSHO 1541); cer-yh.116 in Bowman (PI 483237)*4 (GSHO 1947).1521281GSHO 1541
182GSHO 1542cer-yi.254Hordeum vulgare L. subsp. vulgare Skåne län, SwedenNSGCKERNELS1995DEVELOPED58.98333300, 13.50000000Genetic materialSurface wax coating on the leaf blade appears to be absent (wax code ++ ++ -). A translocation is probably present in the original stock and may be associated with the cer-yi.254 allele. Inheritance: Monofactorial recessive. Location is unknown. Previous nomenclature and gene symbolization: None. Mutational events: cer-yi.254 in Foma, -yi.956 in Bonus (PI 189763); cer-yi.1213 in Kristina, -yi.1353 in Bonus. Mutant used for description and seed stocks; cer-yi.254 in Foma (GSHO 1542); cer-yi.254 in Bowman (PI 483237)*3 (GSHO 2213).1521282GSHO 1542
183GSHO 1543cer-yj.667Hordeum vulgare L. subsp. vulgare Skåne län, SwedenNSGCKERNELS1995DEVELOPED58.98333300, 13.50000000Genetic materialSurface wax coating on the leaf blade appears to be absent (wax code ++ ++ -). Plants of the original stock in Bonus are semidwarf (about 3/4 normal height). Inheritance: Monofactorial recessive. Location is unknown. Previous nomenclature and gene symbolization: None. Mutational events: cer-yj.667 in Bonus, -yj.1217 in Kristina; cer-yj.937 in Bonus; cer-yj.1150 in Kristina, -yj.1811 in Bonus. Mutant used for description and seed stocks: cer-yj.667 in Bonus (GSHO 1543); cer-yj.667 in Bowman (PI 483237)*3 (GSHO 2214).1521283GSHO 1543
184GSHO 1544cer-yk.627Hordeum vulgare L. subsp. vulgare Skåne län, SwedenNSGCKERNELS1995DEVELOPED58.98333300, 13.50000000Genetic materialSurface wax coating on the spike, leaf sheath, and stem appears to be reduced (wax code + + ++). The original stock in Bonus has narrow leaves, a lax spike and a strong wax coating on the kernels. The Bowman backcross-derived line has wax covering on the peduncle, but nodes are not covered with wax (glossy node). Inheritance: Monofactorial recessive. Location is unknown. Previous nomenclature and gene symbolization: None. Mutational events: cer-yk.627 in Bonus. Mutant used for description and seed stocks: cer-yk.627 in Bonus (GSHO 1544); cer-yk.627 in Bowman (PI 483237)*4 (GSHO 2215).1521284GSHO 1544
185GSHO 1545cer-yl.187Hordeum vulgare L. subsp. vulgare Skåne län, SwedenNSGCKERNELS1995DEVELOPED58.98333300, 13.50000000Genetic materialSurface wax coating on the spike, the leaf sheath, and the stem appears to be absent (wax code - - ++). The original mutant stock produces weak plants (about 3/4 normal height) with a high level of sterility. Plants in the Bowman backcross-derived line are semidwarf with waxy nodes, coiled peduncles, globose shaped seeds, and poorly attached hull. Reduced seed set can be caused by adverse environmental conditions. Inheritance: Monofactorial recessive. Location is unknown. Previous nomenclature and gene symbolization: None. Mutational events: cer-yl.187, -yl.188 in Bonus; cer-yl.407 in Foma (CIho 11333), -yl.821 in Bonus. Mutant used for description and seed stocks: cer-yl.187 in Bonus (GSHO 1545); cer-yl.187 in Bowman (PI 483237)*5 (GSHO 2216).1521285GSHO 1545
186GSHO 1546cer-ym.753Hordeum vulgare L. subsp. vulgare Skåne län, SwedenNSGCKERNELS1995DEVELOPED58.98333300, 13.50000000Genetic materialSurface wax coating on the spike, the leaf sheath and stem, and the leaf blade appears to be absent (wax code - - -). Plants are weak and semidwarf (2/3 normal height) and heading is delayed. In the Bowman backcross-derived line, plants are weak, culms have a kinked peduncle, nodes are wax covered, and seeds are small, globe shaped, and nearly hull-less. Inheritance: Monofactorial recessive. Location is unknown. Previous nomenclature and gene symbolization: None. Mutational events: cer-ym.130, -ym.753 in Bonus. Mutant used for description and seed stocks: cer-ym.753 in Bonus (GSHO 1546); cer-ym.753 in Bowman (PI 483237)*5 (GSHO 2217).1521286GSHO 1546
187GSHO 1547cer-yn.1112Hordeum vulgare L. subsp. vulgare Skåne län, SwedenNSGCKERNELS1995DEVELOPED58.98333300, 13.50000000Genetic materialSurface wax coating on the spike, leaf sheath, and stem appears to be reduced (wax code + + ++). In the Bowman backcross-derived line, the wax coating is only slightly reduced, but nodes are glossy. Inheritance: Monofactorial recessive. Location is unknown. Previous nomenclature and gene symbolization: None. Mutational events: cer-yn.1112 in Kristina. Mutant used for description and seed stocks: cer-yn.1112 in Kristina (GSHO 1547); cer-yn.1112 in Bowman (PI 483237)*2 (GSHO 2218).1521287GSHO 1547
188GSHO 1548cer-yo.647Hordeum vulgare L. subsp. vulgare Skåne län, SwedenNSGCKERNELS1995DEVELOPED58.98333300, 13.50000000Genetic materialSurface wax coating on the leaf blade appears to be reduced (wax code ++ ++ +). The original stock in Bonus has a high level of sterility. Seedlings of the original stock and the Bowman backcross-derived line have slightly pale green color. Inheritance: Monofactorial recessive. Location is unknown. Previous nomenclature and gene symbolization: None. Mutational events: cer-yo.647 in Bonus. Mutant used for description and seed stocks: cer-yo.647 in Bonus (GSHO 1548); cer-yo.647 in Bowman (PI 483237)*3 (GSHO 2219).1521288GSHO 1548
189GSHO 1549cer-yp.949Hordeum vulgare L. subsp. vulgare Skåne län, SwedenNSGCKERNELS1995DEVELOPED58.98333300, 13.50000000Genetic materialSurface wax coating on the leaf blade appears to be reduced (wax code ++ ++ +). Leaves have a pale green color similar that of some chlorina mutants. Inheritance: Monofactorial recessive. Location is unknown. Previous nomenclature and gene symbolization: None. Mutational events: cer-yp.949, -yp.955 in Bonus. Mutant used for description and seed stocks: cer-yp.949 in Bonus (GSHO 1549); cer-yp.949 in Bowman (PI 483237)*2 (GSHO 2220).1521289GSHO 1549
190GSHO 1550cer-yq.1246Hordeum vulgare L. subsp. vulgare Skåne län, SwedenNSGCKERNELS1995DEVELOPED58.98333300, 13.50000000Genetic materialSurface wax coating on the leaf blade appears to be absent (wax code ++ ++ -). In the Bowman backcross-derived line, the wax coating is reduced only on upper leaf blades (wax code ++ ++ +). Inheritance: Monofactorial recessive. Location is unknown. Previous nomenclature and gene symbolization: None. Mutational events: cer-yq.1246 in Kristina. Mutant used for description and seed stocks: cer-yq.1246 in Kristina (GSHO 1550); cer-yq.1246 in Bowman (PI 483237)*3 (GSHO 2221).1521290GSHO 1550
191GSHO 1551cer-yr.492Hordeum vulgare L. subsp. vulgare Skåne län, SwedenNSGCKERNELS1995DEVELOPED58.98333300, 13.50000000Genetic materialSurface wax coating on the spike appears to be greatly reduced or absent, and surface wax coating on the leaf sheath and stem is reduced (wax code -/+ + ++). Nodes are glossy and appear to lack surface wax. Inheritance: Monofactorial recessive. Location is unknown. Previous nomenclature and gene symbolization: None. Mutational events: cer-yr.492 in Foma, -yr.1418 in Bonus (PI 189763). Mutant used for description and seed stocks: cer-yr.492 in Foma (GSHO 1551).1521291GSHO 1551
192GSHO 1552cer-ys.680Hordeum vulgare L. subsp. vulgare Skåne län, SwedenNSGCKERNELS1995DEVELOPED58.98333300, 13.50000000Genetic materialSurface was coating on the leaf blade appears to be absent (wax code ++ ++ -). The original stock in Bonus has a lax spike. Inheritance: Monofactorial recessive. Location is unknown. Previous nomenclature and gene symbolization: None. Mutational events: cer-ys.680, -ys.1352 in Bonus; cer-1182, -ys.1247 in Kristina; cer-ys.1826 in Bonus. Mutant used for description and seed stocks: cer-ys.680 in Bonus (GSHO 1552); cer-ys.680 in Bowman (PI 483237)*3 (GSHO 2222).1521292GSHO 1552
193GSHO 1553cer-yt.938Hordeum vulgare L. subsp. vulgare Skåne län, SwedenNSGCKERNELS1995DEVELOPED58.98333300, 13.50000000Genetic materialSurface wax coating on the spike appears to be absent (wax code - ++ ++). In the Bowman backcrossd-derived line, nodes appear to lack surface wax. Inheritance: Monofactorial recessive. Location is unknown. Previous nomenclature and gene symbolization: None. Mutational events: cer-yt.758, -yt.938, -yt.1333 in Bonus; cer-yt.1127, -yt.1304 in Kristina, -yt.1427, -yt.1458, -yt.1493, -yt.1766, -yt.1822 in Bonus; cer-yt.1379 in Bonus. Mutant used for description and seed stocks: cer-yt.938 in Bonus (GSHO 1553); cer-yt.938 in Bowman (PI 483237)*4 (GSHO 2223).1521293GSHO 1553
194GSHO 1554cer-yu.158Hordeum vulgare L. subsp. vulgare Skåne län, SwedenNSGCKERNELS1995DEVELOPED58.98333300, 13.50000000Genetic materialSurface wax coating on the leaf blade appears to be absent only on upper leaves (wax code ++ ++ -). Mutant plants have slightly coiled awns. The Bowman backcross-derived line is weak in the field and has short leaves, but these traits are not as obvious in greenhouse grown plants. Inheritance: Monofactorial recessive. Location is unknown. Previous nomenclature and gene symbolization: None. Mutational events: cer-yu.158, -yu.159, -yu.174, -yu.992, -yu.1799 in Bonus, -yu.1168, -yu.1170 in Kristina. Mutant used for description and seed stocks: cer-yu.159 in Bonus (GSHO 1554); cer-yu.159 in Bowman (PI 483237)*3 (GSHO 2224).1521294GSHO 1554
195GSHO 1555cer-yx.421Hordeum vulgare L. subsp. vulgare Skåne län, SwedenNSGCKERNELS1995DEVELOPED58.98333300, 13.50000000Genetic materialSurface wax coating on the spike, the leaf sheath, and stem appears to be reduced (wax code + + ++). Inheritance: Monofactorial recessive. Location is unknown. Previous nomenclature and gene symbolization: None. Mutational events: cer-yx.421 in Foma, -yx.1226 in Kristin; cer-yx.1092, -yx.1094 in Kristina. Mutant used for description and seed stocks: cer-yx.421 in Foma (GSHO 1555); cer-yx.421 in Bowman (PI 483237)*2 (GSHO 2225).1521295GSHO 1555
196GSHO 1556Cer-yy.849Hordeum vulgare L. subsp. vulgare Skåne län, SwedenNSGCKERNELS1995DEVELOPED58.98333300, 13.50000000Genetic materialSurface wax coating on the spike appears to be absent (wax code - ++ ++). Concerning the chemical epicuticular wax composition, the Cer-yy mutant in Bonus determines the waxes lacking ( )-hydroxy-( )-diketones and having relatively increased proportions of primary alcohols. The distribution of the lipid classes is similar to that of wild type plants. Inheritance: Monofactorial dominant. Located in chromosome 1 HS [5S]. about 4.2 cM distal from the Hor2 (hordein 2, HorB) locus. Previous nomenclature and gene symbolization: Glaucous head=ge. Glossy spike 1=Gle1. Mutational events: Gle1.a in Cristal; Cer-yy.437 in Foma (CIho 11333), -yy.649, -yy.650, -yy.849, -yy.898, -yy.919, -yy.920, -yy.923, -yy.929, -yy.935, -yy.941, -yy.966, -yy.968, -yy.975, -yy.982, -yy.983, -yy.984 in Bonus; Cer-yy.974, -yy.1717 in Bonus.1521296GSHO 1556
197GSHO 1557cer-yz.1452Hordeum vulgare L. subsp. vulgare Skåne län, SwedenNSGCKERNELS1995DEVELOPED58.98333300, 13.50000000Genetic materialSurface wax coating on the spike, leaf sheath, and stem appears to be slightly reduced (wax code + + ++). Inheritance: Monofactorial recessive. Location is unknown. Previous nomenclature and gene symbolization: None. Mutational events: cer-yz.1452, -yz.1453 in Bonus. Mutant used for description and seed stocks: cer-yz.1452 in Bonus (GSHO 1557); cer-yz.1452 in Bowman (PI 483237)* 2(GSHO 2226).1521297GSHO 1557
198GSHO 1558cer-xa.585Hordeum vulgare L. subsp. vulgare Skåne län, SwedenNSGCKERNELS1995DEVELOPED58.98333300, 13.50000000Genetic materialSurface wax coating on the leaf blade appears to be absent only on upper leaves (wax code ++ ++ -). Inheritance: Monofactorial recessive. Location is unknown. Previous nomenclature and gene symbolization: None. Mutational events: cer-xa.585 in Foma, -xa.838 in Bonus (PI 189763), -xa.1113 in Kristina. Mutant used for description and seed stocks: cer-xa.585 in Foma (GSHO 1558); cer-xa.585 in Bowman (PI 483237)*2 (GSHO 2227).1521298GSHO 1558
199GSHO 1559cer-xb.943Hordeum vulgare L. subsp. vulgare Skåne län, SwedenNSGCKERNELS1995DEVELOPED58.98333300, 13.50000000Genetic materialSurface wax coating on the spike appears to be absent (wax code - ++ ++). In the Bowman backcross-derived line, the peduncle appears to lack surface wax. Inheritance: Monofactorial recessive. Location is unknown. Previous nomenclature and gene symbolization: None. Mutational events: cer-xb.943 in Bonus, -xb.1166 in Kristina. Mutant used for description and seed stocks: cer-xb.943 in Bonus (GSHO 1559); cer-xb.943 in Bowman (PI 483237(*4 (GSHO 2228).1521299GSHO 1559
200GSHO 1560cer-xc.1371Hordeum vulgare L. subsp. vulgare Skåne län, SwedenNSGCKERNELS1995DEVELOPED58.98333300, 13.50000000Genetic materialSurface wax coating on the spike, leaf sheath, and stem appears to be reduced (wax code + + ++). Inheritance: Monofactorial recessive. Location is unknown. Previous nomenclature and gene symbolization: None. Mutational events: cer-xc.1371, -xc.1457 in Bonus. Mutant used for description and seed stocks: cer-xc.1371 in Bonus (GSHO 1560); cer-xc.1371 in Bowman (PI 483237)*2 (GSHO 2229).1521300GSHO 1560
201GSHO 1561cer-xd.1455Hordeum vulgare L. subsp. vulgare Skåne län, SwedenNSGCKERNELS1995DEVELOPED58.98333300, 13.50000000Genetic materialSurface wax coating on the spike, leaf sheath, and stem appears to be reduced (wax code + + ++). The Bowman backcross-derived line has surface wax on nodes. Inheritance: Monofactorial recessive. Location is unknown. Previous nomenclature and gene symbolization: None. Mutational events: cer-xd.1455, -xd.1492 in Bonus. Mutant used for description and seed stocks: cer-xd.1455 in Bonus (GSHO 1561); cer-xd.1455 in Bowman (PI 483237)*2 (GSHO 2230).1521301GSHO 1561
202GSHO 1562ert-q.101Hordeum vulgare L. subsp. vulgare Skåne län, SwedenNSGCKERNELS1995DEVELOPED58.98333300, 13.50000000Genetic materialSpikes are semicompact and have slightly irregular placement of kernels. Seedlings leaves are slightly wider than normal, awns are about 3/4 normal length, and plants are semidwarf (3/4 of normal culm length). Inheritance: Monofactorial recessive. Located in chromosome 2H, based on linkage drag with the Gth (toothed lemma) locus. Previous nomenclature and gene symbolization: Erectoides-101=ert-101. Mutational events: ert-q.101 in Bonus; ert-q.392 (GSHO 491), -q.421 in Foma (CIho 11333). Mutant used for description and seed stocks: ert-q.101 in Bonus (GSHO 1562); ert-q.101 in Bowman (PI 483237)*7 (GSHO 1903).1521302GSHO 1562
203GSHO 1563abr1.aHordeum vulgare L. subsp. vulgare Skåne län, SwedenNSGCKERNELS1995DEVELOPED58.98333300, 13.50000000Genetic materialRachis internodes in the upper part of the spike are about 12% longer than normal. Secondary and tertiary shoots may form at the accordian-like basal rachis nodes. Manifestation of the extra shoots may be a day-length response and genotype dependent. Heading is delayed in the Bowman backcross-derived line. Recombination with the eog1.a (elongated outer glume) allele has not been observed, and recombination with the vrs1.a (six-rowed spike 1) allele is low. Inheritance: Monofactorial recessive. Located in chromosome 2HL, about 7.4 cM proximal from the fch15 (chlorina seedling 15) locus, and abouit 6.3 cM proximal from the Gth (toothed lemma) locus. Previous nomenclature and gene symbolization: Laxatum-3=lax-3. Mutational events: abr1.a (lax.3), abr1.b (lax.12) in Bonus, abr1.c (lax.337) in Foma (CIho 11333). Mutant used for description and seed stocks: abr1.a in Bonus (GSHO 1563); abr1.a in Bowman (PI 483237)*7 (GSHO 1887).1521303GSHO 1563
204GSHO 496ert-u.56Hordeum vulgare L. subsp. vulgare Skåne län, SwedenNSGCKERNELS1995DEVELOPED58.98333300, 13.50000000Genetic materialSpikes are slightly denser than those of the parent with a rachis internode length estimate of 2.7 mm, and culms are about 3/4 normal length. Plants have a brachytic-like pattern of growth. Spike density is decreased by GA3 treatment of plants as the flag leaf emerges. Awns are about 3/4 normal length, and the basal rachis internode is slightly elongated. Inheritance: Monofactorial recessive. Located in chromosome 2H, based on linkage drag with the Gth (toothed lemma) locus. Previous nomenclature and gene symbolization: Erectoides-56=ert-56. Brachytic 5=br5. Mutational events: ert-u.56 in Bonus. Mutant used for description and seed stocks: ert-u.56 in Bonus (GSHO 496); ert-u.56 in Bowman (PI 483237)*8 (GSHO 1904).1512450GSHO 496
205GSHO 1732eam9.1Hordeum vulgare L. subsp. vulgare North Dakota, United StatesNSGCKERNELS1995DONATED05/1995Genetic materialPlants are about 4 weeks earlier under short-day conditions. At Kurashiki, Japan, maturity and agronomic effects of the eam9.l allele are similar to those of the photoperiod insensitive gene eam8.k (early maturity 8). The eam9 is not allelic to several other early maturity genes which alter photoperiod sensitivity. Inheritance: Monofactorial recessive. Located in chromosome 4HL, over 17.4 cM distal from the Kap (hooded lemma) locus, and over 15.6 cm proximal from the glf3 (glossy leaf 3) locus. Previous nomenclature and gene symbolization: Early maturity c=ea,,c. Mutational events: eam9.l in Tayeh 8 (OUC34); Pao-an-chen 1 (OUC336), Pai-sha-pu 1 (OUC029), Mushin-chiang 3 (Mokusekko 3, OUC627, PI 420938), Yanghsin 2 (OUC327). Mutant used for description and seed stocks: eam9.l in Tayeh 8 (GSHO 1732).1005723GSHO 1732
206GSHO 1741flo-a.1Hordeum vulgare L. subsp. vulgare Skåne län, SwedenNSGCKERNELS1995DEVELOPED58.98333300, 13.50000000Genetic materialExtra bracts develop occasionally at the base of the central spikelet on the abaxial side. Fromation of the extra bracts is most common in the centeral portion of the spike, but rarely will the bracts form another spikelet. Inheritance: Monofactorial ressive. Location is unknown. Previous nomenclature: none. Mutational events: flo-a.1 in Foma(2). Mutant used for description and seed stocks: flo-a.1 in Foma (GSHO 1741); flo-a.1 in Bowman (PI 483237)*5(GSHO 2005).1545324GSHO 1741
207GSHO 908wax1.aHordeum vulgare L. subsp. vulgare JapanNSGCKERNELS1995DEVELOPEDGenetic materialThe endosperm contains primarily amylopectin, chains of alpha-(1-4) D-glucopyranose units branched through alpha-(1-6) linkages, and has a waxy texture. The starch content is reduced, the amount of sucrose and maltose is increased significantly, and more amylodextrin may be deposited. The starch in the endosperm and pollen grains has reddish-brown reaction to iodine. The estimated spontaneous reversion to normal for six wax1 alleles ranged from 1.9 to 5.6 x 10-5, and the intralocus recombination frequencies ranged from 0.4 to 194x10-5. Instability of the wax1.a allele from R.I Wolfe's Multiple Recessive Stock is induced by crossing to mutant line 152. Inheritance: Monofactorial recessive. Located in chromosome 7HS [1S], about 1.5 cM proximal from the fch12 (chlorina seedling 12) locus, adn about 12.6 cM proximal from the Run1 (reaction to Ustilago nuda 1) locus. Previous nomenclature and gene symbolization: Waxy endosperm=Wx. High amylopectin endosperm=glx. Mutational events: wax1.a in Murasaki Mochi (GSHO 84); wax1.b, wax1.c, wax1.d, wax1.e, wax1.f and wax1.g in Steptoe (CIho 15229). Mutant used for description and seed stocks: wax1.a in Odebrucker (CIho 4666)*10 (Waxy Oderbrucker, CIho 7563, GSHO 908); wax1.a from R.I. Wolfe's Multiple Recessive Stock in Bowman (PI 483237)*8 (GSHO 1828).1513700GSHO 908
208GSHO 988Blp1.bHordeum vulgare L. subsp. vulgare Utah, United StatesNSGCKERNELS1995DEVELOPEDGenetic materialBlack pigmentation of the lemma and pericarp develops slightly before maturation of the spike. Pigmented organs may include all parts of the spike, awns, the upper portion of the stem, and upper leaves. The intensity of pigmentation associated with each of the dominant alleles at the Blp locus is characteristic of that allele, and is relatively stable over environments. Black seed is produced by melanin-like pigment in the pericarp. Woodward reports that the dominance ranking of alleles at the Blp locus is related to the intensity of black pigmentation they confer with the Blp.1.b (B) allele conferring extreme black pigmentation. The Blp1.mb (Bmb) allele is associated with medium black and a reduced distribution pattern; and the Blp1.g (Bg) allele is associated with light black or gray coloration. Inheritance: Monofactorial dominant. Located in chromosome 1HL [5L], about 16.0 cM proximal from the trd (third outer glume) locus. Previous nomenclature and gene symbolization: Black lemma and caryopis=B. Black pericarp=Bk. Black lemma and pericarp=B. Mutational events: Blp1.b (B) in Hordeum distichon var nigrinudum No 1; Blp1.mb (Bmb) in CIho 2970 (GSHO 226); Blp1.g (Bg) in Blackhull (CIho 878, GSHO 199) and Black smyrna (CIho 191, GSHO 222). Mutant used for description and seed stocks: Blp1.b in Hordeum distichon var nigrinudum No 1 (GSHO 988); Blp1.b from R.I. Wolfe's Multiple Dominant Marker Stock (GSHO 1580) in Bowman (PI 483237)*6 (GSHO 2054).1513889GSHO 988
209GSHO 1324Run1.aHordeum vulgare L. subsp. vulgare Colorado, United StatesNSGCKERNELS1995DONATED05/1994Genetic materialA dominant gene which controls a resistant reaction to "non-Trebi-attacking" cultures of Ustilago nuda and to cultures capable of infecting the "Jet" type of resistance. Inheritance: Monofactorial dominant. Located in chromosome 7HS[1S], about 2.0 cM proximal from the Rpg1 (reaction to Puccinia graminis 1) locus, about 12.7 cM distal from the wax (waxy endosperm) locus, and about 15.1 cM distal from the nud (naked caryopsis) locus. Previous nomenclature and gene symbolization: Resistance to Ustilago nuda=Un. Mutational events: Run1.a in Trebi. Mutant used for description and seed stocks: Run1.a in Trebi (GSHO 1324).1514223GSHO 1324
210GSHO 536lbi3.cHordeum vulgare L. subsp. vulgare Ontario, CanadaNSGCKERNELS1995DEVELOPEDGenetic materialMutant plants in the cultivar Montcalm have a marked elongation and weakness of the basal rachis internode of the spike. This region may be 10 to 13 cm long in some tillers. The spike hangs vertically downward from the collar on emergence from the sheath and is often broken off in this region by the wind. When not broken off, spikes have normal fertility, contain well-filled grain, and show normal maturity. Expression of the lbi3.c allele in the Bowman-derived line is limited to a slight elongation of the basal rachis internode and a slightly lax spike. Inheritance: Monofactorial recessive. Located in chromosome 7HL [1L], over 20.0 cM distal from the nud (naked caryopsis) locus. Precivous nomenclature and gene symbolization: Long basal internode 3=lb3. Mutational events: lbi3.c (Alb Acc 291), lbi3.d (Alb Acc 292), lbi3.e (Alb Acc 293), lbi3.m (Alb Acc 290) in Montcalm. Mutant used for description and seed stocks: lbi3.c in Montcalm (GSHO 536); lbi3.c in Bowman (PI 483227)*7 (GSHO 1840).1512187GSHO 536
211GSHO 585abo13.pHordeum vulgare L. subsp. vulgare Montana, United StatesNSGCKERNELS1995DEVELOPEDGenetic materialSeedlings have a white color and die at the two to three leaf stage. The abo13.p gene must be maintained as a heterozygous stock. Inheritance: Monofactorial recessive. Located in chromosome 2HL, about 1.2 cM from the msg 2 (male sterile genetic 2) locus. Previous nomenclature and gene symbolization: Albino seedling e=alb,,e. Albino seedling p=alb,,p. Mutational events: abo13.p in Titan; 12 additional mutants isolated in the progeny of diethyl sulfate treated msg2 plants (generated from a cross to a balanced tertiary trisomic stock) crossed to Arivat (CIho 7543). Mutant used for description and seed stocks: abo13.p in Titan (GSHO 585).1512272GSHO 585
212GSHO 629fst1.aHordeum vulgare L. subsp. vulgare JapanNSGCKERNELS1995DEVELOPEDGenetic materialLeaves and stems of mutant plants are easily broken when physical bent. This extraordinary fragility is exhibited even after maturity. Generally, stems are more flexible than in normal plants, and straw breakage prior to harvest is rare. When the fst1.a is present in a line, straw collapse and/or lodging occurs more frequently. Forage yield is reduced slightly by the fst1.a allele. Inheritance: Monofactorial recessive. Located in chromosome 5HL [7L], about 24.3 cM proximal from the srh (short rachilla hair) locus. Previous nomenclature and gene symbolization: Fragile stem 1=fs1. Mutational events: fst1.a in Kamairazu (OUL008); fst1.d probably originating from Kamairazu in a Western Plant Breeders selection. Mutant used for description and seed stocks: fst1.a in Kamairazu (GSHO 629); fst1.a in Bowman (PI 483237)*5 (GSHO 2100); fst1.d in Bowman*7 (GSHO 2101).1512509GSHO 629
213GSHO 785raw5Hordeum vulgare L. subsp. vulgare JapanNSGCKERNELS1995DEVELOPEDGenetic materialo-r,,e, orange lemma smooth awn, (sterility).1514245GSHO 785
214GSHO 985Kap1.aHordeum vulgare L. subsp. vulgare Colorado, United StatesNSGCKERNELS1995DONATED05/1994Genetic materialThe hooded ("Kapuze") trait is chraracterized by an appendage to the lemma, which develops as a trifurcate structure consisting of a deformed floret at its center with two triangular leaf-like projections called lemma wings. The supernumerary floret often contains stamens with fertile pollen grains and occasionally bears a kernel within it. The ectopic expression of the Kap1.a allele forms the extra floret and is associated with the presence of a 305-base pair duplication in intron 4 of the Knox3 sequence. Inheritance: Monofactorial dominant. Located in chromosome 4HS, about 25.1 cM proximal from the glf3 (glossy leaf 3) locus. Previous nomenclature and gene symbolization: Hooded lemma=K. Hooded lemma=A. Hooded=C1. Mutational events: Kap1.a in an unknown cultivar; Kap1.e (Ke, elevated hooded) a mutant of the Kap1.a allele in an unknown cultivar (GSHO 763, Chengchou 5). Mutant used for description and seed stocks: Kap1.a in Colsess (GSHO 985); Kap1.a from Strip Tease (CIho 6837) in Betzes (PI 129430)*7 (PI 533600, Haybet), Kap1.a from CIho 2220 in Compana (PI 537442)*7 (PI 534506); Kap1.a from Wolfe's Multiple Dominant Marker Stock in Bowman (PI 483237)*8 (GSHO 2007).1513904GSHO 985
215GSHO 1065als1.aHordeum vulgare L. subsp. vulgare Ontario, CanadaNSGCKERNELS1994DEVELOPEDGenetic materialLateral spikelets at the base of the spike fail to develop or are partially developed. Tillers are large, coarse, and stiff, and only 1 or 2 tillers are produced in the six-rowed stock. The plant resembles the uniculm mutant. Plants of the Bowman backcross-derived line commonly produce 3 to 5 tillers. Inheritance: Monofactorial recessive. Located in chromosome 3HL, about 31.2 cM distal from the uzu (uzu) locus, and about 39.7 cM proximal from the cur2 (curly 2) locus. Previous nomenclature and gene symbolization: None. Mutational events: als1.a in Montcalm (Alb Acc 281). Mutant used for description and seed stocks: als1.a in Montcalm (GSHO 1065); als1.a in Bowman (PI 483237)*7 (GSHO 1990).1513839GSHO 1065
216GSHO 107fch2.cHordeum vulgare L. subsp. vulgare Colorado, United StatesNSGCKERNELS1994DONATED05/1994Genetic materialSeedlings have a pale yellow-green color, which persists until near maturity under greenhouse or field conditions. Development of homozygous is delayed, but the stunted plants survive to produce seed. A complete absence or greatly reduced levels of chlorophyll b were found in mutant plants. Respiratory and photosynthetic rates of detached leaves and intact plants were not significantly different from those of normal plants. However, the growth rate of mutant plants decreased after the endosperm was depleted. An absence of chlorophyll b-binding proteins in the photosynthetic membranes was found in fch2 mutants. Inheritance: Monofactorial recessive. Located in chromosome 3HL, about 7.7 cM distal from the cur2 (curly 2) locus. Previous nomenclature and gene symbolization: Chlorina seedling 2=f2. Light green 5=lg5. Chlorina seedling f2=clo-f2. Mutational events: fch2.c in 28-3398 Chlorina; fch2.y (lg5) in CIho 6151; clo-f2.2800, -f2.2807, -f2.3613 in Donaria (PI 209784), -f2.101, -f2.102, -f2.103, -f2.105, -f2.107, -f2.108, -f2.109 in Tron, -f2.122, -f2.123, -f2.133 in Bonus (PI 189763). Mutant used for description and seed stocks: fch2.c in 28-3398 (GSHO 107); fch2.c in Bowman (PI 483237)*7 (GSHO 1993).1510706GSHO 107
217GSHO 112fch1.aHordeum vulgare L. subsp. vulgare Colorado, United StatesNSGCKERNELS1994DONATED05/1994Genetic materialSeedling and immature leaves are very pale yellow-green (chlorophyll deficient). The leaves gradually change to a green color beginning with the leaf tip. Plants may grow to maturity in the field, but they have a slightly yellow-green color and are stunted easily by stress. Inheritance: Monofactorial recessive. Located in chromosome 2HS, in the centromeric region and about 4.0 cM distal from the eog (elongated outer glume) locus. Previous nomenclature and gene symbolization: Chlorina seedling=f f. Light green=lg. Light green 8=lg8. Light green 10=lg10. Mutational events: fch1.a in Gull; fch1.p (lg) (Minn 80, GSHO 548); fch1.q (lg8) (No 155, GSHO 8); and fch1.r (lg10)(No 157, GSHO 154) in Himalaya (CIho 1312). Mutant used for description and seed stocks: fch1.a in Minn 84-7 (GSHO 112); fch1.a in Bowman (PI 483237)*9 (GSHO 1886).1510685GSHO 112
218GSHO 1130msg3.ccHordeum vulgare L. subsp. vulgare Ontario, CanadaNSGCKERNELS1994DEVELOPEDGenetic materialSelfing-none. Outcrossing-complete female fertility. Stamens-anthers much smaller than fertile sib, no stomium or filament elongation. Pollen-non-staining, no free pollen grains. The male sterile plants are also about 1/3 normal size with short, wide leaves. Spikes are dense and spikelets are small and malformed, awns are 1/3 normal length, and double or triple pistils (fasciation) occur in some spikelets. Inheritance: Monofactorial recessive. Located in chromosome 2HS, about 4.6 cM distal from the eog (elongated outer glume) locus, and 0.6 cM distal from the fch1 (chlorina seedling 1) locus. Previous nomenclature and gene symbolization: Male sterile 10=ms10. Male sterile 3=ms3. Mutational events: msg3.cc in Gateway (MSS051). Mutant used for seed stocks: msg3.cc in Gateway (GSHO 1130); msg3.cc in Bowman*7 (GSHO 1885).1514181GSHO 1130
219GSHO 1140yst1.aHordeum vulgare L. subsp. vulgare Ontario, CanadaNSGCKERNELS1994DEVELOPEDGenetic materialLongitudinal yellow stripes appear on seedling leaves and persist until the preripening stage. In some crosses and under certain environmental conditions, the most heavily striped F2 plants fail to reach maturity. There is no evidence for a cytoplasmic association in the inheritance patterns. Expression of the yst1.a allele in the Bowman backcross-derived line is reduced to very narrow streaks that occur infrequently. Inheritance: Monofactorial recessive. Located in chromosome 3HS, about 2.9 cM proximal from the zeb1 (zebra stripe 1) locus, and about 7.6 cM proximal from the uzu (uzu) locus. Previous nomenclature and gene symbolization: Yellow stripe=ys. Yellow streak=yst. Mutational events: yst1.a in Gateway. Mutant used for description and seed stocks: yst1.a in Gateway (GSHO 1140); yst1.a in Bowman (PI 483237)*7 (GSHO 1959).1514332GSHO 1140
220GSHO 115nud1.aHordeum vulgare L. subsp. vulgare Colorado, United StatesNSGCKERNELS1994DONATED05/1994Genetic materialThe lemma and palea do not adhere to the caryopsis and the grain will thresh free of the hull at maturity. The naked caryopsis trait is expressed in all environments. Inheritance: Monofactorial recessive. Located in chromosome 7HL{1L]; near the centromere, about 9.6 cM proximal from the lks2 (short awn 2) locus, and about 10.5 cM proximal from the dsp1 (dense spike 1) locus. Previous nomenclature and gene symbolization: Naked caryopsis=k. Naked caryopsis=s. Naked caryopsis=n. Hulless=h. Mutational events: nud1.a in Himalaya (CIho 1312); nud1.b in Haisa (Mut 4129), nud1.c (Mut 3041/62) in Ackermann's Donaria (PI 161974). Mutant used for description and seed stocks: nud1.a in Himalaya (GSHO 115), nud1.a from Sermo (CIho 7776) in Betzes (PI 129430)*7 (CIho 16559, GP 37), nud1.a from Sermo in Compana (CIho 5438)*7(CIho 16185, GP 41), nud1.a from Sermo in Decap (CIho 3351)*7 (CIho 16563, GP 45); nud1.a from Stamm (PI 194555) in Betzes*7 (CIho 16566, GP 48), nud1.a from Stamm in Compana* (CIho 16183, GP 50), nud1.a from Stamm*7 in Freja (CIho 7130)*7(CIho 16568, GP 52); nud1.a from Wolfe's Multiple Recessive Stock 1 in Bowman (PI 483237)*8 (GSHO 1847).1510623GSHO 115
221GSHO 117xnt6.fHordeum vulgare L. subsp. vulgare Colorado, United StatesNSGCKERNELS1994DEVELOPEDGenetic materialSeedlings have a bright yellow (citron green) to yellow-green color and do not survive beyond the seedling stage. The xnt6.f gene must be maintained as a heterozygous stock. Inheritance: Monofactorial recessive. Located in chromosome 3HS, over 15.5 cM distal from the abo9 (albino seedling 9) locus, over 15.3 cM distal from the msg5 (male sterile genetic 5) locus, and over 22.0 cM distal from the wst1 (white streak 1) locus. Previous nomenclature and gene symbolization: Xantha seedling s=xs. Mutational events: xnt6.f in Smyrna (Smyrna I). Mutant used for description and seed stocks: xnt6.f in Smyrna (GSHO 117).1510619GSHO 117
222GSHO 1211crl1.aHordeum vulgare L. subsp. vulgare Ontario, CanadaNSGCKERNELS1994DEVELOPEDGenetic materialAwns on lateral spikelets are curly or wavy, and lateral spikelets, and occasionally central spikelets, may be malformed and partially sterile. The curly lateral trait is not expressed in all tillers. In the Bowman backcross-derived line, awns seem thinner and the awn of a few spikelets is bent at an odd angle. Inheritance: Monofactorial recessive. Location is unknown; originally reported to be in 5HL [7L] over 30.4 cM distal from the raw1 (smooth awn 1) locus, but linkage to the Rph12 (reaction to Puccinia hordei 12) locus was not found. Previous nomenclature and gene symbolization: Curly laterals=cu. Curly laterals=cl. Mutational events: crl1.a in Montcalm (Alb Acc 305). Mutant used for description and seed stocks: crl1.a in Montcalm (GSHO 1211); crl1.a in Bowman (PI 483237)*3 (GSHO 2146).1513702GSHO 1211
223GSHO 1212clh1.aHordeum vulgare L. subsp. vulgare Arizona, United StatesNSGCKERNELS1994DEVELOPEDGenetic materialLeaf blades are erect, narrow and folded inward and appear thicker than normal. Plants show reduced vigor, fertility, and height (3/4 normal height) when grown in the greenhouse; further reduction in vigor and fertility were observed when mutants are grown in the field. Spikes emerge poorly from the boot. Inheritance: Monofactorial recessive. Located in chromosome 1HL [5L], about 7.0 cM from the msg1 (male sterile genetic 1) locus. Previous nomenclature and gene symbolization: None. Mutational events: clh1.a in Hannchen. Mutant used for description and seed stocks: clh1.a in Hannchen (GSHO 1212); clh1.a in Bowman (PI 483237)*2 (GSHO 2051).1513958GSHO 1212
224GSHO 1214fch4.gHordeum vulgare L. subsp. vulgare Ontario, CanadaNSGCKERNELS1994DEVELOPEDGenetic materialSeedlings are bright yellow-green on emergence and maintain this color until they are nearly mature. Seedlings and plants are relatively vigorous, but heading is delayed. Inheritance: Monofactorial recessive. Located in chromosome 7HL [1L], slightly distal from the nud (naked caryopsis) locus and about 4.4 cM proximal from the lks2 (short awn 2) locus. Previous nomenclature and gene symbolization: Chlorina seedling 4=f4. Yellow viable=yv. Mutational events: fch4.g in Montcalm (Alb Acc 258). Mutant used for description and seed stocks: fch4.g in Montcalm (GSHO 1214); fch4.g in Bowman (PI 483237)*6 (GSHO 1851).1513783GSHO 1214
225GSHO 1215fch5.fHordeum vulgare L. subsp. vulgare Ontario, CanadaNSGCKERNELS1994DEVELOPEDGenetic materialSeedlings are pale yellow to ivory in color at emergence and remain this color until they are nearly mature. Seedlings are relatively weak and rarely survive to maturity in the field at Fargo, North Dakota, USA; however, in the greenhouse they gradually become darker green and produce late-maturing plants. Inheritance: Monofactorial recessive. Located in chromosome 7HS {1S], proximal from the brh1 (brachytic 1) locus, about 25.9 cM proximal from the Rpg1 (reaction to Puccinia graminis tritici 1) locus. Previous nomenclature and gene symbolization: Chlorina seedling 5=f5. Yellow viable=yv2. Mutational events: fch5.f in Gateway (Alb Acc 328). Mutant used for description and seed stocks: fch5.f in Gateway (GSHO 1215); fch5.f in Bowman (PI 483237)*5(GSHO 1827).1513694GSHO 1215
226GSHO 1217snb1.aHordeum vulgare L. subsp. vulgare Ontario, CanadaNSGCKERNELS1994DEVELOPEDGenetic materialA glume-like or stick-like bract arises immediately under the node-base (below and between the outer glumes) of the central spikelets. The bracts are present at only a few internodes on randomly arranged central spikelets. Not all spikes on affected plants have the extra bract. Inheritance: Monofactorial recessive. Located in chromosome 7HS [1S], near the centromere, about 36.3 cM proximal from the brh1 (brachytic 1) locus. Previous nomenclature and gene symbolization: Subnodal bract=sb. Mutational events: snb1.a in L50-200 (Alb Acc 67A), a recessive allele at the brh1 locus is present in this stock also. Mutant used for description and seed stocks: snb1.a in L50-200 (GSHO 1217).1514081GSHO 1217
227GSHO 1224nec2.bHordeum vulgare L. subsp. vulgare Denmark Historic1994DEVELOPEDGenetic materialSmall, dark brown necrotic spots appear on the leaves of field grown plants 3 to 6 weeks after germination. Margins of the spots have poor demarcation, and the expression of necrotic spot is not good some years. Inheritance: Monofactorial recessive. Located in chromosome 6HS, about 5.2 cM from the rob (orange lemma) locus. Previous nomenclature and gene symoblization: Mutant no 7. Necrotic leaf spots 2=nec2.b. Mutational events: nec2.b (Mutant no 7), nec2.c (Mutant no 9) in Carlsberg II. Mutant used for description and seed stocks: nec2.b in Carlsberg II (GSHO 1224).1513959GSHO 1224
228GSHO 1232dsp1.aHordeum vulgare L. subsp. vulgare Colorado, United StatesNSGCKERNELS1994DONATED05/1994Genetic materialSpike length is reduced because rachis internode length is reduced to about 2/3 normal. The reduction in rachis internode length caused by the dsp1.a and uzu1.a (uzu) alleles is similar and additive. In F2 progenies, the dsp1.a gene has pleiotropic effects on coleoptile length, culm length, and grain size. Inheritance: Monofactorial recessive. Located in chromosome 7HS [1S], about 10.5 cM distal from the nud (naked caryopsis) locus. Previous nomenclature and gene symbolization: Dense spike=l. Lax spike=L1. Short spike=e. Mutational events: dsp1.a in many cultivars of Oriental origin, often associated with the short awn gene. Mutant used for description and seed stocks: dsp1.a in Honen 6 (OUJ469, PI 307495, GSHO 1232); dsp1.a from DWS 1081 in Bowman (PI 483237)*7 (GSHO 1833).1513803GSHO 1232
229GSHO 1277var3.cHordeum vulgare L. subsp. vulgare Ontario, CanadaNSGCKERNELS1994DEVELOPEDGenetic materialWhite stripes appear on the foilage and stems, but they are less extensive than the white streak 7 (wst7) type of streaks. Expression of var3.c is poor in the Bowman backcross-derived line. Inheritance: Monofactorial recessive. Located in chromosome 5HL [7L], about 29.7 cM distal from the raw1 (smooth awn 1) locus; Karlsson was unable to confirm the presence of va3 in chromosome 5H. Previous nomenclature and gene symbolization: Variegated 3=va3. Mutational events: var3.c in Montcalm (Alb Acc 306). Mutant used for description and seed stocks: var3.c in Montcalm (GSHO 1277); var3.c in Bowman (PI 483237)*5 (GSHO 2143).1514403GSHO 1277
230GSHO 1278var1.aHordeum vulgare L. subsp. vulgare Ontario, CanadaNSGCKERNELS1994DEVELOPEDGenetic materialWhite stripes on young leaves are less extensive and well-defined than those of white streak 7 (wst7). White stripes appear on the foilage and stems of older plants. Plots grown form homozygous lines have a whitish cast until heading. Inheritance: Monofactorial recessive. Located in chromosome 5HL [7L], about 4.6 cM proximal from the raw1 (smooth awn 1) locus. Previous nomenclature and gene symbolization: Variegated=va. Mutational events: var1.a in Montcalm (Alb Acc 311). Mutant used for description and seed stocks: var1.a in Montcalm (GSHO 1278); var1.a in Bowman (PI 483237)*7 (GSHO 2121).1513963GSHO 1278
231GSHO 1279zeb1.aHordeum vulgare L. subsp. vulgare Minnesota, United StatesNSGCKERNELS1994DEVELOPEDGenetic materialSeedlings may have pale green to yellow bands across the leaves when germinated under cold conditions in the field. Cold room studies showed that development of a mutant phenotype in seedlings could be induced by 10 to 15 days of cold treatment (0 to 1 C) if initiated within one day after wetting of the seed. Alternating cold and warm temperatures in the field induced diagnostic symptoms. Inheritance: Monofactorial recessive. Located in chromosome 3HL, about 7.0 cM proximal from the uzu (uzu) locus. Previous nomenclature and gene symbolization: Zebra stripe=zb. Mutational events: zeb1.a in Mars (C3-1). Mutant used for description and seed stocks: zeb1.a in Mars (GSHO 1279); zeb1.a in Bowman (PI 483237)*3 (GSHO 1964).1513696GSHO 1279
232GSHO 1297lks5.fHordeum vulgare L. subsp. vulgare Montana, United StatesNSGCKERNELS1994DEVELOPEDGenetic materialAwns on both central and lateral spikelets are reduced to 1/4 or less of normal length in six-rowed cultivars. Only the central spikelets exhibit reduced awn length in two-rowed cultivars. The rachilla is often modified, and in extreme cases the rachilla may develop as an malformed additional floret. Awns are thin and brittle. Inheritance: Monofactorial recessive. Located chromosome 4HL, about 6.9 cM distal from the Kap (hooded lemma) locus. Previous nomenclature and gene symbolization: Short awn 5=lk5. Breviaristatum-2=ari-2. Breviaristatum-c=ari-c.2. Mutational events: lks5.f in CIho 5641; lks5.g (Kmut 218) in Ashai 5 (OUJ509); ari-c.2, -c.59 in Bonus (PI 189763), -c.103 in Foma (CIho 11333); ari-c.106 in Foma; ari-c.109, -c.110, -c.111, -c.112, -c.120, -c.139, -c.157, -c.159, -c.179, -c.180, -c.199, -c.201, -c.203, -c.204, -c.206, -c.210, -c.229 in Foma, -c.259, -c.262, -c.272, -c.276, -c.289, in two-rowed Glacier and a mutant in Morex (CIho 15773), have been reported. Mutant used for description and seed stocks: lks5.f in CI5641 (GSHO 1297); ari-c.2 in Bonus (GSHO 1651); lks5.f in Bowman (PI 483237)*6 (GSHO 2014); ari-c.2 in Bowman*7 (GSHO 2013).1514379GSHO 1297
233GSHO 1300uzu1.aHordeum vulgare L. subsp. vulgare Colorado, United StatesNSGCKERNELS1994DONATED05/1994Genetic materialThe uzu1.a gene has pleiotropic effects on the elongation of the coleoptile, leaf, culm, rachis internode, awn, glume, and kernel. These organs are often reduced in length and increased in width. Changes in organ length are temperature sensitive, but heading date and maturity are unaltered. The coleoptile of uzu plants shows a prominent projection or hook near the apex. Sometimes the coleoptile of the mutant shows a V-shaped notch on the side opposite from the projection. Thus, the apex of the coleoptile has two notches, one on each side. The temperature sensitive reduction in culm length associated with uzu1.a gene produces plants ranging from less than 1/4 normal in warm environments to 5/6 normal in cool ones. Inheritance: Monofactorial recessive. Located in chromosome 3HL, about 17.6 cM proximal from the alm (albino lemma) locus. Previous nomenclature and gene symbolization: Normal vs uzu=h. Uzu=u. Uzu (semi-brachytic)=uz. Uzu 2=uz2. Uzu 3=uz3. Mutational events: uzu1.a in many Japanese cultivars. Mutant used for description and seed stocks: uzu1.a in Baitori 11 (OUJ371, PI 182624, GSHO 1300); uzu1.a in Bowman (PI 483237)*7 (GSHO 1963).1513817GSHO 1300
234GSHO 1313Rph1.aHordeum vulgare L. subsp. vulgare Colorado, United StatesNSGCKERNELS1994DONATED05/1994Genetic materialThe seedling reaction type is O-n, -1c with race 4 culture 57-19; heterozygotes have reactiontypes ranging from 1 to 3, depending on parents. Responses will vary for homozygotes and heterozygotes when different rust cultures are tested. Inheritance: Monofactorial incomplete dominant. Located in chromosome 2H, probably in the short arm. Previous nomenclature and gene symbolization: Resistance to Puccinia hordei=Pa1. Resistance to Puccinia hordei C=C. Resistance to Puccinia hordei Otth=Pa. Resistance to Puccinia hordei Otth=Rph1a. Mutational events: Rph1.a in Manchuria; Rph1.a in Cape (PI 39403), OAC 21 (CIho 1470); Rph1.a in Oderbrucker (CIho 940); Rph1.a in Sudan (CIho 6489), Speciale (CIho 7536). Mutant used for description and seed stocks: Rph1.a in Oderbrucker (GSHO 1313).1514111GSHO 1313
235GSHO 1314Rph4.dHordeum vulgare L. subsp. vulgare Colorado, United StatesNSGCKERNELS1994DONATED05/1994Genetic materialThe seedling reaction type is 0-n - 1c with race 4 culture 57-19. Heterozygotes may have a type 2 or 3 reaction under certain conditions. Inheritance: Monofactorial dominant. Located in chromosome 1HS [5S], about 16.0 cM distal from the Reg1 or mla (reaction to Erysiphe graminis 1) locus. Previous nomenclature and gene symbolization: Resistance to Puccinia hordei D=D. Resistance to Puccinia hordei Otth 4=Pa4. Mutational events: Rph4.d in Gull, Lectaler; Franger (CIho 8811). Mutant used for description and seed stocks: Rph4.d in Gull (GSHO 1314); Rph4.d in Bowman (PI 483237)*2 (GSHO 2322).1513967GSHO 1314
236GSHO 1315Ryd2.bHordeum vulgare L. subsp. vulgare Colorado, United StatesNSGCKERNELS1994DONATED05/1994Genetic materialAn intermediate reaction to BYDV that limits expression of dwarfing and yellow discoloration of leaves. Plants with the Ryd2.b gene show less yield and seed size reduction when infected with BYDV and under drought and heat stress. Inheritance: Monofactorial incomplete dominant. Located in chromosome 3HL, about 15.9 cM proximal from the dsp10 (dense spike 10) locus. Previous nomenclature and gene symbolization: Resistance to BYDV 2=Yd2. Resistance to BYDV 2=Ryd2. Mutational events: Ryd2.b in CIho 1227, CIho 1237, CIho 2376, and CIho 14119 (four accessions from Ethiopia). Mutant used for description and seed stocks: Ryd2.b in Abyssinian (CIho 2376, GSHO 1315); Ryd2.b from CIho 2376 in Glenn (CIho 15769)*8 (GSHO 2370); Ryd2.b from CIho 2376 in Bowman (PI 483237)* 7 (GSHO 1969).1514446GSHO 1315
237GSHO 1316Early maturityHordeum vulgare L. subsp. vulgare Colorado, United StatesNSGCKERNELS1994DONATED05/1994Genetic materialThis gene controls the strong response of some cultivars to photoperiod. The Eam1 gene is described as a simply inherited dominant and the main factor for early maturity. Under long-day condition, a near-isogenic line with Eam1 is reported to be 7 to 8 days earlier than the eam1 line 'ea1'. The early maturity gene is weakly linked to the vrs1 (six-rowed spike 1) locus. In North Dakota, USA, progenies from crosses of Bowman to Hordeum vulgare subsp spontaneum and Coast, facultative winter, northern spring (from Finland and Alaska), and winter types of barley, extremely early segregates are found. A linkage between early maturity and malformed spikes (com2) is observed when an accession from the ICARDA-CIMMYT collection of barley freaks is backcrossed to Bowman. Positive linkage data are found also in progenies from some crosses between a line having the com2 - eam1 recombinant and early segregates from the above crosses. The Eam1 locus in barley is likely homoeologous to the Ppd1, Ppd2, and Ppd3 loci in wheat, which control strong photoperiod response.1513721GSHO 1316
238GSHO 1317Rsg1.aHordeum vulgare L. subsp. vulgare Colorado, United StatesNSGCKERNELS1994DONATED05/1994Genetic materialResistant seedlings infested with greenbugs (aphids) are not killed eight weeks after infestation by the buildup of the greenbug population as are susceptible seedlings. Inheritance: Monofactorial dominant. Located in chromosome 7H [1]. Previous nomenclature and gene symbolization: Greenbug resistance=Grb. Resistance to Schizaphis graminum Rondani (greenbug)=Rsg,,a. Mutational events: Rsg1.a in Bozu Omugi, Derbent, Kearney, Dobaku (PI 87817), and CIho 5087 (PI 82683). Mutant used for description and seed stocks: Rsg1.a in Bozu Omugi (GSHO 1317).1514404GSHO 1317
239GSHO 1318Rph7.gHordeum vulgare L. subsp. vulgare Colorado, United StatesNSGCKERNELS1994DONATED05/1994Genetic materialThe seedling reaction type is 0-n - 1c. The resistant reaction associated with the Rph7.g allele is not expressed well above 20 degrees Celsius. Cebada Capa is indistinguishable from the cultivar Forrajera Klein. Inheritance: Monofactorial dominant. Located in chromosome 3HS, linkage to markers in the centromeric region was reported, and about 24.0 cM from the ant17 (anthocyanin-less 17) locus. Previous nomenclature and gene symbolization: Resistance to Puccinia hordei Otth y=Pay. Resistance to Puccinia hordei Otth 5=Pa5. Resistance to Puccinia hordei Otth 7=Pa7, Mutational events: Rph7.g in Cebada Capa; Rph7.g in France 7 and France 21; Rph7.g in Dabat, Gonda, and La Estanzuela; Rph7.ac in Tu17a, a Bowman backcross-derived line from Tunisia. Mutant used for description and seed stocks: Rph7.g in Cebada Capa (GSHO 1318); Rph7.g in Bowman (PI 483237)*8 (GSHO 1994).1513725GSHO 1318
240GSHO 132Pvc1.aHordeum vulgare L. subsp. vulgare Colorado, United StatesNSGCKERNELS1994DONATED05/1994Genetic materialPurple pigment is produced by the presence of anthocyanins. When Pvc is expressed, purple pigmentation of the kernel is confined mainly to the lemma veins. The pigments are formed late during grain fill when the lemma is exposed to sunlight, and they tend to fade as the spike matures. Inheritance: Monofactorial dominant, but Buckley suggested that three pairs of genes controlled expression of purple pigmentation of lemma veins. Located in chromosome 2HL, about 22.2 cM distal from the vrs1 (six-rowed spike 1) locus. Previous nomenclature and gene symbolization: Purple veined lemma=C. Purple veined lemma=Pc. Mutational events: Pvc1.a in Buckley 2223-6. Mutant used for description and seed stocks: Pvc1.a in Buckley 2223-6 (GSHO 132); Pvc1.a from R.I. Wolfe's Multiple Dominant Stock in Bowman (PI 483237)*7 (GSHO 1929).1510814GSHO 132
241GSHO 1326glo-b.1001Hordeum vulgare L. subsp. vulgare Germany Historic1994DEVELOPEDGenetic materialKernels are globe-shaped, plants are stunted (2/3 normal height), and 1000-kernel weights are low. Plants in the Bowman backcross-derived line have some brachytic traits (short culms, leaf blades, and awns) and slightly coiled leaves and awns. Inheritance: Monofactorial recessive. Located in chromosome 5HL [7L], over 31.9 cM proximal from the ari-e (breviaristatum-e) locus. Previous nomenclature and gene symbolization: None. Mutational events: glo-b.1001 (1345/72, GSHO 1327) in Villa, glo-b.1002 (916/65) in Donaria (PI 161974); glo-b.1005 (859/74) in Villa; glo-b.1007 (1296/65) in Donaria; glo-b.1011 (1786/67) in Proctor (PI 280420); glo-b.1012 (1055/74) in Villa. Mutant used for description and seed stocks: glo-b.1002 in Donaria (GSHO 1326); glo-b.1002 in Bowman (PI 483237)*7 (GSHO 2095).1513735GSHO 1326
242GSHO 1328glo-a.1003Hordeum vulgare L. subsp. vulgare GermanyNSGCKERNELS1994DEVELOPEDGenetic materialFertile spikelets are shortened and the resulting kernels are nearly round or globe-shaped. Sterile lateral spikelets are 1/2 normal length and twisted. In the Bowman backcross-derived line, the glo-a.1003 allele reduces kernel size, test weight, and yield. Inheritance: Monofactorial recessive. Located in chromosome 4HS. Previous nomenclature and gene symbolization: None. Mutational events: glo-a.1003 (1343/63) in Proctor. Mutant used for description and seed stocks: glo-a.1003 in Proctor (GSHO 1328); glo-a.1003 in Bowman (PI 483237)*7 (GSHO 2006).1513703GSHO 1328
243GSHO 1329glo-c.1004Hordeum vulgare L. subsp. vulgare GermanyNSGCKERNELS1994DEVELOPEDGenetic materialThe kernels are more rounded or globe-shaped than in the parental cultivar. The spike is very lax with long rachis internodes. Inheritance: Monofactorial recessive. Located in chromosome 2H. Previous nomenclature and gene symbolization: None. Mutational events: glo-c.1004 (1080/72) in Villa. Mutant used for description and seed stocks: glo-c.1004 in Villa (GSHO 1329); glo-c.1004 in Bowman (PI 483237)*4(GSHO 1879).1514152GSHO 1329
244GSHO 1330nec3.dHordeum vulgare L. subsp. vulgare Bayern, Germany LEAFHistoric1994DEVELOPED49.00000000, 11.50000000Genetic materialTan or brown necrotic blotches develop on the leaf sheath and blade prior to heading and persist to maturity. Plants homozygous for nec3.d develop brown blotches having irregular margins. Plants homozygous for nec3.e develop large tan or faded blotches having distinct margins. After maturation, plants expressing the nec3.d gene can be identified easily by brown blotches on the leaves, but those expressing the nec3.e gene are difficult to identify. Inheritance: Monofactorial recessive. Located in chromosome 6HS, about 29.2 cM distal from the rob (orange lemma) locus, and about 16.7 cM distal from the msg36 (male sterile genetic 36) locus. Previous nomenclature and gene symbolization: Necrotic leaf spot 3=nec3a. Necrotic leaf spot 3=nec3b. Mutational events: nec3.d in Proctor (1339/62); nec3.e (1558/74) in Villa (PI 399506). Mutant used for description and seed stocks: nec3.d in Proctor (GSHO 1330); nec3.e in Villa (GSHO 2423); nec3.d in Bowman (PI 483237)*4 (GSHO 2065); nec3.e in Bowman*3 (GSHO 2066).1513878GSHO 1330
245GSHO 1379gpa1.bHordeum vulgare L. subsp. vulgare Idaho, United StatesNSGCKERNELS1994DEVELOPEDGenetic materialSeedlings display a pattern of transverse or alternating white and green bands on the first, second, and occasionally the third foliage leaves. Plants have a slightly pale green color prior to heading. Grandpa plants are sensitive to flooding and produce an albino flag leaf, peduncle, and spike. Plants are viable in the field, but the seed is thin and yields are low. Inheritance: Monofactorial recessive. Located in chromosome 2HL, about 24.5 cM distal from the lig (liguleless) locus. Previous nomenclature and gene symbolization: Grandpa=gp. Grandpa 2=gp2. Mutational events: gpa1.a in Lyallpur (GSHO 519), CIho 6854); gpa1.b (gp2) in Montcalm (CIho 7149). Mutant used for description and seed stocks: gpa1.b in Montcalm (GSHO 1379); gpa1.b in Bowman (PI 483237)*7 (GSHO 1934).1514266GSHO 1379
246GSHO 1390fch6.gHordeum vulgare L. subsp. vulgare Texas, United StatesNSGCKERNELS1994DEVELOPEDGenetic materialSeedlings appear yellowish-white initially, but green pigments develop near the leaf tip within one week. Green pigmentation is not as intense as in normal seedlings. Chlorophyll content is 15% of normal in 3-week old seedlings and free amino acid content is increased by 75% or more. Plants do mature in the field, but development is much delayed. Inheritance: Monofactorial recessive. Located in chromosome 5HL [7L], over 36.2 cM proximal from the srh (short rachilla hair) locus. Previous nomenclature and gene symbolization: Yellow virescent=yv. Chlorina seedling 6=f6. Mutational events: fch6.g in Himalaya (GBC380). Mutant used for description and seed stocks: frch6.g in Himalaya (GSHO 1390); fch6.g in GBC681 was used for linkage studies; fch6.g in Bowman (PI 483237)*7 (GSHO 2097).1514272GSHO 1390
247GSHO 1393blf1.aHordeum vulgare L. subsp. vulgare Ontario, CanadaNSGCKERNELS1994DEVELOPEDGenetic materialPlants are somewhat lighter green than normal. All leaf blades are very broad, about twice normal width, and markedly crinkled, especially at the margins. Inheritance: Monofactorial recessive. Located in chromsome 5HL [7L], about 20.6 cM proximal from the srh (short rachilla hair) locus. Previous nomenclature and gene symbolization: Bredbladig=bb. Latifolium. Broad leaf=bb. Mutational events: blf1.a in Bonus (Alb Acc 55). Mutant used for description and seed stocks: blf1.a in Bonus (GSHO 1393).1514485GSHO 1393
248GSHO 1394mul2.bHordeum vulgare L. subsp. vulgare Ontario, CanadaNSGCKERNELS1994DEVELOPEDGenetic materialSpikelets may have supernumerary florets and occasionally nodal branches appear at random on the spike. The character shows variable expression in some crosses, and is seen in only a few spikes of affected F2 plants. The trait isolated in the Bowman backcross-derived line is expressed in six-rowed progeny as a malformed rachilla in some lateral spikelets. The rachilla malformation is most pronounced in the middle of the spike and ranges from formation of a fertile floret to slight elongation of the rachilla. However, confirmation that this trait is the same gene as in the original stock has not been completed. Inheritance: Monofactorial recessive. Located in chromosome 6HL, about 24.2 cM distal from the rob (orange lemma) locus. Previous nomenclature and gene symbolization: Multiflorous 2=mlt2. The change to mul2 was recommended by Robertson. Mutatioinal events: mul2.b in Montcalm (Alb Acc 299, GBC326). Mutant used for description and seed stocks: mul2.b in Montcalm (GSHO 1394); mul2.b plus vrs1.a in Bowman (PI 483237)*6 (GSHO 2089).1514275GSHO 1394
249GSHO 1398mtt2.bHordeum vulgare L. subsp. vulgare Ontario, CanadaNSGCKERNELS1994DEVELOPEDGenetic materialYellow to ivory bands, rarely necrotic, appear at emergence on the seedling leaf. They gradually disappear and are absent at heading. Expression is better when alternating temperatures occur, but even under constant temperatures, the tip of the first leaf is yellow or pale green. Plant vigor and maturity seem unaffected by the presence of the mtt2.b gene. Inheritance: Monofactorial recessive. Located in chromosome 5HL [7L], about 26.1 cM proximal from the srh (short rachilla hair) locus. Previous nomenclature and gene symbolization: Mottled leaf 2=mt2. Mutational events: mtt2.b in Montcalm (Alb Acc 285). Mutant used for description and seed stocks: mtt2.b in Montcalm (GSHO 1398); mtt2.b in Bowman (PI 483237)*7 (GSHO 2099).1514416GSHO 1398
250GSHO 1404mss1.aHordeum vulgare L. subsp. vulgare Ontario, CanadaNSGCKERNELS1994DEVELOPEDGenetic materialBroad yellow to ivory vertical streaks develop in the leaves of some tillers prior to heading. Affected tillers are weaker and often abort under moderate to severe heat and moisture stress. In some environments, the whole plant becomes a golden yellow prior to maturation. Inheritance: Monofactorial recessive, the deviation from 3:1 reported by Walker et al. may be caused by genotype-environment interactions. Located in chromosome 2H, about 19.9 cM distal (probably) from the vrs1 (six-rowed spike 1) locus. Previous nomenclature and gene symbolization: None. Mutational events: mss1.a in Montcalm (Alb Acc 257). Mutant used for description and seed stocks: mss1.a in Montcalm (GSHO 1404); mss1.a in Bowman (PI 483237)*2 (GSHO 1918).1513715GSHO 1404
251GSHO 1451zeb3.cHordeum vulgare L. subsp. vulgare Utah, United StatesNSGCKERNELS1994DEVELOPEDGenetic materialEarly leaves of homozygous plants have bright yellow bands across a light yellow background when grown under alternating temperatures, warm day and cool night. Later leaves have patches of bright yellow. Cool temperatures enhance the yellow banding while warm temperatures increase green pigments. Under very cold conditions, leaves become yellowish white and deteriorate. Heterozygotes have slightly pale green color compared to normal plants. The zeb3.c gene must be maintained as a heterozygous stock. Mutational events: zeb3.c in an unknown cultivar (Utah 41, Alb Acc 43). Mutant used for description and seed stocks: zeb3.c in Utah 41 (GSHO 145`); zeb3.c in Bowman (PI 483237)*3 (GSHO 2057).1514228GSHO 1451
252GSHO 153vrs1.cHordeum vulgare L. subsp. vulgare Colorado, United StatesNSGCKERNELS1994DONATED05/1994Genetic materialThe spike has long awns on the lemma of the central spikelet and awn-like appendages on the lemma of the lateral spikelets. Lateral spikelets have perfect flowers and complete fertility. The lr trait is expressed only in six-rowed barley, and heterozygotes have a short awn on the lemma of lateral spikelets. Since recombination with neither vrs1.a(v) nor Vrs1.b (V) alleles was observed, the lr or vrs1.c phenotype is likely controlled by an allele at the vrs1 locus. The Bowman backcross- derived line for the vrs1.c gene has short, stiff awns on the central spikelets (about 1/3 normal length) and no awns on the lateral spikelets.Inheritance: An incomplete dominant in the multiple allelic series at the vrs1 (six-rowed spike 1) locus. Located in chromosome 2HL, about 15.4 cM distal from the Gth (toothed lemma) locus, and 26.5 cM distal from the eog (elongated outer glume) locus. Previous nomenclature and gene symbolization: Reduced lateral spikelet appendage on the lemma=lr. Reduced lateral spikelet appendage on the lemma=vlr. Mutational events: vrs1.c or lr in Hordeum intermedium var Nudihaxtoni. Mutant used for description and seed stocks: vrs1.c or lr in Hordeum intermedium var Nudihaxtoni (PI 32368, GSHO 153); vrs1.c in Bowman (PI 483237)*7(GSHO 1908).1510717GSHO 153
253GSHO 159wst1.aHordeum vulgare L. subsp. vulgare Colorado, United StatesNSGCKERNELS1994DONATED05/1994Genetic materialVery narrow white stripes are present in the leaves from the seedling stage until near maturity. Plants have nearly normal vigor. The size of the white sectors is variable from mutant to mutant, plant to plant, and leaf to leaf. All affected plants may produce albino seedlings among their selfed progeny with reported frequencies between 1.0 and 25.6%. Albino seedlings have not been observed when wst1 plants are used as the male parent in crosses. Caldecott's streak has wider and larger white sectors and produces a higher frequency of albino seedlings. Inheritance: Monofactorial recessive. Located in chromosome 3HL, very close to the uzu (uzu) locus. Previous nomenclature and gene symbolization: White stripe=wst. Stripe=st. White streak 3=wst3. Mutational events: wst1.a in an unknown cultivar (USDA 163, CIho 11767); wst1.c (OUL074, GSHO 569) in Akashinriki (PI 467400, OUJ659); wst1.e in Himalaya (GSHO 278, Caldecott's streak); wst1.f (Grandpa fine stripe) in an unknown cultivar (CIho6856, GSHO 854), wst1.g in CIho 6758 (GSHO 314); wst1.h in CIho 11766 (GSHO 797); wst1.i in Moravian (PI 537135). Mutant used for description and seed stocks: wst1.a in an unknown cultivar (GSHO 159); wst1.a in Bowman (PI 483237)*6 (GSHO 1966); wst1.c plus uzu1.b in Bowman*8 (GSHO 1967).1510718GSHO 159
254GSHO 16fch13.vHordeum vulgare L. subsp. vulgare Colorado, United StatesNSGCKERNELS1994DEVELOPEDGenetic materialThe mutant was classified as white seedlings (devoid of chlorophyll and carotenoid pigments) and maintained as a heterozygous stock in the USDA-ARS Barley Genetic Stocks Collection. Mutant plants die at the seedling stage if planted in the field. When grown in the greenhouse, however, seedlings gradually develop a pale green color, leaves have a light green color at heading, and plants survive to maturity and produce plump seed. Inheritance: Monofactorial recessive. Location is unknown. Previous nomenclature and gene symbolization: Chlrorina seedling 13=f13. Mutational events: fch13.v in Nigrinudum (Nigrinudum II). Mutant used for description and seed stocks: fch13.v in Nigrinudum (GSHO 16).1510774GSHO 16
255GSHO 165abo3.cHordeum vulgare L. subsp. vulgare Skåne län, SwedenNSGCKERNELS1994DEVELOPED58.98333300, 13.50000000Genetic materialSeedlings lack normal pigmentation; they are white and die at a two or three leaf stage. The abo3.c gene must be maintained as a heterozygous stock. Inheritance: Monofactorial recessive. Location is unknown. Previous nomenclature and gene symbolization: Albino 2=a2. Albino seedling 2=a2. Albina seedling-za=alb-za. Mutational events: abo3.c in an unknown cultivar; alb-za.111 in Bonus (PI 189763). Mutant used for description and seed stocks: abo3.c in an unknown cultivar (B14-9).1510626GSHO 165
256GSHO 1656ari-h.127Hordeum vulgare L. subsp. vulgare Skåne län, SwedenNSGCKERNELS1994DEVELOPED58.98333300, 13.50000000Genetic materialAwns are about 3/4 normal length, but plant height is normal. Inheritance: Monofactorial recessive. Located in chromosome 5HL [7L], very close to the raw1 (smooth awn 1) locus based on linkage drag. Previous nomenclature and gene symbolization: Breviaristatum-127=ari-127. Short awn 11=lk11. Mutational events: ari-h.127 in Foma (GSHO 1656); ari-h.127 in Bowman (PI 483237)*7 (GSHO 2125). Mutant used for description and seed stocks: ari-h.127 in Foma (GSHO 1656); ari-h.127 in Bowman (PI 483237)*7 (GSHO 2125).1004274GSHO 1656
257GSHO 167abo4.dHordeum vulgare L. subsp. vulgare Skåne län, SwedenNSGCKERNELS1994DEVELOPED58.98333300, 13.50000000Genetic materialSeedlings lack normal pigmentation; they are white and die at a two or three leaf stage. The abo4.d gene must be maintained as a heterozygous stock. Inheritance: Monofactorial recessive. Located in chromosome 2H, 5.0 to 10.2 cM from the fch1 (chlorina seedling 1) locus, and about 12.5 cM from the bo5 (albino seedling 5, a4) locus. Previous nomenclature and gene symbolization: Albino 3=a3. Albino seedling 3=a3. Mutational events abo4.d in an unknown cultivar. Mutant used for description and seed stocks: abo4.d in an unknown cultivar (GSHO 167).1510597GSHO 167
258GSHO 17Fol-b.2Hordeum vulgare L. subsp. vulgare Colorado, United StatesNSGCKERNELS1994DONATED05/1994Genetic materialHomozygous plants are very wead with thread-like, short leaves and often die at the 3 to 4 leaf stage. Heterozygotes have narrow leaves, small spikes and kernels, but vigor is good and fertility is normal. Inheritance: Monofactorial incomplete dominant. Location is unknown, but a possible association with chromosome 1H [5] based on trisomic analysis. Previous nomenclature and gene symbolization: Angustifolia=Ang. Mutational events: Fol-b.2 in an unknown cultivar. Mutant used for description and seed stocks: Fol-b.2 in an unknown cultivar (GSHO 17).1510692GSHO 17
259GSHO 171lgn3.cHordeum vulgare L. subsp. vulgare Minnesota, United StatesNSGCKERNELS1994DEVELOPEDGenetic materialSeedlings have light or pale green color. The pale green color persists until near maturity, and the plants are very weak or non-viable in the field. Plants can be grown to maturity in greenhouses, but tillering is much reduced and seeds are very thin. Inheritance: Monofactorial recessive. Located in chromosome 4HL, over 10.5 cM distal from the Kap (hooded lemma) locus. Previous nomenclature and gene symbolization: Light green 7=lg7. Light green 3=lg3, the lg3 stock studied by Hanson and Kramer may not be the same as the lg3 (GSHO 171) stock. Mutational events: lgn3.c in an unknown cultivar (No 150); lgn3.g (lg7) in an unknown cultivar (GSHO 173, No 154). Mutant used for description and seed stocks: lgn3.c in an unknown cultivar (GSHO 171); lgn3.c in Bowman (PI 483237)*5 (GSHO 2010).1510675GSHO 171
260GSHO 185blx1.aHordeum vulgare L. subsp. vulgare Colorado, United StatesNSGCKERNELS1994DONATED05/1994Genetic materialBlue aleurone color is due to anthocyanin pigments which occur as lumps inside many aleurone granules in some or all aleurone cells. Variaiton in color expression from dark blue to an off-white is caused by environmental factors and modifying genes. Aleurone color is best observed in well filled grain that is magnified to show individual aleurone cells, after more superficial tissues have been peeled off. In heritance: Monofactorial recessive when complementary dominant alleles are present at the Blx2, Blx3, Blx4, and Blx5 loci. Located in chromosome 4HL, over 13.6 cM distal from the glf3 (glossy leaf 3) locus. Previous nomenclature and gene symbolization: Blue aleurone=Bl. Complementary factors for blue vs white aleurone=Bl1 and bl1. Complementary factors for blue vs white aleurone=Blx and blx. Mutational events: blx1.a in Goldfoil (CIho 1866). Mutant used for description and seed stocks: blx1.a in Goldfoil (GSHO 185).1510621GSHO 185
261GSHO 187Rpg2.bHordeum vulgare L. subsp. vulgare Wisconsin, United StatesNSGCKERNELS1994DEVELOPEDGenetic materialModerate resistance to some races of Puccinia graminis f sp tritici is conferred by the Rpg2.b gene. Percent stem and leaf sheath infection and lesion size in adult plants are reduced compared with susceptible plants under field conditions. Patterson used linkage data to demonstrate that the Rpg2 locus differs from the Rpg1 locus. Inheritance: Monofactorial dominant. Location is unknown. Previous nomenclature and gene symbolization: Resistance to Puccinia graminis tritici=T2. Resistance to Puccinia graminis Pers f sp tiritici Eriks and E Henn = Rpg2b. Mutational events: Prg2.b in Hietpas. Mutant used for description and seed stocks: Rpg2.b in Hietpas 5 (GSHO 187).1510786GSHO 187
262GSHO 196vrs1.aHordeum vulgare L. subsp. vulgare Colorado, United StatesNSGCKERNELS1994DONATED05/1994Genetic materialAlleles at this complex locus modify development of the lateral spikelets and the associated lemma awn. The vrs1.a allele (v gene) is present in most six-rowed cultivars and produces well-developed lateral spikelets. The lemma awn of lateral spikelets will vary from 3/4 to nearly as long as those of central spikelets, depending upon alleles present at other loci. The Vrs1.b allele (V gene, distichon) is present in many two-rowed cultivars and reduces lateral spikelets to sterile bracts with a rounded tip. The Vrs1.t allele (Vt gene, deficiens) causes an extreme reduction in the size of lateral spikelets. The lr or vlr (vrs1.c) gene in Nudihaxtoni and Bozu types will not recombine with the vrs1.a allele and produces phenotypes similar to the Vrs1.d allele (Vd gene) of Svanhals. The series of induced mutants in two-rowed barley called hex-v and Int-d mutants differ in the size of lateral spikelets, but they interact with the vrs1.a allele as incomplete dominants. Many heterozygous combinations with vrs1.a have a pointed tip on the lemma of sterile lateral spikelets. Alleles at the int-c (intermedium spike-c) locus modify lateral size in the presence of vrs1.a, Vrs1.b, and Vrs1.d, but not when Vrs1.t is present. Inheritance: A multiple allelic series, incomplete dominant allele interactions based on the size and shape of lateral spikelets. Located in chromosome 2HL, about 30.5 cM distal from the eog (elongated outer glume) locus. Previous nomenclature and gene symbolization:Two-row vs six-row=Zz. Six-row vs two-row=Aa. Two-row=D. Six-row vs two-row=Vv. Six-row vs two-row (distichon) vs two-row (deficiens)=A,as,af. Redcd lateral spikelet append. on the lemma=lr. Allelic series v,Vd,V,Vt. Hex. mut. Intermed. spike-d=Int-d. Reduced lat. spike. append. on lemma-vlr. Mut.events:vrs1.a ... vrs1.o(v1b) in New Golden.Mut. used for descr.:vrs1.a...(GSHO 1907).1510659GSHO 196
263GSHO 2xnt2.bHordeum vulgare L. subsp. vulgare Colorado, United StatesNSGCKERNELS1994DEVELOPEDGenetic materialSeedlings are yellow in color and do not develop green pigmentation. Since they do not survive beyond the two leaf stage, the xnt2.b gene must be maintained as a heterozygous stock. Inheritance: Monofactorial recessive. Location is unknown. Previous nomenclature and gene symbolization: Xantha seedling b=xb. Yellow seedling b=xb. Mutational events: xnt2.b (xb) in Black Hulless (Black Hulless II). Mutant used for description and seed stocks: xnt2.b in Black Hulless (GSHO 2).1510590GSHO 2
264GSHO 20crm1.aHordeum vulgare L. subsp. vulgare Colorado, United StatesNSGCKERNELS1994DEVELOPEDGenetic materialSeedlings are cream in color and survive until they are about two weeks old. The crm1.a gene must be maintained as a heterozygous stock. Inheritance: Monofactorial recessive. Located in chromosome 5HL [7L], closely linked to the raw1 (smooth awn 1) locus, and about 26.0 cM from the srh (short rachilla hair) locus. Previous nomenclature and gene symbolization: White seedling b=ab. Albino seedling b=ab. Cream seedling=cm. Mutational events: crm1.a in Black Hulless (Black Hulless III). Mutant used for description and seed stocks: crm1.a in Black Hulless (GSHO 20).1510640GSHO 20
265GSHO 209blx2.bHordeum vulgare L. subsp. vulgare Colorado, United StatesNSGCKERNELS1994DONATED05/1994Genetic materialBlue aleurone color is due to anthocyanin pigments, which occur as lumps inside many aleurone granules in some or all aleurone cells. Variation in color expression from dark to blue to an off-white is caused by environmental factors and modifying genes. Aleurone color is best observed in well filled grain that is magnified to show individual aleurone cells, after more superficial tissues have been peeled off. Inheritance: Monofactorial recessive when complementary dominant alleles are present at the Blx1, Blx3, Blx4, and Blx5 loci. Located in chromosome 7HL [1L], approximately 9.9 cM distal from the nud (naked caryopsis) locus. Previous nomenclature and gene symbolization: Blue aleurone 1=Bl1 a second factor for blure aleurone. Non-blue aleurone 2=bl2. Mutational events: blx2.b in Nepal (CIho 595). Mutant used for description and seed stocks: blx2.b in Nepal (GSHO 209); also blx2.b in Bowman (PI 483237).1510682GSHO 209
266GSHO 210trp1.aHordeum vulgare L. subsp. vulgare Colorado, United StatesNSGCKERNELS1994DONATED05/1994Genetic materialMutants may have three awns on the lemma instead of one, but expression is variable. The awn or hood on the lemma of the central spikelet forks to produce one normal central appendage and one or two shorter lateral ones. Expression in the Bowman backcross-derived line is reduced to an occasional branch on the awn. Inheritance: Monofactorial recessive. Located in chromosome 2HL, over 35.5 cM distal from the wst7 (white streak 7) locus. Previous nomenclature and gene symbolization: Triple- bearded variation. Triple awned lemma=tr. Mutational events: trp1.a in Triple-awn lemma (CIho 6630). Mutant used for description and seed stocks: trp1. in CIho 6630 (GSHO 210); trp1.a from CIho 6880 (Ahang/ Twisted Flag//Triple-awn Lemma) in Bowman (PI 483237)*7 (GSHO 1936).1510639GSHO 210
267GSHO 216rtt1.aHordeum vulgare L. subsp. vulgare Colorado, United StatesNSGCKERNELS1994DEVELOPEDGenetic materialSpikes are highly modified with numerous immature spikelets and complete sterility. The arrangement of bracts in the spikelets suggests that they contain numerous florets. Because the homozygous recessive is completely sterile, the stock must be maintained in the heterozygous condition. A semidominant interaction between rtt1.a and the Rtt1.b (normal) allele in Okaiku 3 was reported by Takahashi et al. Inheritance: Monofactorial recessive. Located in chromosome 2HS, about 6.0 cM distal from the fch15 (chlorina seedling 15) locus. Previous nomenclature and gene symbolization: Rattail spike=rt. Mutational events: rtt1.a in Goldfoil, Rtt1.b in Okaiku 3 (OUL011). Mutant used for description and seed stocks: rtt1.a in Goldfoil (GSHO 216).1510821GSHO 216
268GSHO 227trd1.bHordeum vulgare L. subsp. vulgare Colorado, United StatesNSGCKERNELS1994DONATED05/1994Genetic materialThe characteristic trait of this mutant is the presence of a bract, third outer glume, outside the two empty glumes of the central spikelet. The bract associated with the lowest spikelet is always the largest, embracing in some cases about one-half the spike, and the bracts become progressively smaller toward the tip of the spike. Basal rachis internodes may be elongated. Allelism tests were not conducted for the first three trd mutants reported, but the linkage value between the trd and Blp loci was similar in all three studies. Takahashi et al. assumed that they used the same material studied by Miyake and Imai. Inheritance: Monofactorial recessive. Located in chromosome 1HL [5L], about 16.0 cM distal from the Blp (black lemma and pericarp) locus, and about 11.c cM proximal from the eam8 (early maturity 8) locus. Previous nomenclature and gene symbolization: Third glume=t. Bracteate. Bracteatum-c=bra-c. Mutational events: trd1.a in Hordeum vulgare var afghanicum; trd1.b in Valki (CIho 5478); trd1.c in a Japanese cultivar; trd1.d (Mut 1969/61) in Proctor (PI 280420); bra-c.1 in Bonus (PI 189763); bra-c.2, -c.3, -c.4, -c.5 in Bonus, -c.6 in Foma (CIho 11333), -c.8 in Kristina. Mutant used for description and seed stocks: trd1.b in Valki (GSHO 227); bra-c.1 in Bonus (GSHO 1695); trd1.b from Wolfe's Chromosome 5 Marker Stock in Bowman (PI 483237)*7 (GSHO 2060); bra-c.1 in Bowman*4 (GSHO 2061).1510754GSHO 227
269GSHO 233abo11.kHordeum vulgare L. subsp. vulgare Colorado, United StatesNSGCKERNELS1994DONATED05/1994Genetic materialSeedlings are albino when grown under low temperature conditions (below 7.5 degrees C), but normal green under high temperature conditions (above 18.5 degrees C). Failure to produce chlorophyll seems confined to early seedling stages. Inheritance: Monofactorial recessive. Location is unknown. Previous nomenclature and gene symbolization: Albino seedling t=albt. Albino seedling t3=at3. Mutational events: abo11.k in Trebi (Trebi II). Mutant used for description and seed stocks: abo11.k in Trebi (GSHO 233).1510828GSHO 233
270GSHO 234Pre2.aHordeum vulgare L. subsp. vulgare Colorado, United StatesNSGCKERNELS1994DONATED05/1994Genetic materialPurple or red coloration of the hull (chaff) and pericarp develops during the soft dough stage of grain fill, and fades as the grain matures. Anthocyanin pigments in an acid cell solution produce the red to violet color observed in the pericarp and lemma. Inheritance: Two dominant complementary genes (Pre1 and Pre2) control purple or red pigmentation of the lemma, palea, and pericarp. Located in chromosome 2HL, about 17.1 cM distal from the vrs1 (six-rowed spike 1) locus, and over 11.2 cM proximal fro the ant2 (anthocyanin-less2) locus. Previous nomenclature and gene symbolization: Purple chaff color=P. Purple chaff color=P1. Purple pericarp=P. Purple lemma =P. Red pericarp=R. Red pericarp 2=Re2. Mutational events: Pre2.a in Buckley (NSL 32606). Mutant used for description and seed stock: Pre2.a in Buckley 3277 (GSHO 234); Pre2.b from R.I. Wolfe's Multiple Dominant Marker Stock in Bowman (PI 483237)*9 (GSHO 1926).1510594GSHO 234
271GSHO 247wst7.kHordeum vulgare L. subsp. vulgare Colorado, United StatesNSGCKERNELS1994DEVELOPEDGenetic materialVertical white streaks of variable width and number develop in the leaf blades of young secondary tillers. Fewer white streaks and fewer tillers with white streaks occur as environmental conditions become warm. White streaks can be found until near maturity, but they are difficult to observe after heading under field conditions. Often the lower or first leaves on early tillers have more and wider streaks. Inheritance: Monofactorial recessive. Located in chromosome 2HL, about 2HL, about 22.0 cM distal from the gpa (grandpa) locus, and over 29.4 cM distal from the lig (liguleless) locus. Previous nomenclature and gene symbolization: Ribbon grass=rb. White streak-k=wst,,k. White streak-B=wst,,B. Mutational events: wst7.k in an unknown cultivar. Mutant used for description and seed stocks: wst7.k in an unknown cultivar (GSHO 247); wst7.k from Wolfe's Multiple Recessive Marker Stock 1 in Bowman (PI 483237)*7 (GSHO 1935).1510773GSHO 247
272GSHO 25brh1.aHordeum vulgare L. subsp. vulgare Missouri, United StatesNSGCKERNELS1994DEVELOPEDGenetic materialPlants have short leaves, culms, spikes, awns, and seeds. The seedling leaf is about 2/3 normal length, and a similar reduction in the size of other organs is observed, but the awns are less than 1/3 normal length. Seeds are small and yields are low. The mutant phenotype is easy to classify at all stages of growth. The brachytic or semidwarf mutant was selected by L.J. Stadler and the gene symbol br was given by Powers. Inheritance: Monofactorial recessive. Located in chromosome 7HS[1S] about 9.3 cM distal from the fch12 (chlorina seedling 12) locus. Previous nomenclature and gene symbolization: Brachytic=br. Breviaristatum-i=ari-i. Dwarf x=dx1. Mutational events:brh1.a in Himalaya;brh1.cGSHO229 in Moravian(PI 539135);ari-i.38 GSHO 1657 in Bonus (PI 189763); brh1.e (GSHO 1690) in Aramir (PI 467786); brh1.f (GSHO 1422) in Domen (CIho 9562); brh1.t(OUM136, GSHO 1691) in Akashinriki (PI 467400, OUJ659); brh1.x (GSHO 1692) in Volla (PI 280423), brh1.ae (FN53) in Steptoe (CIho 15229). Mutant used for description and seed stocks: brh1.a in Himalay(GSHO 25); brh1.a in Bowman (PI 483237)*7 (GSHO 1820); ari-i.38 in Bowman*6 (GSHO 1821); brh1.e in Bowman*7 (GSHO 1822); brh1.t in Bowman*7 (GSHO 1823); brh1.x in Bowman*7 (GSHO 1 824.1510695GSHO 25
273GSHO 2502yst4.dHordeum vulgare L. subsp. vulgare Colorado, United StatesNSGCKERNELS1994DONATED05/1994Genetic materialPlants have a yellow-green color with numerous, vertical yellow streaks in the leaves. The yellow-green color is retained until maturity, but the yellow streaks may be difficult to observe after heading. Plant vigor and height are reduced, heading is delayed, and seed yields are low. Inheritance: Monofactorial recessive. Located in chromosome 2HL, near the vrs1 (six-rowed spike 1) locus. Previous nomenclature and gene symbolization: None. Mutational events: yst4.d in Glenn (DWS 1059). Mutant used for description and seed stocks: yst4.d in Glenn (GSHO 2502); yst4.d in Bowman (PI 483237)*7 (GSHO 1922).1545057GSHO 2502
274GSHO 253mnd1.aHordeum vulgare L. subsp. vulgare District of Columbia, United StatesNSGCKERNELS1994DEVELOPEDGenetic materialWhen grown in the field, plants are about 1/2 normal height with numerous tillers. Each tiller has 12 to 20 small leaves and a spike of 1/2 normal size. Additional tillers with 4 to 7 leaves may develop at the uppermost tiller nodes, but these secondary and tertiary culm branches develop late and have very small spikes. When grown in the greenhouse, plants are taller and later than normal sibs and each tiller may have 15 to 20 nodes. The rate of primordium initiation is normal, but their expansion into leaves is much more rapid. Inheritance: Monofactorial recessive. Location is unknown. Previous nomenclature and gene symbolization: Many noded dwarf=m. Mutational events: mnd1.a in Mesa. Mutant used for description and seed stocks: mnd1.a in Mesa (GSHO 253); mnd1.a in Bowman (PI 483237)*9 (GSHO 2038).1512491GSHO 253
275GSHO 266en-minHordeum vulgare L. subsp. vulgare Colorado, United StatesNSGCKERNELS1994DONATED05/1994Genetic materialPlants are extremely dwarfed with the adult plant being less than 5 cm tall. The leaf blade and sheath are very short and thick and have a whitish dark green color. Roots are thick and short with C-tumor-like swelling at their tips. No spikes are formed; hence, the stock must be maintained as a heterozygous plants, those that will give rise to minute plants have a markedly shrunken endosperm. Tissues of the plant are mixoploid with 2x, 3x, 4x, and up to 60x cells being formed by abnormal cytokinesis. Inheritance: Monofactorial recessive, but expressed only in plants homozygous for the min1.a (semi-minute dwarf 1) allele. Location is unknown. Previous nomenclature and gene symbolization: None. Mutational events: Natural occurrence of en-min in Kaiyo Bozu (OUL921). Mutant used for description and seed stocks: en-min in a line of Kaiyo Bozu that is heterozygous at the min1 locus (GSHO 266).1512211GSHO 266
276GSHO 267sbk1.aHordeum vulgare L. subsp. vulgare Colorado, United StatesNSGCKERNELS1994DONATED05/1994Genetic materialThe spike of this mutant is similar to that of Kap (hooded lemma), but the central cup-like cavity of the lemma appendage or its modification is situated somewhat below the top of the lemma, and no lema wings exist with the hood. There are no rudiments of a floral organ within the hood-like cavity. In the homozygous condition, the sbk1.a gene is epistatic to Kap1.a and to Lks1.a (awnless) genes. The Bowman backcross-derived line with the sbk1.a gene is earlier, shorter, and less vigorous than the recurrent parent. Inheritance: Monofactorial recessive. Located in chromosome 2HS, over 30.4 cM distal from the eog (elongated outer glume) locus and probably a little distal from the com2 (compositum 2) locus. Previous nomenclature and gene symbolization: Subjacent hood=sk. Hooded awn 2=k2. Hoods=kr. Calcaroides-d=cal-d. Mutational events; sbk1.a in Tayeh 13; cal-d.4 ( GSHO 1697) in Foma (CIho 11333), -d.14 in Kristina, -d.18, -d.22 in Bonus (PI 189763); sbk1.b (kr) in Montcalm (CIho 7149). Mutant used for description and seed stocks: sbk1.a in Tayeh 13 (GSHO 267); sbk1.a in Bowman (PI 483237)*7 (GSHO 1874); cal-d.4 in Bowman*3 (GSHO 1875).1512415GSHO 267
277GSHO 27smooth awn 1Hordeum vulgare L. subsp. vulgare Colorado, United StatesNSGCKERNELS1994DONATED05/1994Genetic materialBarbs or teeth on the margins of the awn are absent and the size and number of barbs on the central vein are reduced. Plants homozygous for the raw1.a allele have a semi-smooth awn when other recessive genes reducing the number and/or size of the barbs are absent. The presence of these other genes may cause a complete absence of barbs on the awn. The number of stigma hairs is decreased greatly in plants homozygous for raw1.a, and reduced seed set may occur in some genotype-environment combinations. Inheritance: Monofactorial recessive. Located in chromosome 5HL [7L], about 26.8 cM distal from the srh (short rachilla hair) locus. Previous nomenclature and gene symbolization: Smooth awn=r. Smooth awn=a. Mutational events: raw1.a in Lion (CIho 923). Mutant used for description and seed stocks: raw1.a in Lion (GSHO 27); raw1.a in Bowman (PI 483237), probably from Lion via Midwestern six-rowed cultivars; Raw1 from CIho 2376 in Bowman*8 (GSHO 2124).1510653GSHO 27
278GSHO 270alm1.aHordeum vulgare L. subsp. vulgare Colorado, United StatesNSGCKERNELS1994DONATED05/1994Genetic materialThe lemma and palea are white in color and mostly devoid of chlorophyll, but they terminate into green tips with green awns. The basal part of lower leaf sheaths and stem nodes are devoid of chlorophyll. Ligules and joints between the leaf sheath and blade are white in color. Plant vigor is reduced slightly and maturity is delayed in the Bowman backcross-derived line. Inheritance: Monofactorial recessive. Located in chromosome 3HS, about 16.5 cM distal from the uzu (uzu) locus. Previous nomenclature and gene symbolization: Albino lemma=al. Eburatum=ebu-a. Mutational events: alm1.a in Russia 82; alm1.b in Liberty (CIho 9549); alm1.c (Mut 966/61) in Proctor (PI 280420); ebu-a.2, -a.3 in Foma (CIho 11333); ebu-a.4, -a.5 in Foma. Mutant used for description and seed stocks: alm1.a in Russia 82 (GSHO 270); alm1.a in Bowman (PI 483237)*8 (GSHO 1953).1512375GSHO 270
279GSHO 274cur2.bHordeum vulgare L. subsp. vulgare Colorado, United StatesNSGCKERNELS1994DONATED05/1994Genetic materialThe awn, lemma, and palea are extremely curly, the rachis is twisted slightly in most spikes, and the stem internodes are strongly curved. Leaves are short and severely twisted. Roots are extremely curled compared to the straight roots of normal sister plants. Inheritance: Monofactorial recessive. Located in chromosome 3HL, over 36.8 cM distal from the als (absent lower laterals) locus and about 7.7 cM proximal from the fch2 (chlorina seedling 2) locus. Previous nomenclature and gene symbolization: Deformed plant, Bikini. Curly 2=cu2. Mutational events: cur2.b in Choshiro (OUL006); cur2.c (dwarf 2) in an unknown cultivar; cur2.d (deformed plant, Bikini) in Bonus (PI 189763); cur2.g (DWS1182, GSHO 1706) in Carina (PI 371632). Mutant used for description and seed stocks: cur2.b in Choshiro-hen (GSHO 274); cur2.b in Bowman (PI 483237)*5 (GSHO 1991); cur2.g in Bowman*5 (GSHO 1992).1512099GSHO 274
280GSHO 29eog1.aHordeum vulgare L. subsp. vulgare California, United StatesNSGCKERNELS1994DEVELOPEDGenetic materialA multiple allelic series exists at this locus which controls increased size of the outer glumes (glumes) of the spikelet. Glume width varies from 2.5 to 4.0 mm depending on genetic background and specific allele. Glume awn length varies from awnless to nearly as long as the lemma awn (92 mm) depending on the eog allele present and genes at other loci, but some confusion has occurred regarding alleles at the eog locus and expression of other glume characteristics. The size of the glume may range from less than twice normal width in CIho 14955 to lemma-like glumes in Triple Bearded Mariout. Inheritance: Monofactorial recessive. Located in chromosome 2HL, about 30.5 cM proximal from the vrs1 (six-rowed spike 1) locus. Previous nomenclature and gene symboliation: Extended outer glumes=e. Wide glumes=w. Macrolepis=lep-e(lep-a). Long outer glume=log. Elongated outer glume=e. Mutational events: eog1.a in Triple Bearded Mariout (CIho 2523); lep-e.1, -e.2 in Gull (CIho 1145), -e.3, -e.4, -e.5, -e.6, -e.7, -e.8, -e.9, -e.10, -e.11, -e.12, -e.13, -e.29, -e.34, -e.42, -e.45, -e.49, -e.50, -e.51, -e.52, -e.53, -e.54 in Bonus (PI 189763), -e.14, -e.15, -e.16, -e.17, -e.18, -e.19, -e.20, -e.21, -e.22, -e.23, -e.24, -e.25, -e.26, -e.27, -e.28, -e.30, -e.31, -e.32, -e.33, -e.38 in Foma (CIho 11333), -e.35, -e.36, -e.37, -e.39, -e.40, -e.41, -e.42, -e.43, -e.44, -e.46, -e.47, -e.48 in Kristina; eog 1.b in PI 173567 (G 8); eog1.c in CIho 14955; eog1.d (Mut 3447/64) in Proctor (PI 280420). Mutant used for description and seed stocks: eog1.a in Triple Bearded mariout (GSHO 29); lep-e.1 in Gull (GSHO 960); eog1.a in Bowman (PI 483237)*7(GSHO 1891); eog1.c from CIho 14955 in Bowman*7 (GSHO 1892).1510696GSHO 29
281GSHO 30abo6.fHordeum vulgare L. subsp. vulgare Colorado, United StatesNSGCKERNELS1994DEVELOPEDGenetic materialSeedlings are white in color, devoid of normal pigments, and die about 10 days after emergence. Later, plants with this phenotype were classified as albino mutants. The abo6.f gene must be maintained as a heterozygous stock. Inheritance: Monofactorial recessive. Located in chromosome 3HS, about 10.7 cM distal from the uzu (uzu) locus, and about 2.4 cM from the msg5 (male sterile genetic 5) locus. Previous nomenclature and gene symbolization: White seedling c=ac. Albino seedling c=ac. Mutational events: abo6.f in Colsess [Colsess I, Colsess II, Colsess III (GSHO 63)]. Mutant used for description and seed stocks: abo6.f in Colsess (GSHO 30).1510818GSHO 30
282GSHO 309Gth1.aHordeum vulgare L. subsp. vulgare Colorado, United StatesNSGCKERNELS1994DONATED05/1994Genetic materialLarge teeth or barbs are present on the upper part of lateral lemma nerves. Ubisch distinguished three sizes of teeth, including one that could be seen only with magnification. She reported that one gene pair controlled the presence and absence of large teeth and one or two other pairs of genes were responsible for less developed teeth. Wexelsen observed segregation for one pair of factors. Inheritance: Monofactorial dominant. Located in chromosome 2HL, about 15.4 cM proximal from the vrs1 (six-rowed spike 1) locus. Previous nomenclature and gene symbolization: Toothed lemma=G. Mutational events: Ght1.a in Machine (CIho 4982). Mutant used for description and seed stocks: Gth1.a in Machine (GSHO 309); Gth1.a in Bowman (PI 483237); gth1.b from ND11894 in Bowman*3 (GSHO 1897).1512164GSHO 309
283GSHO 346Vrs1.dHordeum vulgare L. subsp. vulgare Colorado, United StatesNSGCKERNELS1994DONATED05/1994Genetic materialPlants homozygous for Vrs1.d have more lateral kernels than those with Vrs1.b (V), which is the common allele in two-rowed barley. Homozygosity for the Vrs1.d allele and the int-c.b (formally i) allele at the int-c (intermedium-c) locus prevents the development of ovules and awns in lateral spikelets. The Vrs1.d allele interacts with the Int-c.a (formally l) allele to cause development of a few fertile lateral spikelets. Presence of the Int-c.h (formally lh) allele contributes to a greater number of lateral kernels than Int-c.a. Inheritance: A partially dominant allele in the multiple allelic series at the vrs1 (six-rowed spike 1) locus. Located in chromosome 2HL, about 15.4 cM proximal from the Gth (toothed lemma locus, and 30.5 cM f from the eog (elongated outer glume) locus. Previous nomenclature and gene symbolization: Two-rowed barley of Hordeum distichon=Vd. Renamed Vrs1.d based on allelism tests. Mutational events: Vrs1.d or Vd in Svanhals (PI 5474). Mutant used for description and seed stock: Vrs1.d or Vd in Svanhals (GSHO 346).1512104GSHO 346
284GSHO 348abo9.iHordeum vulgare L. subsp. vulgare Colorado, United StatesNSGCKERNELS1994DEVELOPEDGenetic materialSeedlings are white in color and devoid of normal pigments. Later, plants with this phenotype were classified as albino mutants. The abo9.i gene must be maintained as a heterozygous stock. Inheritance: Monofactorial recessive. Located in chromosome 3HS, about 9.4 cM distal from the xnt3 (xantha seedling 3) locus, about 15.5 cM proximal from the xnt6 (xantha seedling 6) locus, and about 9.9 cM distal from the yst1 (yellow streak 1) locus. Previous nomenclature and gene symbolization: White seedling n=an. Albino seedling n=an. Mutational events: abo9.i in Nigrinudum (Nigrinudum I). Mutant used for description and seed stocks: abo9.i in Nigrinudum (GSHO 348).1512105GSHO 348
285GSHO 35ddt1.aHordeum vulgare L. subsp. vulgare Idaho, United StatesNSGCKERNELS1994DEVELOPEDGenetic materialThe recessive allele, ddt1.a confers resistance to the insecticide DDT. Susceptible plants show severe chlorosis followed by yellowing and death about 10 days after spraying seedlings at the 2- to 3- leaf stage with a concentration of 1 part per 500 in water. A second application about 7 days after the first will insure complete kill of susceptible seedlings. The DDT solution affects only the leaf it comes in contact with it; thus, plants can be classified for reaction to DDT about anthesis by dipping a single leaf into the DDT solution. The part of the leaf blade covered by solution on susceptible plants will turn yellow and die, whereas it will remain green on resistant plants. The frequency DDT susceptibility in collections is about 10%, nearly 40% in two-rowed entries and 1% in six-rowed ones. Inheritance: Monofactorial recessive. Located in chromosome 5HS [7S], about 3.0 cM distal from the T2-7a break point. Previous nomenclature and gene symbolization: Resistance to DDT=ddt. Mutational events: ddt1.a in Spartan (CIho 5027). Mutant used for description and seed stocks: ddt1.a in Spartan (GSHO 35).1510832GSHO 35
286GSHO 36fch12.bHordeum vulgare L. subsp. vulgare Colorado, United StatesNSGCKERNELS1994DEVELOPEDGenetic materialSeedling leaves are yellow with green tips and new leaves show a yellow base and a green tip. As the plant develops, leaf color changes to pale green. Plants are vigorous, but anthesis is delayed and seed yielld is low. Inheritance: Monofactorial recessive. Located in chromosome 7HS [1S], about 3.6 cM distal from the gsh3 (glossy sheath 3) locus, and about 9.3 cM proximal from the brh1 (brachytic 1) locus. Previous nomenclature and gene symbolization: Chlorina seedling-c=fc. Chlorina seedling-fc=clo-fc. Mutational events: fch12.b(fc) in Colsess (Colsess V); fch12.l (Trebi chlorina 453, GSHO 155), fch12.m (Trebi V, GSHO 158), fch12.n (Trebi IX, GSHO 18), fch12.o (Trebi XI, GSHO 163) in Trebi (PI 537442); clo-fc.110 in Bonus (PI 189763); fch 12.b may be present in the brachytic chlorina stocks (GSHO 124 and GSHO 174). Mutant used for description and seed stocks: fch12.b in Colsess (GSHO 36); fch12.b in Bowman (PI 483237) *7 (GSHO 1826).1510689GSHO 36
287GSHO 4fch7.hHordeum vulgare L. subsp. vulgare Colorado, United StatesNSGCKERNELS1994DEVELOPEDGenetic materialSeedlings are light green and remain light green until near maturity. Plant vigor and seed yield are reduced, and developmental stages are delayed. Inheritance: Monofactorial recessive. Located in chromosome 1HL, about 4.7 cM proximal from the trd (third outer glume) locus, and about 22.7 cM distal from the Blp (black lemma and pericarp) locus. Previous nomenclature and gene symbolization: Chlorina seedling 7=f7. Chlorina seedling 7=clo-f7f7. Mutational events: fch7.h in Smyrna (Smyrna II; fch7.h in Smyrna (Smyrna III). Mutant used for description and seed stocks: fch7.h in Smyrna (GSHO 4); fch7.h in Bowman (PI 483237)*7 (GSHO 2058).1510817GSHO 4
288GSHO 40fch8.jHordeum vulgare L. subsp. vulgare Colorado, United StatesNSGCKERNELS1994DEVELOPEDGenetic materialSeedlings have a yellow-green color and plants retain a chlorina (yellow-green) phenotype until ripening. Plants are viable in the field, but heading is delayed and seed yield is much reduced. Inheritance: Monofactorial recessive. Located in chromosome 7HS[1S]; about 13.0 cM distal from the nud (naked caryopsis) locus, and about 2.7 cM distal from the msg10 (male sterile genetic 10) locus. Previous nomenclature and gene symbolization: Chlorina seedling 8-f8. Chlorina seedling-f8=clo-f8 Mutational events: fch8.j in Comfort (Comfort No II). Mutant used for description and seed stock: fch8.j in Comfort (GSHO 40); fch8.j in Bowman (PI 483237)*7(GSHO 1829).1510697GSHO 40
289GSHO 41yvs2.cHordeum vulgare L. subsp. vulgare Colorado, United StatesNSGCKERNELS1994DEVELOPEDGenetic materialThe seedling leaf is white with a very marked green tip, which remains green. The green area on the second leaf is very small or lacking. Homozygous recessive seedlings seldom survive beyond the two leaf stage. The ycs2.c gene is generally maintained as a heterozygous stock. In heritance : Monofactorial recessive. Located in chromosome 7HS[1S], about 24.4 cM proximal from the gsh3 (glossy sheath 3) locus, and 17.0 cM distal from the cer-f (eceriferum-f) locus. Previous nomenclature and gene symbolization: Virescent seedling-c=yc. Mutational events: yvs2.c in Coast (Coast III). Mutant used for description and seed stocks: yvs2.c in Coast (GSHO 41).1510721GSHO 41
290GSHO 42xnt4.dHordeum vulgare L. subsp. vulgare Colorado, United StatesNSGCKERNELS1994DEVELOPEDGenetic materialSeedlings have a yellow color and die at a two to three leaf stage. The xnt4.d gene must be maintained as a heterozygous stock. Inheritance: Monofactorial recessive. Located in chromosome 7HL [1L], over 22.0 cM from the centromere using T1-6a as a tester stock. Previous nomenclature and gene symbolization: Yellow seedling. Xantha seedling c2-xc2. Mutational events: xnt4.d(xc2) in Coast (Coast I). Mutant used for description and seed stocks: xnt4.d in Coast (GSHO 42).1510678GSHO 42
291GSHO 425cer-d.5Hordeum vulgare L. subsp. vulgare Skåne län, SwedenNSGCKERNELS1994DEVELOPED58.98333300, 13.50000000Genetic materialSurface wax coating on the spike appears to be reduced (wax code + ++ ++). A translocation is present in the mutual stock cer-d.5, but it is not linked to the cer-d locus. The original stock has a lax spike and heads early. The Bowman backcross-derived line has narrow leaves and reduced vigor, and heads a few days earlier than Bowman. Inheritance: Monofactorial recessive. Location is unknown. Previous nomenclature and gene symbolization: None. Mutational events: cer-d.5 (trans), -d.40, -d.104 in Bonus; cer-d.111, -d.149 in Bonus; cer-d.713 in Bonus; cer-d.809, -d.826 in Bonus; cer-d.965 in Bonus; cer-d.712, -d.1326 in Bonus; cer-d.1818, -d.1819, -d.1820 in Bonus. Mutant used for description and seed stocks: cer-d.5 in Bonus (GSHO 425); cer-d.5 in Bowman (PI 483237)*5 (GSHO 2189).1512131GSHO 425
292GSHO 427cer-f.9Hordeum vulgare L. subsp. vulgare Skåne län, SwedenNSGCKERNELS1994DEVELOPED58.98333300, 13.50000000Genetic materialIn the original stock surface wax coating on the spike, leaf sheath, and stem is reduced (wax code + + ++). Reduced surface wax coating is apparent only on the spike (wax code + ++ ++) in the Bowman backcross-derived line. Inheritance: Monofactorial recessive. Located in chromosome 7HS [1S], near the centromere and about 4.5 cM proximal from the ert-d (erectoides-d) locus, and about 6.3 cM distal from the nud (naked caryopsis) locus. Previous nomenclature and gene symbolization: None. Mutational events: cer-f.9 in Bonus; cer-f.230 in Foma (CIho 11333); cer-f.1002 in Carlsberg II (CIho 10114); cer-f.763 in Bonus; cer-f.1423 in Bonus. Mutant used for description and seed stocks: cer-f.9 in Bonus (GSHO 427); cer-f.9 in Bowman (PI 483237)*6 (GSHO 1846).1512447GSHO 427
293GSHO 428cer-g.10Hordeum vulgare L. subsp. vulgare Skåne län, SwedenNSGCKERNELS1994DEVELOPED58.98333300, 13.50000000Genetic materialMutants have reduced surface wax coating on spike, leaf sheath, and stem (wax code + + ++) with the wax coating of the leaf sheath and stem wax present in broad horizontal bands. Double and triple stomatal complexes are produced during stomatal development. All cer-g mutants have globe-shaped or globosum kernels. Inheritance: Monofactorial recessive. Located in chromosome 2HL, about 15.9 cM distal from the vrs1 (six-rowed spike 1) locus. Previous nomenclature and gene symbolization: None. Mutational events: cer-g.10, -g.11, -g.23, -g.47, -g.55 in Bonus; cer-g.106 in Bonus, -g.202, -g.108, -g.210, -g.494 in Foma (CIho 11333); cer-g.166 in Bonus, -g.553, -g.568, -g.700, -g.709, -g.710 in Foma, -g.1026 in Carlsberg II (CIho 10114); cer-g.746, -g.818, -g.875 in Bonus, -g.1033, -g.1071 in Carlsberg II, -g.1097, -g.1116 in Kristina; cer-g.893, -g.976 in Bonus, -g.1172 in Kristina; cer-g.1161, -g.1169 in Kristina; cer-g.1038 in Carlsberg II, -g.1289 in Kristina, -g.1332 in Bonus; cer-g.1410, -g.1451, -g.1454 in Bonus; cer-g.1495 in Bonus, -g.1510 in Nordal, -g.1736, -g.1740, -g.1741 in Bonus; cer-g.1573 in Nordal. Mutant used for description and seed stocks: cer-g.10 in Bonus (GSHO 428); cer-g.10 in Bowman (PI 483237)*6 (GSHO 1906).1512218GSHO 428
294GSHO 429cer-h.13Hordeum vulgare L. subsp. vulgare Skåne län, Sweden SPIKEHistoric1994DEVELOPED58.98333300, 13.50000000Genetic materialSurface wax coating on the spike appears to be absent (wax code - ++ ++). The original stock has a wide leaf trait, pointed lateral spikelets, and a high level of sterility (10 to 25% seed set). In the Bowman backcross-derived line, plants are semidwarf (about 3/4 normal height) and show the same level of sterility. The original cer-h.13 stock may have a translocation, but sterility in the Bowman stock does not appear to be caused by a translocation. Inheritance: Monofactorial recessive. Location is unknown. Previous nomenclature and gene symbolization: None. Mutational events: cer-h.13 (trans) in Bonus; cer-h.125 in Bonus; cer-h.233, -h.355 in Foma (CIho 11333); cer-h.721 in Bonus; cer-h.1698 in Bonus. Mutant used for description and seed stocks: cer-h.13 in Bonus (GSHO 429); cer-h.13 in Bowman (PI 483237)*5 (GSHO 2190).1512500GSHO 429
295GSHO 43xnt5.eHordeum vulgare L. subsp. vulgare Colorado, United StatesNSGCKERNELS1994DEVELOPEDGenetic materialSeedlings have a pale yellow color and do not survive beyond the two or three leaf stage. The xnt5.e gene must be maintained as a heterozygous stock. Inheritance: Monofactorial recessive. Located in chromosome 6HL, about 25.0 cM distal from the rob (orange lemma) locus, and about 18.8 cM distal from the lax-b (laxatum-b) locus. Previous nomenclature and gene symbolization: Yellow seedling. Xantha seedling n=xn. Mutational events: xnt5.e in Nepal. Mutant used for description and seed stocks: xnt5.e in Nepal (GSHO 43).1510687GSHO 43
296GSHO 430cer-i.16Hordeum vulgare L. subsp. vulgare Skåne län, SwedenNSGCKERNELS1994DEVELOPED58.98333300, 13.50000000Genetic materialSurface wax coating on the spike appears to be absent (wax code - ++ ++). A translocation is present in the cer-i.25 stock, with one of the break points being close to the cer-i locus. Concerning epicuticular wax composition, the wax tubes are much shorter in the mutant, and the amount of two types of diketones (( )-diketone and hydroxy-( )-diketone) is proportionally reduced. Inheritance: Monofactorial recessive. Located in chromosome 5HL [7L], about 5.6 cM distal from the ari-e (breviaristatum-e) locus. Previous nomenclature and gene symbolization: None. Mutational events: cer-i.16, -i.25 (trans), -i.27, -i.30, -i.32, -i.45, -i.51, -i.112h, -i.112l in Bonus; cer-i.92, -i.110 in Bonus; cer-i.156, -i.168, -i.195 in Bonus; cer-i.205, -i.212, -i.255, -i.256, -i.257, -i.263, -i.415, -i.419, -i.430, -i.435, -i.477, -i.486 in Foma (CIho 11333); cer-i.426, -i.434, -i.535, -i.554 in Foma, -i.651, -i.653, -i.655, -i.679, -i.702, -i.704, -i.720, -i.722, -i.732, -i. 734, -i.740 in Bonus; cer-i.802, -i.805, -i.811, -i.814, -i.867, -i.878 in Bonus; cer-i.747, i-.894, -i.921, -i.922, -927, -i.928, -i.936, -i.958 in Bonus, -i.1175, -i.1181 in Kristina; cer-i.891, -i.953 in Bonus, -i.1196, -i.1220, -i.1257 in Kristina; cer-i.1354 in Bonus; cer-i.1397, -i.1711, -i.1743, -i.1823 in Bonus; cer-i.1764 in Bonus. Mutant used for description and seed stocks: cer-i.16 in Bonus (GSHO 430); cer-i.25 in Bonus is used for allelism tests; cer-i.16 in Bowman (PI 483237)*7 (GSHO 2112).1512493GSHO 430
297GSHO 432cer-k.39Hordeum vulgare L. subsp. vulgare Skåne län, SwedenNSGCKERNELS1994DEVELOPED58.98333300, 13.50000000Genetic materialSurface wax coating on the spike only appears to be slightly reduced compared to the parental line (wax code + ++ ++). Thin seeds and slightly narrower leaves are observed in the Bowman backcross-derived line. Inheritance: Monofactorial recessive. Located in chromosome 7HS [1S], based on a linkage drag association with the ant1 (anthocyanin-less 1) locus. Previous nomenclature and symbolization: None. Mutational events: cer-k.39 in Bonus. Mutant used for description and seed stocks: cer-k.39 in Bonus (GSHO 432); cer-k.39 in Bowman (PI 483237)*6 (GSHO 1837).1512446GSHO 432
298GSHO 433cer-l.14Hordeum vulgare L. subsp. vulgare Skåne län, SwedenNSGCKERNELS1994DEVELOPED58.98333300, 13.50000000Genetic materialSurface wax coating on the spike only appears to be slightly reduced (wax code + ++ ++). Inheritance: Monofactorial recessive. Location is unknown. Previous nomenclature and symbolization: None. Mutational events: cer-l.14 in Bonus. Mutant used for description and seed stocks: cer-l.14 in Bonus (GSHO 433); cer-l.14 in Bowman (PI 483237)*3 (GSHO 2191).1512338GSHO 433
299GSHO 434cer-m.15Hordeum vulgare L. subsp. vulgare Skåne län, SwedenNSGCKERNELS1994DEVELOPED58.98333300, 13.50000000Genetic materialSurface wax coating on the spike appears to be reduced or normal, the wax coating on the leaf sheath and stem is reduced (wax code +/++ + ++). The mutant is a semidwarf (5/6 normal height) and shows semi-sterility (25 to 50% seed set). Inheritance: Monofactorial recessive. Location is unknown. Previous nomenclature and gene symbolization: None. Mutational events: cer-m.15 in Bonus. Mutant used for description and seed stocks: cer-m.15 in Bonus (GSHO 434); cer-m.15 in Bowman (PI 483237)*6 (GSHO 2192).1512382GSHO 434
300GSHO 435cer-n.20Hordeum vulgare L. subsp. vulgare Skåne län, SwedenNSGCKERNELS1994DEVELOPED58.98333300, 13.50000000Genetic materialMutants appear to lack surface wax coating on the spike, surface wax on the sheath may be reduced or absent (wax codes - - ++ and - +/- ++). The allele cer-n.20 does not recombine with the eog locus because a translocation is present in the original stock. The cer-n.20 stock may have a translocation breakpoint about 13.0 cM proximal from the vrs1 locus. Inheritance: Monofactorial recessive, except Cer-n.969. Located in chromosome 2HL, gsh9.al is about 8.8 cM distal fromt he eog (elongated outer glume) locus, and about 23.3 cM proximal from the vrs1 (six-rowed spike 1) locus. Previous nomenclature and gene symbolization: Eceriferum-zm=cer-zm. Glossy sheath 9=gs9. Mutational events: cer-n.20 (trans), -n.26, -n.53, -n.101 in Bonus; cer-n.86 (cer-zm.86), cer-n.731 (cer-zm.731) in Bonus; cer-n.93, -n.97 in Bonus, -n.211, -n.236, -n.289 in Foma (CIho 11333); cer-n.155, -n.162, -n.163 -n.170 in Bonus, -n.422, -n.529, -n.547, n.562, -n.580, -n.599, -n.666, -n.715 in Foma, -n.1077 in Carlsberg II (CIho 10114); cer-n.608 in Foma, -n.829, -n.877, -n.880 in Bonus, -n.1085, -n.1139 in Kristina; cer-n.624, -n.926, -n.948, -n.954, -n.957 in Bonus, -n.1173 in Kristina; cer-n.963, -n.985 in Bonus, n.1133 in Kristina; Cer-n.969, cer-n.978 in Bonus, -n.1296, -n.1303, -n.1307 in Kristina, -n.1337 in Bonus; cer-n.998 in Bonus, -n.1093 in Kristina, -n.1391, -n.1437, -n.1449, -n.1456 in Bonus; cer-n.996, -n.1438 in Bonus, -n.1700 in Kristina, -n.1787 in Bonus; cer-n.1765 in Bonus; gsh9.al (OUM029) in Akashinriki (PI 467400, OUJ659). Mutant used for description and seed stocks: cer-n.20 in Bonus (GSHO 435); gsh9.al in Akashinriki (GSHO 1761); cer-n.20 in Bowman (PI 483237)*8 (GSHO 1893); gsh9.al in Bowman *7 (GSHO 1894).1512356GSHO 435
301GSHO 436cer-o.28Hordeum vulgare L. subsp. vulgare Skåne län, SwedenNSGCKERNELS1994DEVELOPED58.98333300, 13.50000000Genetic materialSurface wax coating on the spike appears to be absent or reduced (wax code -/+ ++ ++). All cer-o mutants have pointed lateral spikelets and semi-sterility (50 to 75% seed set). The mutant is a semidwarf (3/4 normal height) in the Bowman backcross-derived line. Inheritance: Monofactorial recessive. Location is unknown. Previous nomenclature and gene symbolization: None. Mutational events: cer-o.28 in Bonus; cer-o.687, -o.787 in Bonus; cer-o.251 in Foma (CIho 11333), -o.1144 in Kristina. Mutant used for description and seed stocks: cer-o.28 in Bonus (GSHO 436); cer-o.28 in Bowman (PI 483237)*5 (GSHO 2193).1512133GSHO 436
302GSHO 437cer-p.43Hordeum vulgare L. subsp. vulgare Skåne län, SwedenNSGCKERNELS1994DEVELOPED58.98333300, 13.50000000Genetic materialSurface wax coating on the leaf blade apears to be reduced (wax code ++ ++ +). Concerning the chemical epicuticular wax composition, a new form of wax crystals, lobe shaped bodies, was observed in the leaf blade. The amount of wax per unit surface area is reduced, but the wax class distribution is the same as in the wild type. Inheritance: Monofactorial recessive. Location is unknown. Previous nomenclature and gene symbolization: None. Mutational events: cer-p.37, -p.43, -p.57 in Bonus; cer-p.77 in Bonus, -p.252, -p.259, -p.300 -p.339 in Foma (CIho 11333); cer-p.522, -p.573 in Foma, -p.707 in Bonus; cer-p.322 in Foma, -p.777, -p.795, -p.874 in Bonus, -p.1118, -p.1142 in Kristina; cer-p.631, -p.646, -p.888, -p.889, -p.905, -p.972 in Bonus; cer-p.939 in Bonus, -p.1201, -p.1206, -p.1207, -p.1210, -p.1235 in Kristina; cer-p.1292 in Kristina, -p.1339, -p.1344 in Bonus; cer-p.1058 in Carlsberg II (CIho 10114), -p.1383, -p.1424 in Bonus; cer-p.1442, -p.1797 in Bonus. Mutant used for description and seed stocks: cer-p.43 in Bonus (GSHO 437); cer-p.37, -p.43, and -p.57 in Bonus were used for wax chemistry and wax structure studies; cer-p.43 in Bowman (PI 483237)*5(GSHO 2194).1512531GSHO 437
303GSHO 439cer-r.19Hordeum vulgare L. subsp. vulgare Skåne län, SwedenNSGCKERNELS1994DEVELOPED58.98333300, 13.50000000Genetic materialSurface wax coating on the spike appears to be greatly reduced or absent, and the wax coating on the leaf sheath and stem appears to be greatly reduced (wax code +/- + ++). The wax coating on the spike appears absent in the Bowman backcross-derived line. Inheritance: Monofactorial recessive. Located in chromosome 3HL, about 4.3 cM distal from the uzu (uzu) locus. Previous nomenclature and gene symbolization: None. Mutational events: cer-r.19 in Bonus; cer-r.127 in Bonus; cer-r.181 in Bonus; cer-r.231 in Foma (CIho 11333); cer-r.801 in Bonus; cer-r.773 in Bonus; cer-r.911 in Bonus, -r.1300 in Kristina; cer-r.1290 in Kristina. Mutant used for description and seed stocks: cer-r.19 in Bonus (GSHO 439); cer-r.19 in Bowman (PI 483237)*6 (GSHO 1977).1512377GSHO 439
304GSHO 44Lks1.aHordeum vulgare L. subsp. vulgare Cambridgeshire, United KingdomNSGCKERNELS1994DEVELOPED52.33333000, -0.08333300Genetic materialLittle or no development of the appendage on the lemma occurs. Heterozygotes may be awnless or awnletted depending upon the source stock for the Lks1 gene and the genetic background. The Lks1.a allele in Engleawnless will not recombine with alleles at the vrs1 (six-rowed spike 1) locus because a short paracentric inversion is present in Engleawnless. The complex vrs1 locus may include awnless and reduced awn length mutants, but the Lks1.b gene in CIho 13311 does recombine with vrs1 locus and is associated with a dominant instead of a recessive allele at the Gth (toothed lemma) locus. Inheritance: Monofactorial dominant. Located in chromosome 2HL, about 9.6 cM distal from the vrs1 (six-rowed spike 1) locus. Previous nomenclature and gene symbolization: Awnless=l(lkeno). Awnless=Lk. Awnless=S. Two-rowed-awnless=Vlk. Mutational events: Lks1.a in Engleawnless; Lks1.b in CIho 13311 (PI 316871). Mutant used for description and seed stocks: Lks1.a in Engleawnless (GSHO 44); Lks1.b in CIho 13311 (GSHO 1579); Lks1.a from CIho 2505 in Betzes (PI 129430)*7(PI 534510), Lks1.a in Compana (PI 537442)*7(PI 534512), Lks1.a in Dekap (CIho 3351)*7 (PI 534514), Lks1.a in Ingrid (CIho 10083)*7 (PI 534516, Ridawn); Lks1.a from Ridawn in Bowman (PI 483237)*5 (GSHO 1913); Lks1.b from CIho 13311 in Bowman*6 (GSHO 1912).1510722GSHO 44
305GSHO 441cer-t.22Hordeum vulgare L. subsp. vulgare Skåne län, SwedenNSGCKERNELS1994DEVELOPED58.98333300, 13.50000000Genetic materialSurface wax coating on the spike appears to be greatly reduced or absent (wax code +/- ++ ++). Concerning the chemical epicuticular wax composition on the spike, the amount of the ( )-diketones and hydroxy-( )-diketones is reduced and the alkan-2-ol containing esters are absent. Wax tubes are much shorter than in the wild types. Inheritance: Monofactorial recessive, except Cer-t.977. Located in chromosome 5HL [7L], about 9.4 cM proximal from the var1 (variegated 1) locus, and about 8.5 cM distal from the cer-zp (eceriferum-zp) locus. Previous nomenclature and gene symbolization: None. Mutational events: cer-t.22, -t.46 in Bonus; cer-t.75, -t.87 in Bonus, -t.216, -t.226, -t.243, -t.264, -t.285, -t.356, -.384, -.493 in Foma (CIho 11333); cer-t.172, cer-t.197, -t.198, -t.199 in Bonus, -t.500, -t.537, -t.564 in Foma, -t.657, -t.664, -t.671, -t.719, -t.726, -t.738, -t.743 in Bonus; cer-t.617 in Foma, -t.708, -t.806, -t.819, -t.871 in Bonus, -t.1088, -t.1115 in Kristina; cer-t.959, -t.961 in Bonus; -t.1176 in Kristina; cer-t.625, -t.641, -t.933 in Bonus, -t.1154, -t.1171, -t.1219, -t.1244 in Kristina; cer-t.1245 in Kristina; cer-t.1360, -t.1393, -t.1401 in Bonus; Cer-t.977, -t.1710, -t.1718 in Bonus. Mutant used for description and seed stocks: cer-t.22 in Bonus (GSHO 441); cer-t.46 in Bonus was used for wax chemistry and wax structure studies; cer-t.22 in Bowman (PI 483237)*3 (GSHO 2119).1512134GSHO 441
306GSHO 442Glossy sheath 8Hordeum vulgare L. subsp. vulgare Skåne län, SwedenNSGCKERNELS1994DEVELOPED58.98333300, 13.50000000Genetic materialThe wax coating on the spike, stem and leaf sheath appears reduced (wax code + + ++). The gsh8 locus is the cer-u component of the complex locus cer-cqu sheath and stem. Deposition of ( )-diketone lipids is reduced or eliminated by mutants in the cer-cqu locus, specifically the cer-u mutants have imparied hydroxyl group insertion. The original cer-u.21 stock has reduced seed set. Inheritance: Monofactorial recessive. Located in chromosome 2HS, based on close association with the gsh6 (glossy sheath 6) locus, in a subterminal position, and over 33.7 cM distal from the Eam1 (early maturity 1) locus. Previous nomenclature and gene symbolization: Eceriferum-u=cer-u. Glossy sheath 8=gs8. Mutational events: cer-u.21, ...gsh8.am (OUM031) in Akashinriki (PI 467400, OUJ659). Mutant used for description and seed stocks: cer-u.21 in Bonus (GSHO 442); gsh8.ag in Akashinriki (GSHO 1760); gsh8.ag in Bowman (PI 483237)*4 (GSHO 1870).1512301GSHO 442
307GSHO 443cer-v.49Hordeum vulgare L. subsp. vulgare Skåne län, SwedenNSGCKERNELS1994DEVELOPED58.98333300, 13.50000000Genetic materialSurface wax coating on the spike appears to be absent or greatly reduced (wax code +/- ++ ++). Pointed lateral spikelets were observed in the original stock, but not in the Bowman backcross-derived line. The vigor of the Bowman backcross derived line is reduced somewhat. Inheritance: Monofactorial recessive. Located in chromosome 2HS, near the fch15 (chlorina seedling 15) locus. Previous nomenclature and gene symbolization: None. Mutational events: cer-v.49 in Bonus; cer-v.489, -v.590 in Foma (CIho 11333); cer-v.515 in Foma; cer-v.752 in Bonus; cer-v.1434 in Bonus. Mutant used for description and seed stocks: cer-v.49 in Bonus (GSHO 443); cer-v.49 in Bowman (PI 483237)*7 (GSHO 1895).1512313GSHO 443
308GSHO 446cer-y.72Hordeum vulgare L. subsp. vulgare Skåne län, SwedenNSGCKERNELS1994DEVELOPED58.98333300, 13.50000000Genetic materialSurface wax coating on the spike appears to be greatly reduced, the wax coating on the leaf sheath and stem is reduced or normal (wax code + +/++ ++). The original cer-y.72 stock has semi-sterility. The Bowman backcross-derived line has normal fertility and surface wax on the leaf sheath and stem appears to be normal (wax code + ++ ++). Inheritance: Monofactorial recessive. Location is unknown. Previous nomenclature and gene symbolization: None. Mutational events: cer-y.72 in Bonus; cer-y.884 in Bonus; cer-y.828 in Bonus. Mutant used for description and seed stocks: cer-y.72 in Bonus (GSHO 446); cer-y.72 in Bowman (PI 483237)*5 (GSHO 2195).1512472GSHO 446
309GSHO 447cer-z.52Hordeum vulgare L. subsp. vulgare Skåne län, SwedenNSGCKERNELS1994DEVELOPED58.98333300, 13.50000000Genetic materialSurface wax coating on the spike, leaf sheath, and stem appears to be absent (wax code - - ++). The original stock shows leaf streaks, late maturity, and semi-sterility (partial seed set). The Bowman backcross-derived line has waxy nodes, normal maturity and fertility, and no leaf streaks. Inheritance: Monofactorial recessive. Located in chromosome 7HS [1S], close to the ant1 (anthocyanin-less 1) locus based on linkage drag. Previous nomenclature and gene symbolization: None. Mutational events: cer-z.52 in Bonus; cer-z.74 in Bonus; cer-z.113 in Bonus; cer-z.512 in Foma (CIho 11333); cer-z.705, -z.981, -z.988, -z.989, -z.990 in Bonus. Mutant used for description and seed stocks: cer-z.52 in Bonus (GSHO 447); cer-z.52 in Bowman (PI 483237)*7 (GSHO 1838).1512110GSHO 447
310GSHO 450cer-zc.65Hordeum vulgare L. subsp. vulgare Skåne län, SwedenNSGCKERNELS1994DEVELOPED58.98333300, 13.50000000Genetic materialSurface wax coating on the spike appears to be greatly reduced or absent (wax code +/- ++ ++). The spike is glossy and the peduncle has reduced surface wax in the Bowman backcross-derived line. Inheritance: Monofactorial recessive. Location is unknown. Previous nomenclature and gene symbolization. Mutational events: cer-zc.65 in Bonus; cer-zc.221, -zc.249, -zc.393 in Foma (CIho11333); cer-zc.442, -zc.501, -zc.518, -zc.545 in Foma; cer-zc.616 in Foma, -zc.785 in Bonus, -zc.1149 in Kristina; cer-zc.642 in Bonus, zc.1305 in Kristina; cer-zc.1758 in Bonus. Mutant used for description and seed stocks: cer-zc.65 in Bonus (GSHO 450); cer-zc.65 in Bowman (PI 483237)*5 (GSHO 2197).1512346GSHO 450
311GSHO 451cer-zd.67Hordeum vulgare L. subsp. vulgare Skåne län, SwedenNSGCKERNELS1994DEVELOPED58.98333300, 13.50000000Genetic materialSurface wax coating on the leaf blade appears to be absent (wax code ++ ++ -). Concerning the chemical epicuticular wax composition, the amount of wax per cm2 is reduced and is accompanied by a decrease of the size and morphological complexity of the wax bodies. They are smaller, simpler, and less dense. The wax class distribution is the same as in the mother variety. Inheritance: Monofactorial recessive. Located in chromosome 3HL, about 9.4 cM proximal from the uzu (uzu) locus, and about 4.6 cM distal from the breakpoint in the ert-c.1 (erecetoides-c) line. Previous nomenclature and gene symbolization: None. Mutational events: cer-zd.67 in Bonus; cer-zd.592 in Foma (CIho 11333), -zd.665, -zd.668 in Bonus; cer-zd.946 in Bonus; cer-zd.1351 in Bonus; cer-zd. 1356 in Bonus. Mutant used for description and seed stocks: cer-zd.67 in Bonus (GSHO 451).1512433GSHO 451
312GSHO 452cer-ze.81Hordeum vulgare L. subsp. vulgare Skåne län, SwedenNSGCKERNELS1994DEVELOPED58.98333300, 13.50000000Genetic materialSurface wax coating on the leaf blade appears to be absent (wax code ++ ++ -). Concerning the chemical epicuticular wax composition, the reduction in the amount of surface wax is accompanied by a decrease in the size and morphological complexity of the wax bodies. The wax body structure is of single, double, and triple finger-like forms. Inheritance: Monofactorial recessive. Located in chromosome 7HS [1S], over 23.7 cM proximal from the brh1 (brachytic 1) locus. Previous nomenclature and gene symbolization: Glossy seedling 5=gl5. Mutational events: cer-ze.81, -ze.145, -ze.152 in Bonus, -ze.203, -ze.247, -ze.253, -ze.298, -ze.331, -ze.333, -ze.391, -ze.392, -ze.460, -ze.506 in Foma (CIho 11333); cer-ze.179, -ze.196, -ze.660, -ze.662, -ze.694, -ze.714, -ze.725, -ze.729, -ze.736, -ze.745 in Bonus, -ze.584 in Foma; cer-ze.620 in Foma, -ze.816, -ze.817, -ze.824, -ze.834, -ze.864, -ze.885 in Bonus, -ze.1086 in Kristina; cer-ze.621 in Foma, -ze.638, -ze.918 in Bonus; cer-ze.934 in Bonus, -ze.1241, -ze.1274 in Kristina; cer-ze.1160, -ze.1297, -ze.1298, -ze.1299 in Kristina, -ze.1322 in Bonus; cer-ze.1349, -ze.1350, -ze.1367, -ze.1380, -ze.1392, -ze.1417, -ze.1422, -ze.1432 in Bonus; cer-ze.1781, -ze.1782, -ze.1804, -ze.1809, -ze.1814, -ze.1815 in Bonus; cer-ze.1817 in Bonus, -ze.1832, -ze.1834, -ze.1836, -ze.1838, -ze.1840, -ze.1842, -ze.1845, -ze.1846, -ze.1848, -ze.1849, -ze.1851, -ze.1854, -ze.1855, -ze.1856, -ze.1869 in Sv 79353; glf5.e in Kirin Choku 1 (Kmut 239, OUM294). Mutant used for description and seed stocks: cer-ze.81 in Bonus (GSHO 452); cer-ze.81 in Bowman (PI 483237)*6 (GSHO 1831); glf5.e in Bowman*7 (GSHO 1830).1512258GSHO 452
313GSHO 453cer-zf.70Hordeum vulgare L. subsp. vulgare Skåne län, SwedenNSGCKERNELS1994DEVELOPED58.98333300, 13.50000000Genetic materialSurface wax coating on the leaf blade appears to be reduced, but only on the three upper leaf blades (wax code ++ ++ +). The original stock in Bonus has short awns, but the Bowman backcross-derived line does not. Inheritance: Monofactorial recessive. Location is unknown. Previous nomenclature and gene symbolization: None. Mutational events: cer-zf.70 in Bonus. Mutant used for description and seed stocks: cer-zf.70 in Bonus (GSHO 453); cer-zf.70 in Bowman (PI 483237)*3 (GSHO 2198).1512425GSHO 453
314GSHO 454cer-zg.214Hordeum vulgare L. subsp. vulgare Skåne län, SwedenNSGCKERNELS1994DEVELOPED58.98333300, 13.50000000Genetic materialSurface wax coating on the leaf blade appears to be reduced, but only on the three upper leaf blades (wax code ++ ++ +). Inheritance: Monofactorial recessive. Located in chromosome 4HL, about 7.0 cM distal from the lks5 (short awn 5, ari-c) locus, and about 14.0 cM distal from the lbi2 (long basal rachis internode 2) locus. Previous nomenclature and gene symbolization: None. Mutational events: cer-zg.214 in Foma; cer-zg.1195 in Kristina. Mutant used for description and seed stocks: cer-zg.214 in Foma (GSHO 454).1512506GSHO 454
315GSHO 456cer-zi.68Hordeum vulgare L. subsp. vulgare Skåne län, SwedenNSGCKERNELS1994DEVELOPED58.98333300, 13.50000000Genetic materialSurface wax coating on the spike, leaf sheath, and stem appears to be somewhat reduced (wax code + + ++). Inheritance: Monofactorial recessive. Located in chromosome 1HL [5L], about 1.3 cM proximal from the cer-e (eceriferum-e) locus, and about 16.1 cM proximal from the nec1 (necrotic leaf spot1) locus. Previous nomenclature and gene symbolization: None. Mutational events: cer-zi.68 in Bonus, -zi.219, -zi.409, -zi.410 in Foma (CIho 11333); cer-zi.138, -zi.674, -zi.701 in Bonus; cer-zi.815, -zi.846, -zi.971 in Bonus; cer-zi.1104, -zi.1119, -zi.1145 in Kristina; cer-zi.1314 in Kristina; cer-zi.979 in Bonus; cer-zi.633, -zi.1414, -zi.1465 in Bonus. Mutant used for description and seed stocks: cer-zi.68 in Bonus (GSHO 456); cer-zi.68 in Bowman (PI 483237)*6 (GSHO 2047).1512219GSHO 456
316GSHO 457cer-zj.78Hordeum vulgare L. subsp. vulgare Skåne län, SwedenNSGCKERNELS1994DEVELOPED58.98333300, 13.50000000Genetic materialSurface wax coating on the leaf blade appears to be absent (wax code ++ ++ -). Concerning the chemical epicuticular wax composition, the amount of wax per cm2 is reduced and is accompanied by a decrease of the size and morphological complexity of the wax bodies. The amount of aldehydes is slightly increased, and no diketones are observed. Wax bodies are smaller and simpler and less densely scattered over the surface than those of the wild type. Inheritance: Monofactorial recessive. Located in chromosome 5HL [7L], about 11.4 cM proximal from the cer-t (eceriferum-t) locus, and about 4.8 cM distal from the ert-g (erectoides-g) locus. Previous nomenclature and gene symbolization: None. Mutational events: cer-zj.78, ... cer-zj.1441 in Bonus. Mutant used for description and seed stocks: cer-zj.78 in Bonus (GSHO 457); cer-zj.78 in Bowman (PI 483237)*5 (GSHO 2113).1512136GSHO 457
317GSHO 458cer-zk.85Hordeum vulgare L. subsp. vulgare Skåne län, SwedenNSGCKERNELS1994DEVELOPED58.98333300, 13.50000000Genetic materialSurface wax coating on the spike, leaf sheath, and stem appears to be greatly reduced, and surface wax on the leaf blade is greatly reduced or absent (wax code + + +/-). In the Bowman backcross-derived line, reduction in the surface wax coating on the leaf sheath and stem was not observed (wax code + ++ +/-). Inheritance: Monofactorial recessive. Located in chromosome 2H, based on the presence of the gth1.a allele at the Gth (toothed lemma) locus in the BC3F3 stock. Previous nomenclature and gene symbolization: None. Mutational events: cer-zk.85, -zk.117 in Bonus; cer-zk.91 in Bonus; cer-zk.157 in Bonus, -zk.593, -zk.611, -zk.612 in Foma (CIho 11333); cer-zk.952 in Bonus; cer-zk.872 in Bonus, -zk.1214, -zk.1218 in Kristina; cer-zk.165 in Bonus, -zk.614 in Foma, -zk.749, -zk.1396 in Bonus; cer-zk.1716, -zk.1720 in Bonus. Mutant used for description and seed stocks: cer-zk.85 in Bonus (GSHO 458); cer-zk.85 in Bowman (PI 483237)*4 (GSHO 2199).1512267GSHO 458
318GSHO 459cer-zl.17Hordeum vulgare L. subsp. vulgare Skåne län, Sweden SPIKEHistoric1994DEVELOPED58.98333300, 13.50000000Genetic materialSurface wax coating on the spike, leaf sheath, and stem appears to be absent (wax code - - ++). The original stock and derived lines containing cer-zl mutants have low seed set and poor plant vigor. Inheritance: Monofactorial recessive. Location is unknown. Previous nomenclature and gene symbolization: None. Mutational events: cer-zl.17 in Bonus; cer-zl.269 in Foma (CIho 11333); cer-zl.1027 in Carlsberg II (CIho 10114). Mutant used for description and seed stocks: cer-zl.17 in Bonus (GSHO 459).1512137GSHO 459
319GSHO 462cer-zo.229Hordeum vulgare L. subsp. vulgare Skåne län, SwedenNSGCKERNELS1994DEVELOPED58.98333300, 13.50000000Genetic materialSurface wax coating on the spike appears to be absent in the original stock (wax code - ++ ++). Pointed lateral spikelets were observed in the original stock, but not in the Bowman backcross-derived line. Only reduced surface wax coating on the spike was observed in the Bowman backcross-derived line (wax code + ++ ++). Inheritance: Monofactorial recessive. Location is unknown. Previous nomenclature and gene symbolization: None. Mutational events: cer-zo.229, -zo.352 in Foma; cer-zo.565 in Foma; cer-zo.718, -zo.882 in Bonus (PI 189763). Mutant used for description and seed stocks: cer-zo.229 in Foma (GSHO 462); cer-zo.229 in Bowman (PI 483237)*3 (GSHO 2201).1512231GSHO 462
320GSHO 463cer-zp.313Hordeum vulgare L. subsp. vulgare Skåne län, SwedenNSGCKERNELS1994DEVELOPED58.98333300, 13.50000000Genetic materialSurface wax coating on the leaf blade appears to be absent (wax code ++ ++ -). Leaf blades are narrow and rolled, and they are maculated with dark and light zones. Expression of the maculated trait is less in the Bowman backcross-derived line, and the plant vigor is much better. Inheritance: Monofactorial recessive. Located in chromosome 5Hl [7L], about 7.5 cM distal from the ert-g (erectoides-g) locus and about 8.9 cM proximal from the cer-t (eceriferum-t) locus. Previous nomenclature and gene symbolization: None. Mutational events: cer-zp.313, -zp.382, -zp.397, -zp.404 in Foma; cer-zp.1205 in Kristina; cer-zp.1795, -zp.1796 in Bonus (PI 189763). Mutant used for description and seed stocks: cer-zp.313 in Bonus (GSHO 463); cer-zp.313 in Bowman (PI 483237)*7 (GSHO 2114).1512320GSHO 463
321GSHO 468ert-a.6Hordeum vulgare L. subsp. vulgare Skåne län, SwedenNSGCKERNELS1994DEVELOPED58.98333300, 13.50000000Genetic materialSpikes of all alleles have a compact appearance caused by a reduction in rachis internode length, estimated from length of ten rachis internodes. Among the alleles at the ert-a locus, rachis internode length ranged from 1.7 to 2.6 mm. The ert-a.6 allele produces plants with shorter internodes than most other ert-a alleles. Plants with an allele at the ert-a locus are 10 to 15 cm shorter than parental cultivars. The effects of the ert-a mutants on spike density are partially reversed by GA3 treatments. Plants of the Bowman backcross-derived line retained the above traits and headed 3 to 6 days earlier than Bowman. Inheritance: Monofactorial recessive. Located in chromosome 7HS [1S], about 11.4 cM distal from the cer-f (eceriferum-f) locus and about 11.7 cM distal from the nud (naked caryopsis) locus. Previous nomenclature and gene symbolization: Erectoides-6=ert-6. Mutational events: ert-a.6, -a.11 (GSHO 215) in Gull, -a.13 in Maja (PI 184884), -a.19, -a.21, -a.23, -a.28 in Bonus (PI 189763), -a.29 in Maja; ert-a.36, -a.38, -a.49 in Bonus; ert-a.77 in Bonus; ert-a.99 in Bonus; ert-a.131, -a.147, -a.160h in Bonus; ert-a.160l -a.161 in Bonus; ert-a.166 in Bonus, -a.315, -a.316 in Foma (CIho 11333); ert-a.323 in Foma; ert-a.334, -a.340a, -a.341, -a.343 in Foma; ert-a.348 in Foma; ert-a.350, -a.364, -a.368, -a.378, -a.388, -a.406, -a.433, -a.481, -a.494 in Foma; ert-a.1180, -a.1187, -a.1193, -a.1197, -a.1203, -a.1210, -a.1219 in Bonus. Mutant used for description and seed stocks: ert-a.6 in Gull (GSHO 468); ert-a.6 in Bowman (PI 483237)*8 (GSHO 1844).1512537GSHO 468
322GSHO 470ert-b.2Hordeum vulgare L. subsp. vulgare Skåne län, SwedenNSGCKERNELS1994DEVELOPED58.98333300, 13.50000000Genetic materialSpikes have a compact appearance caused by a reduction in rachis internode length. Among the alleles at the ert-b locus, rachis internode length varied from 1.9 to 2.3 mm. Culms are shorter than those of parental cultivars. Spike density is decreased by GA3 treatment of plants as the flag leaf emerges. Plants of the Bowman backcross-derived lines have relatively short peduncles and are about 3/4 normal height. Inheritance: Monofactorial recessive. Located in chromosome 1HL [5L], slightly distal from the cer-e (eceriferum-e) locus, and about 19.3 cM proximal from the nec1 (necrotic leaf spot 1) locus. Previous nomenclature and gene symbolization: Erectoides-2=ert-2. Mutational events: ert-b.2, -b.4, -b.5, -b.9 in Gull; ert-b.128, -b.151 in Bonus (PI 189763); ert-b.174 in Bonus, -b.333 in Foma (CIho 11333); ert-b.354, -b.394 in Foma. Mutant used for description and seed stocks: ert-b.2 in Gull (GSHO 470); ert-b.2 in Bowman (PI 483237)*7 (GSHO 2049).1512368GSHO 470
323GSHO 471ert-c.1Hordeum vulgare L. subsp. vulgare Skåne län, SwedenNSGCKERNELS1994DEVELOPED58.98333300, 13.50000000Genetic materialSpikes have a compact or semicompact appearance caused by a reduction in rachis internode length. Among the alleles at the ert-c locus, rachis internode length had a continuous range from 1.7 to 2.9 mm. The spike has fusiform shape with shorter rachis internodes at the base and longer ones near the apex. The culm length is also highly variable among mutants at the ert-c locus. Mutants ert-c.66 and ert-c.153 are partially sterile, much shorter in culm length, and have a more bushy growth pattern than other mutants at this locus. Spike density is decreased by GA3 treatment of plants as the flag leaf emerges. The Bowman backcross-derived line for ert-c.1 is later and taller than Bowman. Inheritance: Monofactorial recessive. Located in chromosome 3HL, about 3.7 cM from the centromere, and about 2.9 cM proximal from the msg5 (male sterile genetic 5) locus. Previous nomenclature and gene symbolization: Erectoides-1=ert-1. Mutational events: ert-c.1 (trans) in Gull; ert-c.14 (trans) (GSHO 473) in Maja (PI 184884), -c.39, -c.47 (trans), -c.48, -c.59 (trans), -c.62, -c.66, -c.70 (trans), -c.75 (trans), -c.82, -c.83, -c.85, -c.88 in Bonus (PI 189763); ert-c.91, -c.95, -c.97, -c.98, -c.108, -c.137, -c.153, -c.173, -c.177 in Bonus; ert-c.176, ert-c.179, -c.188 in Bonus; ert-c.310, -c.311, -c.312 in Foma (CIho 11333); ert-c.320 in Foma; ert-c.327, -c.342, -c.351 in Foma; ert-c.352 in Foma; ert-c.355a, -c.369, -c.386, -c.395 in Foma; ert-c.463 in Foma; ert-c.495 in Foma. Mutant used for description and seed stocks: ert-c.1 in Gull (GSHO 471); ert-c.1 in Bowman (PI 483237)*7 (GSHO 1960).1512329GSHO 471
324GSHO 475ert-d.7Hordeum vulgare L. subsp. vulgare Skåne län, SwedenNSGCKERNELS1994DEVELOPED58.98333300, 13.50000000Genetic materialSpikes have a compact or semicompact appearance caused by a reduction in rachis internode length. Among the alleles at the ert-d locus, rachis internode length ranged from 1.5 to 1.8 mm for mutants of group one and 2.1 to 2.7 mm for mutants in group two. The ert-d.7 allele is in the group two based on rachis internode lengthh and is closely linked to the translocation point. Plants with an allele at the ert-d locus were placed in two plant height groups, but mutants with compact spikes were not all in the short plant height group. Plants with alleles at the ert-d locus responded positively to GA3 in studies on internode length. Inheritance: Monofactorial recessive. Located in chromosome 7HS [1S], near the ert-a (erectoides-a) locus, and about 4.5 cM distal from the cer-f (eceriferum-f) locus. Previous nomenclature and gene symbolization: Erectoides-7=ert-7. Mutational events: ert-d.7 (trans) in Gull; ert-d.14, -d.15, -d.30h (GSHO 476), -d.30l in Maja (PI 184884); ert-d.33, -d.43, -d.60 in Bonus (PI 189763); ert-d.73 in Bonus; ert-d.89 in Bonus; ert-d.96, -d.100, -d.123 (trans), -d.129, -d.152, -d.158 in Bonus, -d.307, -d.322, -d.335, -d.338, -d.339, -d.372, -d.375, -d.385a in Foma (CIho 11333); ert-d.404 in Foma; ert-d.405, -d.420 in Foma. Mutant used for description and seed stocks: ert-d.7 in Gull (GSHO 475); ert-d.7 in Bowman (PI 483237)*7 (GSHO 1845).1512251GSHO 475
325GSHO 477ert-e.17Hordeum vulgare L. subsp. vulgare Skåne län, SwedenNSGCKERNELS1994DEVELOPED58.98333300, 13.50000000Genetic materialSpikes are very compact due to a reduction in rachis internode length. Rachis internode length varied from 1.2 to 1.5 mm among the four mutants examined. Plants are about 2/3 normal height. Partial fertility and reduced vigor are noted among ert-e mutants. The peduncle is very short and spikes often emerge from the side of the flag sheath. A large deficiency of mutant plants is frequently noted in segregating populations. Spike density is greatly decreased by GA3 treatment of plants as the flag leaf emerges. The possibility of allelism at the dsp9 (dense spike 9) locus has not been determined. Inheritance: Monofactorial recessive. Located in chromosome 6HL, over 50.0 cM from the centromere. Previous nomenclature and gene symbolization: Erectoides-17=ert-17. Mutational events: ert-e.17, -e.65 in Bonus; ert-e.94, -e.143 in Bonus, -e.331, -e.396 in Foma (CIho 11333). Mutant used for description and seed stocks: ert-e.17 in Bonus (GSHO 477); ert-e.17 in Bowman (PI 483237)* 7 (GSHO 2091).1512357GSHO 477
326GSHO 478ert-f.18Hordeum vulgare L. subsp. vulgare Skåne län, SwedenNSGCKERNELS1994DEVELOPED58.98333300, 13.50000000Genetic materialSpikes are semicompact with average rachis internode length measurements from 2.2 to 2.6 mm. Culm length is about 5/6 of normal. Sterile lateral spikelets are larger than those of normal sibs, but apices are rounded. Inheritance: Monofactorial recessive, but some partial dominance is expressed. Located in chromosome 4HS?, an association with the int-c (intermedium spike-c) locus is reported, but the association could be caused by a pleiotropic effect of the ert-f gene and not linkage. Previous nomenclature and gene symbolization: Erectoides-18=ert-18. Mutational events: ert-f.18 in Bonus; ert-f.86, -f.145 in Bonus. Mutant used for description and seed stocks: ert-f.18 in Bonus (GSHO 478); ert-f.18 in Bowman (PI 483237)*6 (GSHO 2004).1512140GSHO 478
327GSHO 479ert-g.24Hordeum vulgare L. subsp. vulgare Skåne län, SwedenNSGCKERNELS1994DEVELOPED58.98333300, 13.50000000Genetic materialSpikes are compact due to a reduction in rachis internode length. All mutants at this locus had a similar decrease in rachis intenode length, which varied from 2.1 to 2.4 mm. Awns and lateral spikelets are closely pressed to the spike, the upper part of the straw is spirally twisted, anthers are short ad thick, and the ligule is small. Spikes tend to have a whitish color at maturity. Spike density is decreased by GA3 treatment of plants as the flag leaf emerges. Plants are about 2/3 normal height, have a brachytic-like appearance, have a very strong surface wax covering, and may show reduced seed set. Inheritance:Monofactorial recessive. Located in chromosome 5HL [7L], about 4.7 cM proximal from the cer-i (eceriferum-i) locus, and about 3.1 cM distal from the lax-a (laxatum-a) locus. Previous nomenclature and gene symbolization: Erectoides-24=ert-24. Brachytic 3=br3. Mutational events: ert-g.24, -g.80, -g.84, -g.90 in Bonus; ert-g.78 (GSHO 480), -g.92 in Bonus; ert-g.303, -g.349, -g.359, -g.393 in Foma (CIho 11333). Mutant used for description and seed stocks: ert-g.24 in Bonus (GSHO 479); ert-g.24 in Bowman (PI 483237)*7 (GSHO 2106).1512539GSHO 479
328GSHO 48yst3.cHordeum vulgare L. subsp. vulgare Colorado, United StatesNSGCKERNELS1994DONATED05/1994Genetic materialYellow streaks of variable width and number develop in the leaves, stems, and spikes of homozygous plants. Some plants are almost entirely yellow, similar to chlorina plants. Penetrance is 100%, but expressivity is highly variable and may require careful observation. Cold treatment increases the number and width of yellow streaks. Inheritance: Monofactorial recessive. Located in chromosome 3HS, may be linked to the Btr (brittle rachis) locus based on linkage drag; previous attempts to locate the yst3 locus using trisomic analyses were unsuccessful. Previous nomenclature and gene symbolization: Yellow streak c=yst,,c. Mutational events: yst3.c in Lion. Mutant used for description and seed stocks: yst3.c in Lion (GSHO 48); yst3.c in Bowman (PI 483237)*6 (GSHO 1939).1510810GSHO 48
329GSHO 481ert-h.25Hordeum vulgare L. subsp. vulgare Skåne län, SwedenNSGCKERNELS1994DEVELOPED58.98333300, 13.50000000Genetic materialPlants are semidwarf (about 3/4 normal height), and spike density is increased only slightly. Maturity may be somewhat delayed. Because selection of ert-h plants was difficult in crosses to Bowman, and the identity of the gene in the Bowman backcross-derived line needs to be rechecked. Inheritance: Monofactorial recessive. Location is unknown. Previous nomenclature and gene symbolization: Erectoides-25=ert-25. Mutational events: ert-h.25 in Bonus; ert-h.68 in Bonus. Mutant used for description and seed stocks: ert-h.25 in Bonus (GSHO 481); ert-h.25 in Bowman (PI 483237)*3 (GSHO 2252).1512141GSHO 481
330GSHO 483ert-ii.79Hordeum vulgare L. subsp. vulgare Skåne län, SwedenNSGCKERNELS1994DEVELOPED58.98333300, 13.50000000Genetic materialPlants are about 1/2 normal height and spike has an elongated basal rachis internode. Spikes are relatively short and show a slight reduction in rachis internode length with a range of values from 2.7 to 3.0 mm. Inheritance: Monofactorial recessive. Located in chromosome 3HL, over 6.6 cM distal from the centromere. Previous nomenclature and gene symbolization: Erectoides-79=ert-79. Mutational events: ert-ii.79 in Bonus. Mutant used for description and seed stocks: ert-ii.79 in Bonus (GSHO483); ert-ii.79 in Bowman (PI 483237)*7 (GSHO 1982).1512260GSHO 483
331GSHO 484ert-j.31Hordeum vulgare L. subsp. vulgare Skåne län, SwedenNSGCKERNELS1994DEVELOPED58.98333300, 13.50000000Genetic materialSpikes are semicompact with rachis internode length estimates of 2.4 and 2.6 mm, and culms are about 3/4 normal length. Inheritance: Monofactorial recessive. Located in chromosome 2H, based on linkage drag with the Gth (toothed lemma) locus. Previous nomenclature and gene symbolization: Erectoides-31=ert-31. Mutational events ert-j..31 in Bonus; ert-j.74 in Bonus; ert-j.326 in Foma (CIho 11333). Mutant used for description and seed stocks: ert-j.31 in Bonus (GSHO 484); ert-j.31 in Bowman (PI 483237)*7 (GSHO 1902).1512220GSHO 484
332GSHO 485ert-k.32Hordeum vulgare L. subsp. vulgare Skåne län, SwedenNSGCKERNELS1994DEVELOPED58.98333300, 13.50000000Genetic materialSpikes are semicompact with internode length estimates of 2.4 to 2.8 mm. Culm length is 3/4 to 5/6 of normal. The mutant stock ert-k.32 was released as the cultivar Pallas. Inheritance: Monofactorial recessive. Location is unknown. Previous nomenclature and gene symbolization: Erectoides-32=ert-32. Mutational events: ert-k.32 in bonus; ert-k.76 in Bonus; ert-k.93 in Bonus, -k.309 in Foma (CIho 11333); ert-k.302, -k.435, -k.459, -k.477 in Foma. Mutant used for description and seed stocks: ert-k.32 in Bonus (GSHO 485); ert-k.32 in Bowman (PI 483237)*6 (GSHO 2253).1512142GSHO 485
333GSHO 486ert-l.12Hordeum vulgare L. subsp. vulgare Skåne län, SwedenNSGCKERNELS1994DEVELOPED58.98333300, 13.50000000Genetic materialSpikes are slightly more dense, rachis internode length value from 2.7 to 3.0 mm, and plant height is about 5/6 of normal. Inheritance: Monofactorial recessive. Location is unknown. Previous nomenclature and gene symbolization: Erectoides-12=ert-12. Mutational events: ert-l.12 in Maja; ert-l.45, -l.72 in Bonus (PI 189763); ert-l.114, -l.155, -l.156, -l.162 in Bonus, -l.304, -l.324, -l.371 in Foma (CIho 11333); ert-l.450 in Foma. Mutant used for desciption and seed stocks: ert-l.12 in Maja (GSHO 486); ert-l.12 in Bowman (PI 483237)*4 (GSHO 2254).1512304GSHO 486
334GSHO 487ert-m.34Hordeum vulgare L. subsp. vulgare Skåne län, SwedenNSGCKERNELS1994DEVELOPED58.98333300, 13.50000000Genetic materialSpikes have a compact appearance caused by a reduction in rachis internode length. Rachis internode length averaged from 2.0 to 2.8 mm among the alleles at the ert-m locus. However, the rachis internodes within each spike are often variable in length and the spike appears irregular similar to those of opposite spikelet mutants. Plants with an allele at the ert-m locus are often 10 to 15 cm shorter than parental cultivars, and some tillers of most plants have one or more extremely shortened upper internodes. Alleles at the ert-m locus responded positively to GA3 treatments designed to increase internode length. Some alleles at the ert-m locus lack normal anthocyanin pigmentation. This trait could not be separated from the ert-m allele and was shown to be allele at the ant1 locus. Inheritance: Monofactorial recessive. Located in chromosome 7HS [1S], about 14.7 cM distal from the cer-f (eceriferum-f) locus, and near the ant1 (anthocyanin-less 1) locus. Previous nomenclature and gene symbolization: Erectoides-34=ert-34. Mutational events: ert-m.34, -m.35, -m.40, -m.41, -m.42, -m54, -m.64 in Bonus; ert-m.87, -m.107, -m.115, -m.130, -m.144, -m.168, -m.169 in Bonus, -m.314 in Foma (CIho 11333); ert-m.328 in Foma; ert-m.330, -m.363, -m.384, -m.426 in Foma. Mutant used for description and seed stocks: ert-m.34 in Bonus (GSHO 487); ert-m.34 in Bowman (PI 483237)*8 (GSHO 1843).1512286GSHO 487
335GSHO 488ert-n.51Hordeum vulgare L. subsp. vulgare Skåne län, SwedenNSGCKERNELS1994DEVELOPED58.98333300, 13.50000000Genetic materialSpikes are semicompact due to a reduction in rachis internode length. Most mutants at this locus had a similar decrease in rachis internode length, which varied from 1.9 to 2.4 mm. Mutant lines are about 3/4 normal height. The spike has enlarged lateral spikelets that have a pointed or awnletted tip and plants show partial sterility not associated with a translocation. Spikes tend to have a whitish color at maturity. Spike density is decreased by GA3 treatment of plants as the flag leaf emerges. In the Bowman backcross-derived stock, plants homozygous for the ert-n.51 gene produce few tillers and mature later than normal sibs. Inheritance: Monofactorial recessive. Located in chromosome 5HL [7L], very close to the ert-g (erectoides-g) locus, and about 4.8 cM proximal from the cer-zj (eceriferum-zj) locus. Previous nomenclature and gene symbolization: Erectoides-51=ert-51. Mutational events: ert-n.51 in Bonus; ert-n.61, -n.63 in Bonus; ert-n.133, -n.135 in Bonus; ert-n.325, -n.419 in Foma (CIho 11333); ert-n.409, -n.464 in Foma. Mutant used for description and seed stocks: ert-n.51 in Bonus (GSHO 488); ert-n.51 in Bowman (PI 483237)*3 (GSHO 2107).1512370GSHO 488
336GSHO 489eam8Hordeum vulgare L. subsp. vulgare Skåne län, SwedenNSGCKERNELS1994DEVELOPED58.98333300, 13.50000000Genetic materialEarly heading is associated with decreased culm length, spike length, kernels per spike, and grain yield. When grown in the fall at Kurashiki, Japan, plants head about 20 days earlier than the standard mid-season cultivar, Akashinriki, because they are day-length neutral or photoperiod insensitive. Day-length neutrality is observed in the Swedish early heading mutants, which were induced in spring barley. under controlled environmental conditions, number of days to heading does not change as photoperiod is altered. Plants become yellow green in response to specific growing conditions, 8 to 12 hours of illumination at low temperatures (below 10 degrees C) plus high temperature (20 degrees C or higher) during the dark period. All the mat-a induced mutants are characterized by yellowish-green seedlings at an early stage of development under controlled environmental conditions. The color change is caused by photothermal stress, which increases the zeaxanthin content at the expense of chlorophyll and other pigments. The mutant stock mat-a.8 was released as the cultivar Mari. Inheritance: Monofactorial recessive. Located in chromosome 1HL [5L], about 11.4 cM distal from the trd (third outer glume) locus and 20.9 cM distal from the Blp (black lemma and pericarp) locus. Previous nomenclature and gene symbolization: Early heading k=eak. Early maturity-a=ea-a. Praematurum-a=mat-a. Erectoides-o=ert-o. Mutational events: ert-o.16 in Maja;...eam8.w in Early Russian (CIho 13839). Mutant used for description and seed stocks: ert-o.16 in Maja (GSHO 489); eam8.k in Kinai 5 (OUJ439, GSHO 765); eam8.k in Bonus*5; mat-a.8 in Tochigi Golden*5; eam8.u in Munsing/7*Titan (CIho 16526); eam8.k in Bowman (PI 483237)*7 (GSHO 2063); ert-o.16 in Bowman*7 (GSHO 2064).1512339GSHO 489
337GSHO 49fch15.xHordeum vulgare L. subsp. vulgare Colorado, United StatesNSGCKERNELS1994DEVELOPEDGenetic materialThe normal color of seedlings is altered to "apricot yellow" when plants are grown in the field under low temperature. Plants respond to higher temperatures and may be grown to maturity in suitable environments. The yellow plant color is stable neartly to maturity. Inheritance: Monofactorial recessive. Located in chromosome 2HS, about 18.5 cM distal from the fch1 (chlorina seedling 1) locus. Previous nomenclature and gene symbolization: Orange seedling=or. Mutantional events: fch15.x or or1.a in Trebi. Mutant used for description and seed stocks: fch15.x in Trebi (GSHO 49).1510650GSHO 49
338GSHO 490ert-p.44Hordeum vulgare L. subsp. vulgare Skåne län, SwedenNSGCKERNELS1994DEVELOPED58.98333300, 13.50000000Genetic materialSpikes are semicompact with an estimated rachis internode length of 2.7 mm, and the culm length is 3/4 normal. Inheritance: Monofactorial recessive. Location is unknown. Previous nomenclature and gene symbolization: Erectoides-44=ert-44. Mutational events: ert--.44 in Bonus; ert-p.116, -p.117 in Bonus, -p.301, -p.370, -p.501 in Foma (CIho 11333). Mutant used for description and seed stocks: ert-p.44 in Bonus (GSHO 490); ert-p.44 in Bowman (PI 483237)* 5 (GSHO 2255).1512347GSHO 490
339GSHO 492Ert-r.52Hordeum vulgare L. subsp. vulgare Skåne län, SwedenNSGCKERNELS1994DEVELOPED58.98333300, 13.50000000Genetic materialSpikes are compact in heterozygotes and very compact in homozygotes with rachis internode length ranging from 1.4 to 1.8 mm. Homozygotes are about 2/3 normal height with excellent vigor. The glumes associated with lateral spikelets are 3 to 4 times larger than normal. Lodicule size is reduced. Heterozygotes are intermediate in plant height, have slightly more lax spikes, and have normal glumes in lateral spikelets. Spike density is decreased by GA3 treatment of plants as the flag leaf emerges. Inheritance: Monofactorial incomplete dominant. Location is unknown. Previous nomenclature and gene symbolization: Erectoides-52=ert-52. Mutational events: Ert-r.52, -r.67 in Bonus, -r.329 in Foma (CIho 11333); Ert-r.453 in Foma. Mutant used for description and seed stocks: Ert-r.52 in Bonus (GSHO 492); Ert-r.52 in Bowman (PI 483237)*8 (GSHO 2123).1512143GSHO 492
340GSHO 493ert-s.50Hordeum vulgare L. subsp. vulgare Skåne län, SwedenNSGCKERNELS1994DEVELOPED58.98333300, 13.50000000Genetic materialRachis internode length is reduced slightly to about 3.0 mm in the original mutant and culm length is about 5/6 of normal. Very late segregates were selected in F2 progenies as the representative type for the Bowman backcross-derived line. Awn length is reduced slightly. Inheritance: Monofactorial recessive. Location is unknown. Previous nomenclature and gene symbolization: Erectoides-50=ert-50. Mutational events: ert-s.50 in Bonus. Mutant used for description and seed stocks: ert-s.50 in Bonus (GSHO 493); ert-s.50 in Bowman (PI 483237)*2 (GSHO 2256).1512278GSHO 493
341GSHO 494ert-t.55Hordeum vulgare L. subsp. vulgare Skåne län, SwedenNSGCKERNELS1994DEVELOPED58.98333300, 13.50000000Genetic materialSpikes are semicompact, rachis internode length is about 2.7 mm in the original mutant, and culm length is about 2/3 of normal. These phenotypic traits, included the dense spike and short awn, are inherited together. Based on general appearance of the plants, ert-t was placed in the brachytic class of semidwarf mutants. Awns are about 3/4 normal length. Inheritance: Monofactorial recessive. Location is unknown. Previous nomenclature and gene symbolization: Erectoides-55=ert-55. Brachytic 4=br4. Mutational events: ert-t.55 in Bonus. Mutant used for description and seed stocks: ert-t.55 in Bonus (GSHO 494); ert-t.55 in Bowman (PI 483237)*5 (GSHO 2257).1512144GSHO 494
342GSHO 497ert-v.57Hordeum vulgare L. subsp. vulgare Skåne län, SwedenNSGCKERNELS1994DEVELOPED58.98333300, 13.50000000Genetic materialSpikes appear abnormal with irregular placements of kernels. Rachis internode length is reduced with average values from 1.7 to 3.1 mm. Plants are weak and semi-sterile, anthers are often whitish, and culm length is about 3/4 of normal. The phenotype of this mutant can be classified as an opposite spikelet type. Inheritance: Monofactorial recessive. Location is unknown. Previous nomenclature and gene symbolization: Erectoides-57=ert-57. Mutational events: ert-v.57 in Bonus; ert-v.172 in Bonus, -v.313 in Foma (CIho 11333). Mutant used for description and seed stocks: ert-v.57 in Bonus (GSHO 497); ert-v.57 in Bowman (PI 483237)*6 (GSHO 2258).1512399GSHO 497
343GSHO 498ert-x.58Hordeum vulgare L. subsp. vulgare Skåne län, SwedenNSGCKERNELS1994DEVELOPED58.98333300, 13.50000000Genetic materialSpikes are semicompact with an average rachis internode length of 2.7 mm, and culm length is about 3/4 of normal. Awns are about 2/3 normal length. Inheritance: Monofactorial recessive. Location is unknown. Previous nomenclature and gene symbolization: Erectoides-58=ert-58. Mutational events: ert-x.58 in Bonus. Mutant used for description and seed stocks: ert-x.58 in Bonus (GSHO 498); ert-x.58 in Bowman (PI 483237)*5 (GSHO 2259).1512221GSHO 498
344GSHO 499ert-y.69Hordeum vulgare L. subsp. vulgare Skåne län, SwedenNSGCKERNELS1994DEVELOPED58.98333300, 13.50000000Genetic materialPlants have slightly denser spikes than the parent with an average rachis internode length of 2.8 mm. Culm length is about 5/6 of normal. The mutant phenotype was not observed in crosses to Bowman. Inheritance: Monofactorial recessive. Location is unknown. Previous nomenclature and gene symbolization: Erectoides-69=ert-69. Mutational events: ert-y.69 in Bonus. Mutant used for description and seed stocks: ert-y.69 in Bonus (GSHO 499).1512314GSHO 499
345GSHO 500ert-z.71Hordeum vulgare L. subsp. vulgare Skåne län, SwedenNSGCKERNELS1994DEVELOPED58.98333300, 13.50000000Genetic materialPlants have slightly denser spikes than the parent with an average rachis internode length of 2.8 mm. Culm length is about 3/4 of normal. Variability in rachis internode lengths was observed in the Bowman backcross-derived line. Also, mutant plants have leaf blades that are slightly narrower than those of normal sibs. Inheritance: Monofactorial recessive. Location is unknown. Previous nomenclature and gene symbolization: Erectoides-71=ert-71. Mutational events: ert-z.71 in Bonus. Mutant used for description and seed stocks: ert-1.71 in Bonus (GSHO 500); ert-z.71 in Bowman (PI 483237)*7 (GSHO 2260).1512183GSHO 500
346GSHO 501ert-za.102Hordeum vulgare L. subsp. vulgare Skåne län, SwedenNSGCKERNELS1994DEVELOPED58.98333300, 13.50000000Genetic materialPlants are semidwarf (3/4 normal height) and were placed in the brachytic class of semidwarfs. Peduncles are relatively short, the leaf blade attachment appears more erect than normal, and kernels seem relatively globe shaped. Inheritance: Monofactorial recessive. Location is unknown. Previous nomenclature and gene symbolization: Erectoides-102=ert-102. Brachytic 7=br7. Mutational events: ert-za.102, -za.146 in Bonus. Mutant used for description and seed stocks:1512145GSHO 501
347GSHO 502ert-zb.132Hordeum vulgare L. subsp. vulgare Skåne län, SwedenNSGCKERNELS1994DEVELOPED58.98333300, 13.50000000Genetic materialPlants are 3/4 normal height and spikes tend to emerge prematurely. Inheritance: Monofactorial recessive. Location is unknown. Previous nomenclature and gene symbolization: Erectoides-132=ert-132. Mutational events: ert-zb.132 in Bonus. Mutant used for description and seed stocks: ert-zb.132 in Bonus (GSHO 501); ert-zb.132 in Bowman (PI 483237)*5 (GSHO 2262).1512146GSHO 502
348GSHO 503ert-zc.149Hordeum vulgare L. subsp. vulgare Skåne län, SwedenNSGCKERNELS1994DEVELOPED58.98333300, 13.50000000Genetic materialSpikes are semicompact and plants are semidwarf (3/4 normal height). The basal rachis internode is slightly elongated. Inheritance: Monofactorial recessive. Location is unknown. Previous nomenclature and gene symbolization: Erectoides-149=ert-149. Mutational events: ert-zc.149 in Bonus. Mutant used for description and seed stocks: ert-zc.149 in Bonus (GSHO 503); ert-zc.149 in Bowman (PI 483237)*3 (GSHO 2263).1512252GSHO 503
349GSHO 504ert-zd.159Hordeum vulgare L. subsp. vulgare Skåne län, SwedenNSGCKERNELS1994DEVELOPED58.98333300, 13.50000000Genetic materialPlants have a brachytic-like pattern of growth and are about 3/4 normal height. The basal rachis internode is slightly elongated. Inheritance: Monofactorial recessive. Located in chromosome 2H, based on linkage drag with the Gth (toothed lemma) locus. Previous nomenclature and gene symbolization: Erectoides-159=ert-159. Brachytic 7=br7. Mutational events: ert-zd.159 in Bonus. Mutant used for description and seed stocks: ert-zd.159 in Bonus (GSHO 504); ert-zd.159 in Bowman (PI 483237)*7 (GSHO 1901).1512287GSHO 504
350GSHO 505ert-ze.105Hordeum vulgare L. subsp. vulgare Skåne län, SwedenNSGCKERNELS1994DEVELOPED58.98333300, 13.50000000Genetic materialSpikes are semicompact, but other phenotypic changes were not observed. Inheritance: Monofactorial recessive. Location is unknown. Previous nomenclature and gene symbolization: Erectoides-105=ert-105. Mutational events: ert-ze.105 in Bonus. Mutant used for description and seed stocks: ert-ze.105 in Bonus (GSHO 505); ert-ze.105 in Bowman (PI 483237)*5 (GSHO 2264).1512184GSHO 505
351GSHO 51abo1.aHordeum vulgare L. subsp. vulgare Colorado, United StatesNSGCKERNELS1994DEVELOPEDGenetic materialSeedlings are white in color and devoid of normal pigments. Later, plants with this phenotype were classified as albino mutants. The abo1.a gene must be maintained as a heterozygous stock. The albino seedling mutant (line B17) studied by Daane originated in the cultivar Trebi at the University of Minnesota while Robertson reported that the abo1.a (at) stock was obtained from the University of Minnesota and is in Trebi. Although some phenotypic differences and different map positions were reported, it is possible that both authors studied the same mutant. The mutant studied by Daane has not been found in barley collections; thus, allelism tests are not possible. Inheritance: Monofactorial recessive. Located in chromosome 1HL [5L], over 22.3 cM proximal from the Blp (black lemma and pericarp) locus, and over 36.7 cM proximal from the Blp locus. Previous nomenclature and gene symbolization: White seedling t=at. Albino seedling t=at. Mutational events: abo1.a in Trebi (Trebi 1). Mutant used for description and seed stocks: abo1.a in Trebi (GSHO 51).1510610GSHO 51
352GSHO 531cul2.bHordeum vulgare L. subsp. vulgare Wisconsin, United StatesNSGCKERNELS1994DEVELOPEDGenetic materialPlants usually have but a single culm (stem). Stems are much greater in diameter than normal, and plants are usually earlier than normal. Irregular placement of some spikelets and male fertility in lateral spikelets occur in the Bowman backcross-derived line. Yield of uniculm plants was not restored when grown under high plant populations. Inheritance: Monofactorial recessive. Located in chromosome 6HL, about 1.3 cM distal from the gsh4 (glossy sheath 4) locus, about 11.4 cM from the msg36 (male sterile genetic 36) locus, and 2.2 cM proximal from the rob (orange lemma) locus. Previous nomenclature and gene symbolization: Uniculm 2=uc2. Mutational events: cul2.b in Kindred (GBC379). Mutant used for description and seed stocks: cul2.b in Kindred (GSHO 531, CIho 115530); cul2.b in Bowman (PI 483237)*4 (GSHO 2074); cul2.b plus rob1.a from sel 79Cal in Bowman *8 (GSHO 2075).1512423GSHO 531
353GSHO 566lks2.bHordeum vulgare L. subsp. vulgare Colorado, United StatesNSGCKERNELS1994DONATED05/1994Genetic materialAwns of both central and lateral spikelets are reduced to about 3/5 of the long awned type. Texture of the short awn is finer and more flexible than that of the long awn, especially in non-uzu genotypes. Awn length of heterozygotes in some crosses is shorter than in the normal parent. Inheritance: Monofactorial recessive. Located in chromosome 7HL [1L], estimates range from 7.9 to 10.5 cM distal from the nud (naked caryopsis) locus. Previous nomenclature and gene symbolization: Short awn=a. Short awn=lk. Short awn 2=lk1. Short awn 2=lk2. Short awn 4=lk4. Mutational events: lks2.b in many cultivars of Oriental origin, often associated with the dsp1.a (dense spike 1) gene; a possible mutant in Morex (CIho 15773). Mutant used for description and seed stocks: lks2.b in Honen 6 (OUJ469, PI 307495, GSHO 566); lks2.b from Sermo (CIho 7776) in Betzes (PI 129430)*7 (CIho 16558, GP 36), lks2.b from Sermo in Compana (CIho 5438)*7(CIho 16188, GP 40), lks2.b from Sermo in Decap (CIho 3351)*7 (CIho 16562, GP 44); lks2.b from Wolfe's Multiple Recessive Stock 1 in Bowman (PI 483237)*9 (GSHO 1850).1512441GSHO 566
354GSHO 567ubs4.dHordeum vulgare L. subsp. vulgare JapanNSGCKERNELS1994DEVELOPEDGenetic materialThe stigma has only a few very short branches. This prevents normal pollen reception and reduces seed set to 13 to 30% in uzu type plants. Both the uzu1.a and srh1.a (short rachilla hair) genes interact with ubs4.d to further reduce in seed set. Pollen fertility is normal. Awn length is about 1/4 normal. Seed set on plants of the Bowman backcross-derived line varies from about 10% for plants grown in greenhouses to nearly 50% for plants grown at Aberdeen, Idaho, USA. Inheritance: Monofactorial recessive. Located in chromosome 7HL [1L], about 8.0 cM distal from the nud (naked caryopsis) locus. Previous nomenclature and gene symbolization: Unbranched style 4. Breviaristatum-15=ari-15. Breviaristatum-d=ari-d. Short awn 8=lk8. Mutational events: ubs4.d in Ao Hadaka (Ao Hadaka-hen); ari-d.15, -d.35, -d.51 in Bonus (PI 189763); ari-d.44, -d.57 in Bonus; ari-d.105, -d.107, -d.116, -d.129, -d.130, -d.150, -d.160, -d.186, -d.187, -d.192, -d.193, -d.232, -d.239, -d.240, -d.241, -d.242, -d.243, -d.247 in Foma (CIho 11333), -d.288 in Kristina. Mutant used for description and seed stocks: ubs4.d in Ao Hadaka (GSHO 567); ari-d.15 in Bonus (GSHO 1652); ubs4.d in Bowman (PI 483237)*6 (GSHO 1849); ari-d.15 in Bowman*8 (GSHO 1848).1512223GSHO 567
355GSHO 568wst4.dHordeum vulgare L. subsp. vulgare JapanNSGCKERNELS1994DEVELOPEDGenetic materialMutants grown at low temperatures exhibit numerous, wide, chlorotic (white) streaks on developing leaves. If termperatures are continuously low, plants appear highly chlorotic. As termperature rises, the white streaks gradually disappear. Exposure to 3 degrees Celius for two weeks after sowing is sufficient to induce distinct streaks in the first and second leaves. Inheritance: Monofactorial recessive. Located in chromosome 2HL, about 6.5 cM distal from the eog (elongation outer glume) locus. Previous nomenclature and gene symbolization: None. Mutational events: wst4.d in Kanyo 7 (OUL068). Mutant used for description and seed stocks: wst4.d in Kanyo (GSHO 568); wst4.d in Bowman (PI 483237)*4(GSHO 1905).1512373GSHO 568
356GSHO 57abo10.jHordeum vulgare L. subsp. vulgare Colorado, United StatesNSGCKERNELS1994DEVELOPEDGenetic materialSeedlings are very weak and white in color. The xenia effect of the abo10.j gene on seed size was described in Robertson. Seeds, which are homozygous for this seedling lethal gene, are shriveled with a 1000-kernel weight of 23.4 g compared with 46.5 g for normal kernels. After germination, the endosperm turns watery instead of pasty as found with normal seeds. Later, plants with this phenotype were classified as albino mutants. The abo10.j gene must be maintained as a heterozygous stock. Inheritance: Monofactorial recessive and expressed in the endosperm as a xenia effect. Location is unknown. Previous nomenclature and gene symbolization: White seedling t2=at2. Albino seedling t2=at2. Mutational events: abo10.j in Canadian Thorpe (Canadian Thorpe 1). Mutant used for description and seed stocks: abo10.j in Canadian Thorpe (GSHO 57).1510792GSHO 57
357GSHO 570yst2.bHordeum vulgare L. subsp. vulgare JapanNSGCKERNELS1994DEVELOPEDGenetic materialWhen grown at low temperatures, seedlings develop numerous wide yellow streaks in the seedling leaves. Exposure to 3 degrees Celsius for two weeks after sowing induces distinct streaks in the first and second leaves. As temperatures rise, newly developed leaves have only very fine yellow streaks. These streaks persist until the plant nears maturity. As plants develop, they have a yellow-green color, show delayed maturation, and have low seed yields. Inheritance: Monofactorial recessive. Located in chromosome 3HS, about 0.9 cM distal from the alm (albino lemma) locus. Previous nomenclature and gene symbolization: None. Mutational events: yst2.b in Kuromugi 148/Mensury C (OUJ073). Mutant used for description and seed stocks: yst2.b in Kuromugi 148/Mensury C (GSHO 570); yst2.b in Bowman (PI 483237)*7 (GSHO 1952).1512192GSHO 570
358GSHO 571fch9.kHordeum vulgare L. subsp. vulgare JapanNSGCKERNELS1994DEVELOPEDGenetic materialSeedling leaves are yellow green in color, and the yellow green color persists until near maturity. Plants grown in the field are stunted and maturity is delayed. Inheritance: Monofactorial recessive. Located in chromosome 4HS, over 22.7 cM distal from the Kap (hooded lemma) locus. Previous nomenclature and gene symbolization: Chlorina k=fk. Chlorina 9=f9. Mutational events: fch9.k in Ko A (OUM288, Kmut 174). Mutant used for description and seed stocks: fch9.k in KoA (GSHO 571); fch9.k in Bowman (PI 483237)*7 (GSHO 1996).1512117GSHO 571
359GSHO 572lbi2.bHordeum vulgare L. subsp. vulgare Ontario, CanadaNSGCKERNELS1994DEVELOPEDGenetic materialMutant plants in the cultivar Montcalm have a marked elongation and weakness of the basal rachis internode of the spike. This region may be 10 to 13 cm long in some tillers. The spike hangs vertically downward from the collar on emergence from the sheath and is often broken off in this region by the wind. Even when not broken off, spikes have a high degree of sterility and contain poorly filled grain. Plants show delayed maturity. The ert-i mutants in Bonus have an erect, semicompact (rachis internode length ranging from 2.3 to 2.9 mm) spike, an elongated (2 to 4 cm) basal rachis internode, and reduced plant height (3/4 normal). Removal of modifiers in the Montcalm and Bonus mutant stocks by backcrossing resulted in Bowman backcross- derived lines having similar phenotypes. Inheritance: Monofactorial recessive. Located in chromosome 4HL, less than 12.8 cM proximal from the Kap (hooded lemma) locus, about 2.4 cM proximal from the Kap locus, and about 18.5 cM proximal from the glf1 (glossy leaf 1) locus. Previous nomenclature and gene symbolization: Long basal rachis internode 2=lb2. Erectoides-i=ert-i. Mutational events: lbi2.b in Montcalm (Alb Acc 1028); ert-i.27, -i.53 in Bonus (PI 189763); ert-i.126 in Bonus. Mutant used for description and seed stocks: lbi2.b in Montcalm (GSHO 572); ert-i.27 in Bonus (GSHO 482); lbi2.b in Bowman (PI 483237)*7 (GSHO 2009); ert-i.27 in Bowman*7 (GSHO 2008).1512295GSHO 572
360GSHO 573brh2.bHordeum vulgare L. subsp. vulgare Colorado, United StatesNSGCKERNELS1994DEVELOPEDGenetic materialPlant height and vigor are reduced to about 2/3 normal; the awn is less than 1/4 normal length; the spike is semi-compact; and the leaf, kernel, glume and glume awn, rachilla, and coleoptile are shorter than in the original cultivar. Auricles are well developed and larger than those of the original cultivar. Inheritance: Monofactorial recessive. Located in chromosome 4HL, about 1.5 cM proximal from the glf3 (glossy leaf 3) locus and over 22.8 cM proximal from the Kap (hooded lemma) locus. Previous nomenclature and gene symbolization: Brachytic 2=br2. Breviaristatum-l=ari-l. Mutational events: brh2.b in Svanhals (Kmut 28, OUM283); ari-l.3 in Bonus (PI 189763); ari-l.132 in Foma (CIho 11333); ari-l.135, -l.145, -l.214, -l.237 in Foma, -l.257 in Kristina. Mutant used for description and seed stocks: brh2.b in Svanhals (GSHO 573); ari-l.3 in Bonus (GSHO1660); brh2.b in Bowman (PI 483237)*6 (GSHO 2016); ari-l.3 in Bowman*6 (GSHO 2017).1512224GSHO 573
361GSHO 574yhd1.aHordeum vulgare L. subsp. vulgare JapanNSGCKERNELS1994DEVELOPEDGenetic materialThe spike appears ivory to pale yellow in color; the lemma is ivory-colored, but terminates into green tips with green awns; the plant has a whitish lower leaf sheath, ivory-colored stem nodes and rachis internodes. Mutant plants head and mature slightly earlier than normal sibs. Inheritance: Monofactorial recessive. Located in chromosome 4HL, over 8.7 cM distal from the Hsh (hairy leaf sheath) locus. Previous nomenclature and gene symbolization: Yellow head=yh. Mutational events: yhd1.a in Kimugi (OUL012). Mutant used for description and seed stocks: yhd1.a in Kimugi (GSHO 574); yhd1.a in Bowman (PI 483237)*8 (GSHO 2028).1512482GSHO 574
362GSHO 575sgh1.aHordeum vulgare L. subsp. vulgare Colorado, United StatesNSGCKERNELS1994DONATED05/1994Genetic materialThe sh1.a allele conditions a highly spring habit (grade 1) and plants readily form ear primordia under long-day conditions without prior cold treatment. However, the type cultivar, Iwate Mensury C, and some other cultivars, which are homozygous for the sgh1.a allele, behave as a winter type when sown in the fall (short-day condition). When near-isogenic lines for growth habit were grown at Kurashiki, Japan, the sgh1.a allele had little influence maturity of phenotype in the presence of the other spring habit growth genes. Cultivars having only the sgh1.a allele for spring habit are rare. The sgh1.a allele is found rarely in spring barley cultivars from Tibet. Inheritance: Monofactorial recessive. Located in chromosome 4HL, about 5.1 cM distal from the Hsh (hairy leaf sheath) locus. Previous nomenclature and gene symbolization: Light sensitiveness=ls. Spring habit of growth 1=sh1. Mutational events: sgh1.a in many cultivars in the so-called Occidental region, and in most cases sgh1.a is accompanied by the dominant allele at the Sgh2 (spring growth habit 2) locus. Mutant used for description and seed stocks: sgh1.a in Iwate Mensury C (OUJ308, GSHO 575); sgh1.a in Hayakiso 2 (OUJ064)*11; sgh1.a in Dairokkaku 1 (OUJ645)*11.1512391GSHO 575
363GSHO 576Hln1.aHordeum vulgare L. subsp. vulgare Colorado, United States SPIKEHistoric1994DONATED05/1994Genetic materialA few hairs of 1 to 2 mm are mixed with the ordinary teeth or barbs on the lateral nerves of the lemma. Expression of the Hln1.a gene may be easier to observe as 1mm hairs on the tip of sterile lateral kernels in two-rowed barley. The Hln gene is associated with a recessive short awn trait (2/3 of normal length). Inheritance: Monofactorial dominant when crossed to a cultivar having abundant barbs on the lemma nerves. Located in chromosome 4HL, over 8.7 cM proximal from the yhd1 (yellow head 1) locus, and over 22.5 cM distal from the mlo (reaction to Erysiphe graminis hordei-o) locus; complete linkage with the dominant allele at the Hsh (hairy leaf sheath) locus has been observed. Previous nomenclature and gene symbolization: Hairs on l lemma nerves=Hn. Mutational events: Hln1.a in Kogane-mugi (OUL010) and in several other cultivars. Mutant used for description and seed stocks: Hln1.a in Kogane-mugi (GSHO 576, PI 225020); Hln1.a from CIho 6884 in Bowman (PI 483237)*4 (GSHO 2027).1512193GSHO 576
364GSHO 577glf3.cHordeum vulgare L. subsp. vulgare JapanNSGCKERNELS1994DEVELOPEDGenetic materialSurface wax coating on the leaf blades appears to be absent from the seedling stage to near maturity, and leaves have a shiny appearance (wax code ++ ++ -). Concerning the chemical epicuticular wax composition, the cer-j mutant in Bonus produces 34% less wax compared to the wild type, the primary alcohols are the largest wax class. The wax occurs as many very thin plates and as few large irregularly shaped bodies. Inheritance: Monofactorial recessive. Previous nomenclature and gene symbolization: Glossy leaf-gl. Glossy leaf 3=gl3. Glossy leaves 4=gl4. Eceriferum-j=cer-j. Mutational events: glf3.c in Goseshikoku (Goseshikoku-hen, OUL032); glf3.d (gl4, GSHO 1376) in Gateway (CIho 10072; cer-j.59, -j.62 in Bonus (PI 189763); cer-j.71, -j.142, -j.148 in Bonus, -j.274, -j.288, -j.301, -j.311, -j.340, -j.346, -j.349, -j.358, -j.370, -j.390, -j.429, -j.447, -j.470 in Foma (CIho 11333); cer-j.183, -j.184, -j.696 in Bonus, -j.275, -j.399, -j.517, -j.521, -j.558 in Foma, -j.1024, -j.1081 in Carlsberg II (CIho 10114); cer-j.458 in Foma, -j.780, -j.797, -j.832 in Bonus, -j.1090, -j.1102 in Kristina; cer-j.909 in Bonus, j.1152 in Kristina; cer-j.1204, -j.1231, -j.1233, -j.1239, -j.1251 in Kristina; cer-j.997, -j.1331, -j.1361, -j.1388, -j.1405, -j.1433, -j.1469 in Bonus; cer-j.1712, -j.1744, -j.1761, -j.1803, -j.1808 in Bonus; cer-j.1827 in Bonus, -j.1831, -j.1835, -j.1839, -j.1841, -j.1843, -j.1844, -j.1847, -j.1850, -j.1852, -j.1857, -j.1858, -j.1859, -j.1860, -j.1861, -j.1862 in Sv 79353. Mutant used for description and seed stocks: glf3.c in Goseshikoku (GSHO 577); cer-j.59 in Bonus (GSHO 431) is used for allelic localization tests and for wax chemistry and wax structure studies; glf3.d in Bowman (PI 483237)*6 (GSHO 2019); cer-j. 59 in Bowman*4 (GSHO 2021).1512118GSHO 577
365GSHO 578fst2.bHordeum vulgare L. subsp. vulgare JapanNSGCKERNELS1994DEVELOPEDGenetic materialThe leaf and stem characteristics of fst2 plants are similar to those of fst1 plants. Leaves and stems are very fragile and easily broken when slightly bent. Young leaves generally wither in the afternoon in the greenhouse and in the field during the spring. Most of the leaf blade tips turn yellow prior to heading, and gradual die back is observed. Plants are about one-half normal height when protected in the greenhouse, but in the field plants are weak and are easily damaged by wind. The stems are often curved, and the seeds are small and thin. Inheritance: Monofactorial recessive. Located in chromosome 1HL [5L], over 19.0 cM proximal from the nec1 (necrotic leaf spot 1) locus. Previous nomenclature and gene symbolization: Fragile stem 2= fs2. Mutational events: fst2.b in Oshichi (Oshichi-hen, OUL006). Mutant used for description and seed stocks: fst2.b in Oshichi (GSHO 578); fst2.b in Bowman (PI 483237)*7 (GSHO 2046).1512242GSHO 578
366GSHO 579eam7.gHordeum vulgare L. subsp. vulgare California, United StatesNSGCKERNELS1994DEVELOPEDGenetic materialUnder short-day conditions in California, USA, mutant plants bloom about 4 weeks before California Mariout, produce relatively few tillers, are fine stemmed, and are relatively short. Differences are less pronounced under long-day conditions. Mutants are partially insensitive to photoperiod and have a vernalization requirement. Under long-day conditions, difficulty in identification of segregates expressing the eam7.g gene was reported. When the Eam1.a (early maturity 1) gene is not present, eam7.g plants head to 10 to 14 days earlier than Bowman in nurseries at Yuma, Arizona, USA, but only 3 to 5 days earlier at Fargo, North Dakato, USA. The Eam1.a gene present in California Mariout apparently interacts with the eam7.g allele under short-day conditions to cause extreme earliness. Inheritance: Monofactorial recessive. Located in chromosome 6HS, about 0.5 cM from the rob (orange lemma) locus, and about 3.0 cM from the centromere. Previous nomenclature and gene symbolization: Early heading=ec. Early maturity 7=ea7. Mutational events: eam7.g in BC6 California Mariout (GBC326); eam7.n (Ea1), eam7.o (Ea2), eam7.p (Ea3) in Chikurin Ibaraki 1 (OUJ069, CIho 7370). Mutant used for description and seed stocks: eam7.g in BC6 California Mariout (GSHO 579); eam7.g in Bowman (PI 483237(*4 (GSHO 2068).1512327GSHO 579
367GSHO 580lbi1.aHordeum vulgare L. subsp. vulgare Colorado, United StatesNSGCKERNELS1994DONATED05/1994Genetic materialThe first or basal rachis internode is much longer than the other rachis internodes with an expression range from less than 1 cm to more than 20 cm. Genotype, environmental conditions, and tiller health alter basal rachis internode elongation in plants homozygous for lbi1.a. Expression of the lbi1.a allele is often greater in six-rowed genotypes than in two-rowed ones. Inheritance: Monofactorial recessive. Located in chromosome 5HL [7L], about 16.0 cM distal from the raw1 (smooth awn 1) locus. Previous nomenclature and gene symbolization: Curved peduncle=cr. Long basal spike internode=bi. Long basal spike internode=lb. Rachisextensum=rac-a and rac-b. Mutational events: lbi1.a in Wisconsin Pedigree 38; rac-a.2 and rac-b.1. in Bonus (PI 189763) are not alleles of each other; but both rac-a.2 and rac-b.1 were reported to be alleles at the lbi1 locus. Mutant used for description and seed stocks: lbi1.a in Wisconsin Pedigree 38 (GSHO 580); lbi1.a in Bowman (PI 483237)*5 (GSHO 2144).1512495GSHO 580
368GSHO 581xnt7.gHordeum vulgare L. subsp. vulgare Montana, United StatesNSGCKERNELS1994DEVELOPEDGenetic materialSeedlings have a yellow color and die at the two to three leaf stage. The xnt7.g gene must be maintained as a heterozygous stock. Inheritance: Monofactorial recessive. Located in chromosome 1HL [5L], about 2.0 cM from the msg1 (male sterile genetic 1) locus. Previous nomenclature and gene symbolization: Xantha seedling 66=x66. Albino seedling a=alb,,a. Xantha seedling g=xan,,g. Mutational events: xnt7.g in Erbet. Mutant used for description and seed stocks: xnt7.g in Erbet (GSHO 581).1512502GSHO 581
369GSHO 582xnt8.hHordeum vulgare L. subsp. vulgare Montana, United StatesNSGCKERNELS1994DEVELOPEDGenetic materialSeedlings have a yellow color and die at the two to three leaf stage. The xnt8.h gene must be maintained as a heterozygous stock. Allelism tests with mutants at the xnt3 (xantha seedling 3) locus, present in the same region of chromosome 3H, have not been conducted. Inheritance: Monofactorial recessive. Located in chromosome 3HS, about 2.9 cM from the msg5 (male sterile genetic 5) locus. Previous nomenclature and gene symbolization: Albino seedling b=alb,,b. Xantha seedling h=xan,,h. Mutational events: xnt8.h in Carlsberg II. Mutant used for description and seed stocks: xnt8.h in Carlsberg II (GSHO 582).1512485GSHO 582
370GSHO 583abo12.oHordeum vulgare L. subsp. vulgare Montana, United StatesNSGCKERNELS1994DEVELOPEDGenetic materialSeedlings have a white color and die at the two to three leaf stage. The abo12.o gene must be maintained as a heterozygous stock. Inheritance: Monofactorial recessive. Located in chromosome 5HS [7S], over 26.1 cM distal from the msg16 (male sterile genetic 16) locus, and not linked to the msg19 (male sterile genetic 19) locus. Previous nomenclature and gene symbolization: Albino seedling c=alb,,c. Albino seedling o=alb,,o. Mutational events: abo12.o in Titan. Mutant used for description and seed stocks: abo12.o in Titan (GSHO 583).1512281GSHO 583
371GSHO 584xnt9.iHordeum vulgare L. subsp. vulgare Montana, United StatesNSGCKERNELS1994DEVELOPEDGenetic materialSeedlings have a yellow color and die at the two to three leaf stage. The xnt9.i gene must be maintained as a heterozygous stock. Inheritance: Monofactorial recessive. Located in chromosome 7HL, over 7.4 cM from the msg23 (male sterile genetic 23) l locus. Previous nomenclature and gene symbolization: Albino seedling d = alb,,d. Xantha seedling i = xan,,i. Mutational events: xnt9.i in Erbet. Mutant used for description and seed stocks: xnt9.i in Erbet (GSHO 584).1512464GSHO 584
372GSHO 586abo14.qHordeum vulgare L. subsp. vulgare Montana, United StatesNSGCKERNELS1994DEVELOPEDGenetic materialSeedlings have a white color and die at the two to three leaf stage. The abo14.q gene must be maintained as a heterozygous stock. Inheritance: Monofactorial recessive. Located in chromosome 6HL, about 12.5 cM distal from the rob (orange lemma) locus, and about 8.1 cM distal from the sex1 (shrunken endosperm xenia 1) locus. Previous nomenclature and gene symbolization: Albino seedling f=alb,,f. Albino seedling q=alb,,q. Mutational events: abo14.q in Shabet (G365). Mutant used for description and seed stocks: abo14.q in Shabet (GSHO 586).1512361GSHO 586
373GSHO 591wst5.eHordeum vulgare L. subsp. vulgare DenmarkNSGCKERNELS1994DEVELOPEDGenetic materialWhite streaks with poorly defined margins may develop on the foilage, but expression is influenced by environment and possibly genetic background. Some environmental conditions cause the plants to be nearly albinotic from the seedling stage until heading when the plants turn green, while under other conditions the streaks seem to be absent. Inheritance: Monofactorial recessive. Located in chromosome 1HL [5L], about 34.5 cM distal from the nec1 (necrotic leaf spot 1) locus, and about 21.7 cM proximal from the Blp (black lemma and pericarp) locus. Previous nomenclature and gene symbolization: None. Mutational events: wst5.e in Carlsberg II (Mutant no 10). Mutant used for description and seed stocks: wst5.e in Carlsberg II (GSHO 591).1512516GSHO 591
374GSHO 597des6.iHordeum vulgare L. subsp. vulgare Arizona, United StatesNSGCKERNELS1994DEVELOPEDGenetic materialThe chromosomes are paired during pachytene and undergo desynapsis during diplotene. The degree of desynapsis is 7.9 + 2.1 ranging from 7 ring bivalents (d=0) to 14 univalents (d=14). Many univalents split longitudinally during Anaphase I. Lagging chromosomes and micronuclei are observed frequently at Telophase I. Microspore quartets contain an average of 3.3 micronuclei per quartet with a range of 0 to 12. Ovule fertility is about 16%, and about 22% of the selfed seeds are trisomics. Inheritance: Monofactorial recessive. Located in chromosome 1H [5]. Previous nomenclature and gene symbolization: None. Mutational events: des6.i in Betzes; des6.m in OAC 21 (CIho 1740), des6.o in Betzes. Mutant used for description and seed stocks: des6.i in Betzes (GSHO 597).1512225GSHO 597
375GSHO 598des7.jHordeum vulgare L. subsp. vulgare Arizona, United StatesNSGCKERNELS1994DEVELOPEDGenetic materialThe chromosomes are paired during the pachytene stage of microsporogenesis, but they undergo desynapsis during diplotene. The degree of desynapsis is d=6.1 + 2.6, ranging from 7 ring bivalents (d=0) to 14 univalents (d=14). Many univalents split l longitudinally during anaphase I. Lagging chromosomes and micronuclei are observed frequently at Telophase I. Microspore quartets contain an average of 3.0 micronuclei per tetrad with a range of 0 to 13. Female fertility is about 33%, and about 13% of the selfed seeds are trisomics. Inheritance: Monofactorial recessive. Located in chromosome 2H. Previous nomenclature and gene symbolizaiton: None. Mutational events: des7.j in Betzes (GSHO 598). Mutant used for description and seed stocks: des7.j in Betzes (GSHO 598).1512243GSHO 598
376GSHO 6lig1.myHordeum vulgare L. subsp. vulgare JapanNSGCKERNELS1994DEVELOPEDGenetic materialThe ligule and auricle of all leaves are absent, and the leaf blades are erect along the stem. Liguleless plants can be identified visually at all stages of growth. Reverse mutation of some mutants is possible. Fine structure analysis of mutants at the lig locus was conducted by Konishi. Inheritance: Monofactorial recessive. Located in chromosome 2HL; about 25.1 cM distal from the mtt4 (mottled leaf 4) locus, and about 9.2 cM proximal from the Zeo1 (Zeocriton 1) locus. Previous nomenclature and gene symbolization: Ligule and auricle less=al. Liguleless=li. Exauriculum=aur-a. Mutational events: lig1.my as Muyoji (OUL007); lig1.ky in Koyo (PI 190819), lig1.a1 (OUM001), lig1.a2 in Akashinriki (PI 467400), OUJ659); lig1.c1, lig1.c2, lig1.c3, lig1.c4 in Chikurin Ibaraki 1 (OUJ030, CIho 7370); aur-a.1 (lig1.b1), aur-a.2 (lig1.b2), aur-a.7 (lig1.b7), aur-a.8 (lig1.b8), aur-a.9 (lig1.b9) in Bonus (PI 189763), aur-a.3 (lig1.b3), aur-a.4 (lig1.b4), aur-a.5 (lig1.b5), aur-a.6 (lig1.b6), aur-a.10 (lig1.b10) in Foma (CIho 11333); aur-a.11, aur-a.12 in Kristina, aur-a.13, aur-1.14 in Bonus, aur-a.15 in Golf (PI 488529). Mutant used for description and seed stocks: lig1.my as Muyoji (GSHO 6); lig1.my in Bowman (PI 483237)*8 (GSHO 1930).1510646GSHO 6
377GSHO 608rvl1.aHordeum vulgare L. subsp. vulgare Colorado, United StatesNSGCKERNELS1994DEVELOPEDGenetic materialTips of young leaves tend to roll into a tube through a counter-clockwise spiral, compared to flat surfaces of the control. Segregates can be scored from the three-leaf stage until near maturity. Inheritance: Monofactorial recessive. Located in chromosome 1HL [5L], proximal from the trd (third outer glume) locus, and distal from the abo1 (albino seedling 1) locus. Previous nomenclature and gene symoblization: None. Mutational events: rvl1.a in Hakata 2 (Kmut 103). Mutant used for description and seed stocks: rvl1.a in Hakata 2 (GSHO 608).1512244GSHO 608
378GSHO 61abo8.hHordeum vulgare L. subsp. vulgare Colorado, United StatesNSGCKERNELS1994DEVELOPEDGenetic materialSeedlings are white in color and devoid of normal pigments. Later, plants with this phenotype were classified as albino mutants. The abo8.h gene must be maintatined as a heterozygous stock. Inheritance: Monofactorial recessive. Located in chromosome 7HS[1S], about 27.2 cM distal from the nud (naked caryopsis) locus, over 11.3 cM distal from the nud locus, and about 18.0 cM distal from the fch8 (chlorina seedling 8) locus. Previous nomenclature and gene symbolization: White seedling c2=ac2. Albino seedling c2=ac2. Albino seedling-m=alb-m. Mutational events: abo8.h in Coast (Coast II); alb-m.27 in Bonus (PI 539132). Mutant used for description and seed stocks: abo8.h in Coast (GSHO 61).1510684GSHO 61
379GSHO 622mtt1.aHordeum vulgare L. subsp. vulgare Ontario, CanadaNSGCKERNELS1994DEVELOPEDGenetic materialSeedlings have two to four clearly marked white bands across or partially across the leaves. Necrosis of the white region occurs later and the leaf is constricted in these regions. Temperatures at germination over 20 degrees C reduce the extent of mottling. Plant mortality is low at Edmonton, Alberta, Canada, where temperatures are low, but nearly 100% at St. Paul, Minnesota, USA. Inheritance: Monofactorial recessive. Located in chromosome 1HS [5S], based on linkage drag with the sls1.a (small lateral spikelet) gene. Previous nomenclature and gene symbolization: Mottled leaves=mt. Mottled leaves 3=mt3. Mutational events: mtt1.a (mt) in Montcalm (Alb Acc 284). Mutant used for description and seed stocks: mtt1.a in Montcalm (GSHO 622, OUM070); mtt1.a in Bowman (PI 483237)*4 (GSHO 2036).1512148GSHO 622
380GSHO 66xnt3.cHordeum vulgare L. subsp. vulgare Colorado, United StatesNSGCKERNELS1994DONATED05/1994Genetic materialSeedlings have a yellow (picric yellow) to yellow-green color depending upon temperatures, but they can be grown to maturity under certain environmental conditions. The xnt3.c gene must be maintained as a heterozygous stock. Inheritance: Monofactorial recessive. Located in chromosome 3HS, about 9.4 cM proximal from the abo9 (albino seedling 9) locus, about 14.4 cM distal from the uzu (uzu) locus, and about 3.7 cM proximal from the alm (albino lemma) locus. Previous nomenclature and gene symbolization: Xantha seedling c=xc. Viridis seedling-l=vir-l Mutational events: xnt3.c in Colsess (Colsess IV); vir-l.27 in Maja (PI 184884). Mutant used for desciption and seed stocks: xnt3.c in Colsess (GSHO 66).1510661GSHO 66
381GSHO 68yvs1.aHordeum vulgare L. subsp. vulgare Colorado, United StatesNSGCKERNELS1994DONATED05/1994Genetic materialVirescent seedlings have a slightly green tip, but fail to survive beyond the seedling stage. Recessive alleles at the yvs1 locus must be maintained as heterozygous stocks. Inheritance: Monofactorial recessive. Located in chromosome 2HS, over 13.7 cM proximal from the fch15 (chlorina seedling 15) locus and about 2.0 cM proximal from the fch1 (chlorina seedling 1) locus. Previous nomenclature and gene symbolization: Yellow seedling 1=y. Xantha (yellow) seedling x=yx. Yellow seedling x=yx. Albino seedling-c=alb-c. Mutational events: yvs1.a in a Swedish cultivar; yvs1.b (yx)(GSHO 62) in Faust I (CIho 4579); alb-c.7 in Gull (CIho 1145). Mutant used for description and seed stocks: yvs1.a in Minn 72-8 (GSHO 68).1510837GSHO 68
382GSHO 681lgn4.dHordeum vulgare L. subsp. vulgare Minnesota, United StatesNSGCKERNELS1994DEVELOPEDGenetic materialSeedlings are light green in color, and the light green color persists until near maturity. In the Bowman backcross-derived lines, the plants are weaker than normal sibs and produce thinner seeds. Inheritance: Monofactorial recessive. Located in chromosome 4HL, about 5.0 cM distal from the Kap (hooded lemma) locus and about 4.0 cM proximal from the glf1 (glossy leaf 1) locus. Previous nomenclature and gene symbolization: Light green 1=lg1. Light green 4=lg4. Light green 9=lg9. Mutational events: lgn4.d (lg1) in Himalaya X Ingrescens; lgn4.i (lg9) in No 156, an unknown cultivar (GSHO 1, GSHO 94). Mutant used for description and seed stocks: lgn4.d in Himalaya X Ingrescens (GSHO 681); lgn4.d in Bowman (PI 483237)*7 (GSHO 2011).1512411GSHO 681
383GSHO 684Vrs1.tHordeum vulgare L. subsp. vulgare Utah, United StatesNSGCKERNELS1994DEVELOPEDGenetic materialPlants homozygous for Vrs1.t (Vt) allele always have a deficiens phenotype regardless of the allele present at the int-c (intermedium-c) locus in chromosome 4. Deficiens is characterized by the extreme reduction in size of the lateral spikelet. The Vrs1.t allele in Engleawnless does not recombine with the Lks1.a allele at the Lks1 (awnless) locus because a short paracentric inversion is present in this stock. Inheritance: The Vrs1.t allele is the most dominant allele in the multiple allelic series at the vrs1 (six-rowed spike 1) locus, but interacts as an incomplete dominant with some alleles. Located in chromosome 2HL, about 15.4 cM distal from the Gth (toothed lemma) locus, and 30.5 cM distal from the eog (elongated outer glume) locus. Previous nomenclature and gene symbolization: Hordeum deficiens=Vt. Renamed Vrs1.t based on allelism tests. Mutational events: Vrs1.t or Vt in White Deficiens (CIho 7316). Mutant used for description and seed stocks: Vrs1.t or Vt in White Deficiens (GHSO 684); Vrs1.t in Bowman (PI 483237)*8(GSHO 1909).1512235GSHO 684
384GSHO 70abo2.bHordeum vulgare L. subsp. vulgare Colorado, United StatesNSGCKERNELS1994DONATED05/1994Genetic materialSeedlings lack normal pigmentation; they are white and die at a two or three leaf stage. Robertson suggested that an albino 3 stock [abo4 9a30] may have been used in his linkage studies instead of albino 1 (a1). The abo2.b gene must be maintained as a heterozygous stock. Inheritance: Monofactorial recessive. Located in chromosome 2HS, about 13.0 cM distal from the fch1 (chlorina seedling 1) locus, about 5.5 cM proximal from the fch15 (chlorina seedling 15) locus, and over 9.4 cM distal from the msg2 (male sterile genetic 2) locus. Previous nomenclature and gene symbolization: Albino 1=a1. Albino seedling 2=a2. Mutational events abo2.b in Nilsson-Ehle No 2. Mutant used for description and seed stocks: abo2.b in Nilsson-Ehle No 2 (GSHO 70).1510824GSHO 70
385GSHO 701Rpg1.aHordeum vulgare L. subsp. vulgare Wisconsin, United StatesNSGCKERNELS1994DEVELOPEDGenetic materialThe dominant allele at the Rpg1 locus confers moderate resistance to most races of Puccinia graminis f sp tritici. Lesion size and percent stem and leaf sheath infection in adult plants is reduced compared with susceptible plants under field conditions. At high temperatures (greater than 20 degrees C), differentiation of seedling reaction types is observed with race 56 (10). These differences correlate with adult plant reactions better than at lower temperatures. Differences among cultivars indicate that a modifier of the Rpg1.e allele exists. Plants with Rpg1 gene show a susceptible reaction to pathotype QCC of Puccinia graminis f sp tritici and to Puccinia graminis f sp secalis. Molecular markers closely linked to the three different sources of the Rpg1 region gene suggest that they originated independently. Synteny of the Rpg1 region of chromosome 5HS to the short arm or rice chromosome 6 was demonstrated. Inheritance: Monofactorial dominant. Located in chromosome 7HS [1S], about 13.2 cM distal to the brh1 (brachytic 1) locus, about 9.2 cM distal from the Run1 (reaction to Ustilago nuda 1) locus, about 1.4 cM distal from the Run1 locus, and 0.7 cM from RFLP marker ABG077. Previous nomenclature and gene symbolization: Resistance to Puccinia graminis tritici=T. Resistance to Puccinia graminis Pers f sp tritici Eriks and E Henn. = Rpg1a. Mutational events: Rpg1.a in Chevron; Rpg1.e in Peatland, Rpg1.f in Kindred. Mutant used for description and seed stocks: Rpg1.a in Chevron (GSHO 701); Rpg1.a in 80-TT-29 (CIho 16129); Rpg1.e from Peatland in Bowman (PI 483237).1514437GSHO 701
386GSHO 707rob1.aHordeum vulgare L. subsp. vulgare California, United StatesNSGCKERNELS1994DEVELOPEDGenetic materialThe lemma, palea, and rachis have an orange pigmentation that is present in immature spikes, can be observed at heading, and is retained in mature grain and spikes. The orange pigmentation is visible at the base of sheath of seedlings and in exposed nodes after jointing. Internodes have a layer of orange tissue and stems have an orange color as the straw dries. The mutant stock for rob1.f (OUM189) has a lighter orange lemma color than that in other mutants at the rob locus. Inheritance: Monofactorial recessive. Located in chromosome 6HS, about 10.8 cM proximal from the msg36 (male sterile genetic 36) locus, and about 2.2 cM distal from the cul2 (uniculm 2) locus. Previous nomenclature and gene symbolization: Orange lemma=pl. Orange lemma=br. Orange lemma=o. Robiginosum-o=rob-o. Mutational events: rob1.a in CIho 5649 (GBC340); rob1.b (OUM185), rob1.c (OUM186), rob1.d (OUM187), rob1.e (OUM188), rob1.f (OUM189) in Akashinriki (OUJ659, PI 467400); rob1.1, rob1.2 in Bonus (PI 189763), rob1.3, rob1.4, rob1.5 in Foma (CIho 11333), rob1.6 in Kristina. Mutant used for description and seed stocks: rob1.a in CIho 5649 (GSHO 707); rob1.a in Bowman (PI 483237)*7 (GSHO 2069).1513863GSHO 707
387GSHO 71dsp10.cHordeum vulgare L. subsp. vulgare Colorado, United StatesNSGCKERNELS1994DONATED05/1994Genetic materialAlthough rachis internode length or spike density is not a highly stable character, plants with relatively dense spikes can be selected from crosses to Club Mariout. Rachis internode length values of 2.8 to 3.4 mm, 2.9 to 3.6 mm, and 2.6 to 2.8 mm were recorded for Club Mariout in one experiment. Inheritance: Monofactorial recessive. Located in chromosome 3HS, about 1.3 cM distal from the xnt3 (xantha seedling 3) locus. Previous nomenclature and gene symbolization: Dense spike c=lc. Mutational events: dsp10.c in Club Mariout. Mutant used for description and seed stocks: dsp10.c in Club Mariout (GSHO 71); dsp10.c in Bowman (PI 483237)*3 (GSHO 1956).1510760GSHO 71
388GSHO 735gsh1.aHordeum vulgare L. subsp. vulgare Minnesota, United StatesNSGCKERNELS1994DEVELOPEDGenetic materialA bright green glossy color is apparant on the spike, leaf sheath, stem (wax code --++). The gsh1 locus is the cer-q component of the complex locus named cer-cqu, which controls formation of ()-diketone lipids in epicuticular wax layer. The cer-q mutants affect the ()-ketoacyl condensing enzyme. Inheritance: Monofactorial recessive, except Cer-q.1440. Located in chromosome 2HS, in a subterminal position over 36.5 cM distal from Eam1 (early maturity 1) locus. Previos nomenclature and gene symbolization: Glaucous sheath = gs. Glossy sheath and spike=gs. Glossy sheath 1 = gs1. Eceriferum-q=cer-q. Mutational events: gsh1.a in PI 95285; gsh1.b in Piroline PI 539132.....gsh1.ba in Bozu Omugi 1 (OUJ655). Mutant used for description in seed stock: gsh1.a in PI 95285 (GSHO 735); gsh1.a in Bowman (PI 483237)*7 (GSHO 1868).1513810GSHO 735
389GSHO 736gsh2.fHordeum vulgare L. subsp. vulgare Montana, United StatesNSGCKERNELS1994DEVELOPEDGenetic materialSurface wax coating appears to be absent on the spike, leaf sheath, and stem (wax code - - ++). Inheritance: Monofactorial recessive. Located in chromosome 3HL, about 5.1 cM proximal from the als (absent lower laterals) locus and over 32.5 cM distal from the uzu (uzu) locus. Previous nomenclature and gene symbolization: Glossy sheath 2=gs2. Eceriferum-b=cer-b. Mutational events: gsh2.d in Vantage, ...gsh2.bb in the cross NDB112/Fr926-77. Mutant used for description and seed stocks: gsh2.f in Atlas (GSHO 735); cer-b.2 in Maja (GSHO 423; GSHO 1081); gsh2.f in Bowman (PI 483237)*7 (GSHO 1988); cer-b.2 in Bowman*6 (GSHO 1989).1513976GSHO 736
390GSHO 737gsh3.iHordeum vulgare L. subsp. vulgare Ontario, CanadaNSGCKERNELS1994DEVELOPEDGenetic materialSurface wax coating appears to be absent on the spike, leaf sheath, and stem (wax code - - ++). Concerning the chemical epicuticular wax composition, only traces of the ( )-diketone wax classes and alkan-2-ol containing esters occur in the cer-a.6 mutant in Bonus. Inheritance: Monofactorial recessive. Located in chromosome 7HS [1S], about 2.1 cM proximal from the fch12 (chlorina seedling 12) locus, and 6.5 to 12.9 cM proximal from the brh1 (brachytic 1) locus. Previous nomenclature and gene symbolization: Glossy sheath 3=gs3. Eceriferum-a=cer-a. Mutational events: gsh3.i in Mars;...gsh3.aj (OUM027) in Akashinriki (PI 467400, OUJ659). Mutant used for description and seed stocks: gsh3.i in Mars (GSHO 737); cer-a.1 in Maja (GSHO 422); gsh3.i in Bowman (PI 483237)*8 (GSHO 1825); cer-a.1 in Maja is used for allelism tests; cer-a.6 in Bonus was used for wax chemistry and wax structure studies.1513843GSHO 737
391GSHO 738gsh4.lHordeum vulgare L. subsp. vulgare Minnesota, United StatesNSGCKERNELS1994DEVELOPEDGenetic materialSurface wax coating appears to be appears to be absent on the spike, leaf sheath, and stem (wax code - - ++). Concerning the chemical epicuticular wax composition, only traces of the ( )=diketone wax classes and alkan-2-ol containing esters occur in the cer-x mutant in Bonus. Inheritance: Monofactorial recessive. Located in chromosome 6HL, near the centromere, and less than 3.0 cM distal from the rob (orange lemma) locus. Previous nomenclature and gene symbolization: Glossy sheath 4=gs4. Eceriferum-x=cer-x. Mutational events: gsh4.l Gateway; cer-x.60 in Bonus; cer-x.102, -x.114 in Bonus, -x.228 (GSHO 445), -x.309, -x.350, -x.417, -x.449, -x.450, -x.507 in Foma (CIho 11333); cer-x.164, -x.658 in Bonus, -s.561 in Foma, -x.1049, -x.1055, -x.1066 in Carlsberg II (CIho 10114); ce-x.788 in Bonus, -x.1095 in Kristina; cer-x.1180 in Kristina; cer-x.1236 in Kristina; cer-x.886 in Bonus; cer-x.1398, -x.1435 in Bonus, -x.1508 in Nordal; cer-x.1000, -x.1498 in Bonus, -x.1518, -x.1523, -x.1529, -x.1536, -x.1547, -x.1552 in Nordal, -x.1732, -x.1756 in Bonus. Mutant used for description and seed stocks: gsh4.l in Gateway (GSHO 738); cer-x.60 in Bonus (GSHO 1520); gsh4.l in Bowman (PI 483237)*7 (GSHO 2072); cer-x.60 in Bowman *7 (GSHO 2073).1514455GSHO 738
392GSHO 739gsh5.mHordeum vulgare L. subsp. vulgare Skåne län, SwedenNSGCKERNELS1994DEVELOPED58.98333300, 13.50000000Genetic materialMutants have a reduced surface wax covering on the spikes and stems, and a bright glossy green color on all leaf sheaths (wax code + - ++). The nodes appear to have excessive wax covering. The cer-s.31 allele is associated with delayed heading in the Bowman backcross-derived line, but the gsh5.m allele is not. Inheritance: Monofactorial recessive. Located in chromosome 2HL; gsh5 is in the centromeric region, about 2.5 cM proximal from the eog (elongated outer glume) locus, and about 32.0 cM proximal from the vrs1 (six-rowed spike 1) locus. Previous nomenclature and gene symbolization: Glossy sheath 5=gs5. Eceriferum-s=cer-s. Glossy sheath 8=gs8. Mutational events: cer-s.24 inBonus; cer-s.31 in Bonus; cer-s.41 in Bonus; cer-s.287, -s.303, -s.372, -s.444 in Foma (CIho 11333); cer-s.1047 in Carlsberg II (CIho 10114); cer-s.137 in Bonus, -s.591 in Foma, -s.803 in Bonus, -s.1132 in Kristina; cer-s.914 in Bonus; cer-s.622 in Bonus, -s.1054 in Carlsberg II, -s.1153 in Kristina; cer-s.1323 in Bonus; cer-s.1134 in Kristina; cer-s.1754 in Bonus; gsh5.m in Jotun; gsh5.n in Okaiku 3 (GSHO 832). Mutant used for description and seed stocks: gsh5.m in Jotun (GSHO 739); cer-s.31 in Bonus (GSHO 440); cer-s.31 in Bowman (PI 483237)*5 (GSHO 1888); gsh5.m in Bowman*8 (GSHO 1889).1513922GSHO 739
393GSHO 740gsh6.sHordeum vulgare L. subsp. vulgare Skåne län, SwedenNSGCKERNELS1994DEVELOPED58.98333300, 13.50000000Genetic materialA bright, glossy green color is apparent on the spike, leaf sheath, and stem. The gsh6 locus is the cer-c component of the complex locus cer-cqu, which controls formation of ( )-diketone lipids in epicuticular wax layer. Most alleles have the wax code - - ++, but a few alleles show delayed development of surface waxes on the sheath (wax code - +/- ++). Inheritance: Monofactorial recessive. Located in chromosome 2HS, in a subterminal position, and over 33.7 cM distal from the Eam1 (early maturity 1) locus. Previous nomenclature and gene symbolization: Glossy sheath 6=gs6. Eceriferum-c=cer-c. Eceriferum-cqu=cer-cqu. Mutational events: cer-c.3 in Bonus; ...gsh6.aq in an Ethiopian introduction (PI 151787, OUE120). Mutant used for description and seed stocks: gsh6.s in Betzes (GSHO 740); cer-c.3 in Bonus (GSHO 212, GSHO 424, GSHO 959); gsh6.s in Bowman (PI 483237)*7 (GSHO 1869).1513977GSHO 740
394GSHO 741msg22.eHordeum vulgare L. subsp. vulgare Montana, United StatesNSGCKERNELSNot Available1994DEVELOPEDGenetic materialSelfing-none. Outcrossing-complete female fertility. Stamens-anthers smaller than fertile sibs, no stomium or filament elongation. Inheritance: Monofactorial recessive. Located in chromosome 7H [1]. Previous nomenclature and gene symbolization: Male sterile e=msg,,e. Male sterile 22=ms22. Mutational events: msg22.e in a selection from Glacier/Compana (MSS073); msg22.fc (MSS404) in Proctor (PI 280420), msg22.fo (MSS416) in Zephyr (PI 339815). Mutant used for description and seed stocks: msg22.e in Bowman (PI 483237)*7 (GSHO 1857).1514336GSHO 741
395GSHO 744msg25.rHordeum vulgare L. subsp. vulgare Montana, United StatesNSGCKERNELS1994DEVELOPEDGenetic materialSelfing-0.7% for msg25.r, 2.6% for msg25.dz. Outcrossing-complete female fertility. Stamens-anthers smaller than fertile sib, but some have stomium. Some filament elongation may occur. Inheritance: Monofactorial recessive. Located in chromosome 4HL, near the centromere and proximal from the Blx1 (Blue aleurone xenia 1) locus. Previous nomenclature and gene symbolization: Male sterile=msg,,r. Mutational events: msg25.r in Betzes (MSS086); msg25.dz (MSS374) in Klages (CIho15487). Mutant used for description and seed stocks: msg25.r in Betzes (GSHO 744); msg25.r in Bowman (PI 483237)*7 (GSHO 2020).1513979GSHO 744
396GSHO 745msg26.uHordeum vulgare L. subsp. vulgare Montana, United StatesNSGCKERNELS1994DEVELOPEDGenetic materialSelfing-none. Outcrossing-complete female fertility. Stamens-anthers smaller than fertile sibs, no stomium or filament elongation. Inheritance: Monofactorial recessive. Located in chromosome 7HS [1S], linked to the nud (naked caryopsis) locus, and l linked to the ant1 (anthocyanin-less 1) locus based on linkage drag. Previous nomenclature and gene symbolization: Male sterile u=msg,,u. Mutational events: msg26.u in Unitan (MSS088). Mutant used for description and seed stocks: msg26.u in Unitan (GSHO 745, GSHO 2378); msg26.u in Bowman (PI 483237)*6 (GSHO 1858).1513923GSHO 745
397GSHO 750seg1.aHordeum vulgare L. subsp. vulgare Montana, United States Historic1994DEVELOPEDGenetic materialSeed is long and thin and the 100-kernel weight is about 33% of normal. Good stands can be established in the field if optimum environmental conditions prevail during germination and emergence. This mutant is associated with an increase in percentage lysine in the protein. Inheritance: Monofactorial recessive. Located in chromosome 7HL [1L], linked to the msg23 (male sterile genetic 23) locus. Previous nomenclature and gene symbolization: Shrunken endosperm=se1. Mutational events: seg1.a in Betzes. Mutant used for description and seed stocks: seg1.a in Betzes (GSHO 750); seg1.a in Bowman (PI 483237)*7 (GSHO 1852).1513980GSHO 750
398GSHO 751seg2.bHordeum vulgare L. subsp. vulgare Montana, United States Historic1994DEVELOPEDGenetic materialSeed size is greatly reduced with a 100-kernel weight about 15% of normal. Fertility is normal. There is little or no starch deposited in the seed that develops on homozygous plants. When seed is germinated on blotters in a 10% sucrose solution, seedling establishment was about 25% after transplanting to soil. Inheritance: Monofactorial recessive. Located in chromosome 7HS [1S], about 5.7 cM proximal from the msg10 (male sterile genetic 10) locus, and about 2.8 cM proximal from the sex6 (shrunken endosperm xenia 6) locus. Previous nomenclature and gene symbolization: Shrunken endosperm=se2. Mutational events: seg2.b in Betzes. Mutant used for description and seed stocks: seg2.b in Betzes (GSHO 751), which is a balanced tertiary trisomic (BBT) stock T16c.1513820GSHO 751
399GSHO 752seg3.cHordeum vulgare L. subsp. vulgare Montana, United States Historic1994DEVELOPEDGenetic materialSeed size is reduced to about 33% of normal when grown under field conditions. Seeds are long and thin similar to those from seg1 plants; seeds are viable, and good stand establishment is possible. Inheritance: Monofactorial recessive. Located in chromosome 3H, over 30.8 cM from the centromere. Previous nomenclature and gene symbolization: Shrunken endosperm=se3. Mutational events: seg3.c in Compana. Mutant used for description and seed stocks: seg3.c in Compana (GSHO 752); seg3.c in Bowman (PI 483237)*7 (GSHO 1957).1514393GSHO 752
400GSHO 753seg4.dHordeum vulgare L. subsp. vulgare Montana, United StatesNSGCKERNELS1994DEVELOPEDGenetic materialUnder field conditions in the original mutant, seed size is reduced to about 38% of normal and seed set is about 50% of normal. Stand establishment is poor under field conditions. Xenia effects were not observed in the original study, but the stock presently held as GSHO 753 and the Bowman backcross-derived line show xenia effects in hybrid plants. The lemma side of the kernel has a very distinct depression in mature seed. Inheritance: Monofactorial recessive. Located in chromosome 7HL [1L], over 34.0 cM from the centromere. Previous nomenclature and gene symbolization: Shrunken endosperm=se4. Mutational events: seg4.d in Compana. Mutant used for description and seed stocks: seg4.d in Compana (GSHO 753); seg4.d in Bowman (PI 483237)*5 ( (GSHO 1853).1513844GSHO 753
401GSHO 754seg5.eHordeum vulgare L. subsp. vulgare Montana, United StatesNSGCKERNELS1994DEVELOPEDGenetic materialSeed size is reduced in homozygotes with the 100-kernel weight being about 50% of normal. However, seed size is quite variable within a spike. Seed set is reduced to about 20%, probably due to seed abortion. The caryopsis freqently exceeds the lemma and palea in length. The combination of caryopsis length and reduced seed set is used to classify mutant plants. Inheritance: Monofactorial recessive. Located in chromosome 7HS [1S], probably in the centromeric region of the short arm. Previous nomenclature and gene symbolization: Shrunken endosperm=se5. Mutational events: seg5.e in a Sermo/7*Glacier line. Mutant used for description and seed stocks: seg5.e in a Sermo/7* Glacier line (GSHO 754).1513981GSHO 754
402GSHO 755sex1.aHordeum vulgare L. subsp. vulgare Montana, United StatesNSGCKERNELS1994DEVELOPEDGenetic materialAfter the hard dough stage, kernels develop a depression in the center of the lemma which becomes progressively more distinct with maturity. The 100-kernel weight of the mutant is about 75% of normal. The mature endosperm of the mutant appears to be harder than normal. The mutant expresses xenia permitting classification of kernels from heterozygous plants as normal or shrunken with an expected 3:1 ratio. Field establishment and growth of mutant plants is normal. Mutant plants have a higher lysine content in the endosperm than normal sibs. Inheritance: Monofactorial recessive. Located in chromosome 6HL, about 5.6 cM proximal from the msg36 (male sterile genetic 36) locus. Previous nomenclature and gene symbolization: Shrunken endosperm 6=se6. Shrunken endosperm 6=se6f. High lysine 5=lys5. Mutational events: sex1.a in Compana; lys5.f (Riso 13) in Bomi (PI 43371), sex1.d or lys5.h (Riso 86), sex1.e or lys5.g (Riso 29) in Carlsberg II (CIho 10114). Mutant used for description and seed stocks: sex1.a in Compana (GSHO 755); sex1.e in Bowman (PI 483237)*3 (GSHO 2071).1514338GSHO 755
403GSHO 764Sgh3.cHordeum vulgare L. subsp. vulgare JapanNSGCKERNELS1994DEVELOPEDGenetic materialCultivars with the Sgh3.c allele exhibit a highly spring habit (grade 1) and plants form ear primordia under long-day conditions without prior cold treatment. When near-isogenic lines for growth habit were grown at Kurashiki, Japan, the Sgh3.c allele had major influence on maturity and phenotype in the absence or presence of the other spring habit of growth genes. Plants were 10 days or more earlier, tillers per plant were 30% less or more, kernel number was reduced by about 20%, and spike length was reduced. Spring cultivars having only the Sgh3.c gene have never been found. Spring cultivars with the Sgh2.b Sgh3.c genotype occur most frequently in North Pakistan, North India, Tibet, Ethiopia, and Northern Europe. The Sgh3.c allele is always accompanied by the Sgh2.b or sgh1.a and Sgh2.b alleles in cultivars. Inheritance: Monofactorial dominant. Located in chromosome 1HL [5L], over 35.4 cM proximal from the Blp (black lemma and pericarp) locus. Previous nomenclature and gene symbolization: Spring habit of growth 3=Sh3. Mutational events: Sgh3.c occurs in many spring cultivars distributed mostly in regions of extremely high latitude or high altitude. Mutant used for description and seed stocks: Sgh3.c in a selection from the cross Tammi X Hayakiso 2 (GSHO 764), which has the winter habit alleles at the sgh1 and Sgh2 loci.1513880GSHO 764
404GSHO 766wst2.bHordeum vulgare L. subsp. vulgare JapanNSGCKERNELS1994DEVELOPEDGenetic materialWhen grown at low temperatures, mutants exhibit wide numerous stripes on the developing leaves, becoming almost albinic or chlorotic. Exposure to 3 degrees Celsius for about two weeks immediately after sowing induces distinct streaks on the first and second leaves. As temperatures rise, subsequent leaves have only a few, narrow white stripes. Inheritance: Monofactorial recessive. Located in chromosome 5HL [7L], about 3.6 cM proximal from the srh (short rachilla hair) locus. Previous nomenclature and gene symbolization: White streak 2=wst2. Mutational events: wst2.b in OUL067. Mutant used for description and seed stocks: wst2.b in OUL067 (GSHO 766); wst2.b in Bowman (PI 483237)*5 (GSHO 2102).1514456GSHO 766
405GSHO 769nld1.aHordeum vulgare L. subsp. vulgare JapanNSGCKERNELS1994DEVELOPEDGenetic materialMutant plants have narrow, dark green leaves, which are erect with well developed midribs. Auricles are degenerated to tiny projections, but ligules are normal. Stem internodes are short, and the upper ones are curved. Spikelets are relatively narrow and small and seed set may be low. Plants are short (less than 1/2 normal height), the spike commonly emerges from the side of the sheath before anthesis, and kernels are thin. Inheritance: Monofactorial recessive. Located in chromosome 5HL [7L], about 21.6 cM proximal from the fst1 (fragile stem 1) locus. Previous nomenclature and gene symbolization: None. Mutational events: nld1.a in Nagaoka dwarf (OUL054). Mutant used for description and seed stocks: nld1.a in Nagaoka dwarf (GSHO 769); nld1.a in Bowman (PI 483237)*7 (GSHO 2093).1513874GSHO 769
406GSHO 770Sgh2.lHordeum vulgare L. subsp. vulgare Colorado, United StatesNSGCKERNELS1994DONATED05/1994Genetic materialWhen grown under long-day conditions in the greenhouse, plants with the Sgh2.l gene for spring habit (grade I type) head in about 40 days. Plants with the Sgh2.ll gene (grade II type) are 10 days or more later than those with Sgh2.l, and require about 10 days of pre-chilling to make up the difference in heading time. Complex interactions with genotype and other spring growth havits have been reported. The Sgh2.ll allele is prevalent in the Oriental region, while the Sgh2.l allele is confined to the Occidental region. Inheritance: Monofactorial dominant, but a multiple allelic series exists with the alleles for the highest grade of spring habit dominant over those for less spring-like habit. The dominant alleles are epistatic to the winter habit alleles at the sgh1 and Sgh3 loci. Located in chromosome 5HL [7L], over 15.1 cM distal from the raw1 (smooth awn 1) locus. Previous nomenclature and gene symbolization: Spring habit of growth 2=Sh2. Mutational events: Sgh2.l or Sgh2.ll are present in many spring barley cultivars and probably some wild progenitors. Mutant used for description and seed stocks: Sgh2.l in Indian Barley (OUJ694, GSHO 770); Sgh2.ll in Marumi 16 (OUJ007, GSHO 767).1513755GSHO 770
407GSHO 773vrs2.eHordeum vulgare L. subsp. vulgare JapanNSGCKERNELS1994DEVELOPEDGenetic materialThe spike appears similar to the ordinary six-rowed spike, but the lateral spikelets on the upper and lower portion of the spike are reduced in size, less fertile, and more or less pedicellated. Laterals are pointed with reduced awn length or awnless. Inheritance: Monofactorial recessive. Located in chromosome 5HL [7L], over 19.0 cM proximal from the srh (short rachilla hair) locus. Previous nomenclature and gene symbolization: Six-rowed spike 2=v2. Mutational events: vrs2.e in Svanhals (Kmut 27, OUM284). Mutant used for description and seed stocks: vrs2.e in Svanhals (GSHO 773); vrs2.e in Bowman (PI 483237)*4 (GSHO 2111).1513794GSHO 773
408GSHO 774vrs3.fHordeum vulgare L. subsp. vulgare JapanNSGCKERNELS1994DEVELOPEDGenetic materialThe upper half of the spike appears to be six-rowed, but the lateral spikelets in the lower half are reduced in size with the basal portion being two-rowed. Awns on the lateral spikelets range from normal near the top, to awnletted or pointed in the middle, to awnless at the base of the spike. Lateral spikelets set seed only in the upper two-thirds of the spike. Lateral spikelets may be pedicellated in some stocks. Central spikelets often have double awn, one on the lemma and one on the palea. The rachilla may be deformed. Inheritance: Monofactorial recessive. Located in chromosome 1HL [5L]; near the centromere and proximal from the ert-b (erectoide-b) locus. Previous nomenclature and gene symbolization: Six-rowed spike 3=v3. Intermedium spike-a=int-1. Intermedium spike-g=int-g. Mutational events: vrs3.f in Hakata 2 (Kmut 213, OUM292), vrs3.g (Xb 388.8, Piro-1), vrs3.h (Xd 160.46, Piro-3), vrs3.i (X15 1463, Piro-5), vrs3.j (X17 3555, Piro-8) in Piroline (PI 539132); int-a.1 in Bonus (PI 189763); int-a.2 in Bonus; int-a.8, -a.9, -a.10, -a.64 in Bonus, -a.14, -a.17, -a.21, -a.27, -a.30, -a.31, -a.32, -a.34, -a.35, -a.37 in Foma (CIho 11333), -a.46, -a.51, -a.52, -a.54, -a.55, -a.59, -a.61 in Kristina; int-a.71, -a.74, -a.77, -a.79, -a.86, -a.88 in Bonus, -a.102, -a.103 in Hege. Mutant used for description and seed stocks: vrs3.f in Hakata 2 (GSHO 774); int-a.1 in Bonus (GSHO 170); vrs3.f in Bowman (PI 483237)*7 (GSHO 2056); int-a.1 in Bowman*5 (GSHO 2055).1513988GSHO 774
409GSHO 775vrs4.kHordeum vulgare L. subsp. vulgare Japan Historic1994DEVELOPEDGenetic materialAlleles at this locus enhance the development of lateral spikelets and formation of additonal spikelets to various degrees. The int-e mutants, which were isolated in two-rowed cultivars, have enlarged lateral spikelets that may set seed in the upper two-thirds of the spike. Kernels in lateral spikelets are smaller than those from central spikelets in int-e mutants. The rachilla may be deformed by partial formation of an extra spikelet. The awn size on lateral spikelets range from a pointed apex to 3/4 normal length. The mul1.a and vrs4.k alleles, which were isolated in six-rowed cultivars, may produce numerous extra spikelets at the base of the lateral spikelets and on the rachilla. Phenotypic differences among alleles are retained in the Bowman backcross-derived lines. Inheritance: Monofactorial recessive. Located in chromosome 3HL, about 27.5 cM distal from the uzu (uzu) locus. Previous nomenclature and gene symbolization: Multiflorus=mul. Intermedium spike-e=int-e. Six-rowed spike 4=v4. Mutational events: mul1.a (Alb Acc 321, GSHO 561) in Montcalm (CIho 7149); vrs4.k (Gamma III 3B, MFB-2) in MFB 104 (PI 232921), vrs4.l (Xc 41.5, Piro-2) in Piroline (PI 262210); vrs4.m (X17 2497, Piro-7) in Piroline; int-e.4 (trans) in Bonus (PI 189763), -e.65 in Bonus; int-e.72, -e.87 in Bonus, -e.89, -e.90, -e.91, -e.92, -e.101 in Hege. Mutant used for description and seed stocks: vrs4.k in Gamma III 3B (GSHO 775); int-e.58 in Kristina (GSHO 1766); mul1.a in Bowman (PI 483237)*6 (GSHO 1985); vrs4.k in Bowman*6 (GSHO 1986); int-e.58 in Bowman*6 (GSHO 1987).1513925GSHO 775
410GSHO 776vrs5.nHordeum vulgare L. subsp. vulgare Colorado, United States SPIKEHistoric1994DONATED05/1994Genetic materialAlleles at the int-c (vrs5) locus alter the size of lateral spikelets. The lemma apex of lateral kernels is rounded or weakly pointed, awnless or short-awned. Lower lateral spikelets may develop poorly in some int-c mutants, while seed development may occur in all lateral spikelets of others. Variability in lateral spikelet development among the int-c mutants is responsive, also, to environmental conditions. The Int-c.a (formally l) allele in six-rowed barley increases the size of lateral spikelets, while the int-c.b (formally i) allele in two-rowed barley prevents anther development in lateral spikelets. The int-c.5 mutant n Bonus produces fertile stamens in lateral spikelets. in the presence of the Int-c.h (formally lh) allele of Mortoni, lateral spikelets are male fertile and may occasionally set seed. Spikes of vrs5.n plants appear similar to the six-rowed spike, but lateral spikelets are smaller (less than half the size of the central spikelets) and broader. Inheritance: Monofactorial recessive. Located in chromosome 4HS, about 13.1 cM proximal from the fch9 (chlorina seedling 9) locus, and about 14.5 cM distal from the Kap (hooded lemma) locus. Previous nomenclature and gene symbolization: Intensifier for Z=W. Infertile intermedium=i. Allelic series lh, l, i. Intermedium spike-c=int-c. Six-rowed spike 5=v5. Mutational events: int-c.b in two-rowed barley; Int-c.h (lh) in Mortoni (Ciho 2210, B6-12); vrs5.n (v5) in Gamma4(38X-197, OUM338); int-c.5 in Bonus (PI 189763); int-c.7, -c.62, 0c.63 in Bonus, -c.13, -c.15, -c.16, -c.18, -c.25, -c.29 in Foma (CIho 11333), -c.33, -c.38, -c.45, -c.48, -c.49, -c.53, -c.56, -c.60 in Kristina; int-c.70, -c.76, -c.78, -c.84 in Bonus, -c.95 in Hege. Mutant used for description and seed stocks: vrs5.n in Gamma 4 (GSHO 776); int-c.b in Hordeum distichon var niginudum (GSHO 988); int-c.5 in Bonus (GSHO 1765); ..(GSHO 2003). Ref.: Forster, B.P. 1993. Coordinator's report: Chromosome 4. BGN 22:75-77.1513728GSHO 776
411GSHO 833lnt1.aHordeum vulgare L. subsp. vulgare JapanNSGCKERNELS1994DEVELOPEDGenetic materialThe tiller number is reduced to 2 to 4 per plant. These tillers are formed soon after seedling emergence; hence, no late-emerging tillers are observed. Culms are thick and stiff and leaves are dark green. Spike malformations occur in most environments. The spike may have irregular rachis internode lengths and is relatively short. The lower portion of the spike appears more dense. Lateral spikelets in two-rowed cultivars are enlarged and have a pointed apex. Plants homozygous for a recessive allele at the lnt1 locus head slightly earlier than normal sibs. Inheritance: Monofactorial recessive. Located in chromosome 3HL, about 30.5 cM distal from the uzu (uzu) locus. Previous nomenclature and gene symbolization: Reduced number of tillers=rnt. Intermedium spike-l=int-l. Mutational events: lnt1.a in Mitake (OUJ408); int-l.81 in Bonus (PI 189763). Mutant used for description and seed stocks: lnt1.a in Mitake (GSHO 833); int-l.81 in Bonus (GSHO 1771); lnt1.a in Bowman (PI 483237) *8 (GSHO 1984); int-l.81 in Bowman*6 (GSHO 1961).1514004GSHO 833
412GSHO 835Lga1.aHordeum vulgare L. subsp. vulgare Alberta, Canada KERNELSHistoric1994DEVELOPEDGenetic materialIn spikelets of plants with the dominant allele (Lga1.a), the glume (outer) plus its awn is much longer than the kernel. Heterozygotes have a glume awn of intermediate length. In the short glume awn phenotype (lga1.b), the glume length and width are unaltered, but the glume awn is 4 to 6 mm for the short type and 11 to 13 mm for the long type. Recessive alleles at the eog locus (GSHO 1592) produce an array of glume sizes, some of which have glumes phenotypically similar to those associated with the dominant allele at the Lga locus. However, all eog alleles have wider than normal glumes and show a recessive inheritance pattern. Inheritance: Monofactorial dominant. Location is unknown. Previous nomenclature and gene symbolization: Long glume awn=L. Long outer glume=Log; Helgason used the gene symbol log for the short glume awn type, but this symbol is also used for the elongated outer glume (eog) gene. Mutational events: Lga1.a in Guy Mayle (CIho 620; lga1.b in Canadina Thorpe (PI 2710). Mutant used for description and seed stocks: Lga1.a in Guy Mayle (GSHO 835); lga1.b in Canadian Thorpe (GSHO 834); lga1.b in Bowman (PI 483237) from Hector (CIho 15514); Lga1.a from Akashinriki (OUJ659, PI 467400) mutant OUM113 in Bowman (PI 483237)* 8 (GSHO 2147).1514458GSHO 835
413GSHO 842btr2.bHordeum vulgare L. subsp. vulgare JapanNSGCKERNELS1994DEVELOPEDGenetic materialJoints at the rachis nodes are tough or non-britle, which prevents segmentation or disarticulation of the rachis prior to maturity. Plants homozygous for the btr2.b allele do not shatter prior to harvest in contrast to wild barley (Hordeum vulgare subsp spontaneum). All cultivars with the btr2.b allele have a dominant allele at the Btr1 locus. When crossed to cultivars with the btr1.a allele at the Btr1 locus (genotype btr1btr1Btr2Btr2), F1 plants have a brittle rachis. The segregation ratio in the F2 generation is 1 brittle: 1 tough rachis. The majority of the cultivars (mostly six-rowed) distributed in Oriental region of the Old World have the btr2.b allele. Inheritance: Monofactorial recessive in crosses to Hordeum vulgare subsp spontaneum. Located in chromosome 3HS, very close or semi-allelic to the Btr1 (brittle rachis 1) locus; the Btr1Btr2 complex is about 15.4 cM distal from the alm (albino lemma) locus. Previous nomenclature and gene symbolization: Non-brittle=r. Non-brittle rachis=b. Non-brittle rachis 2=bt2. Mutational events: btr2.b in an unknown race of Hordeum vulgare subsp spontaneum. Mutant used for description and seed stocks: btr2.b in Sakigake (GSHO 842); btr2.b in Bowman (PI 483237); Btr1Btr2 from R.I. Wolfe's Multiple Dominant Marker Stock in Bowman*10 (GSHO 1937).1513875GSHO 842
414GSHO 851fch3.dHordeum vulgare L. subsp. vulgare Minnesota, United StatesNSGCKERNELS1994DEVELOPEDGenetic materialSeedlings have bright yellow leaves, which gradually change color to a light green as the plant grows. Plants are vigorous, but remain chlorina or light green until maturity. Plants expressing the fch3.d gene can be distinguished readily from normal plants at all stages of growth. Inheritance: Monofactorial recessive. Located in chromosome 1HS [5S]. Previous nomenclature and gene symbolization: Chlorina seedling 3=f3. Mutational events: fch3.d in an unknown cultivar (Minn 89-4). Mutant used for description and seed stocks: fch3.d in an unknown cultivar (GSHO 851); fch3.d in Bowman (PI 483237)*6 (GSHO 2033).1513938GSHO 851
415GSHO 93zeb2.bHordeum vulgare L. subsp. vulgare Minnesota, United StatesNSGCKERNELS1994DEVELOPEDGenetic materialLeaves are chlorina throughout the life cycle, independent of light and temperature. Plants may be killed by low temperature during the winter when normal plants are unaffected. A zonate pattern of different degrees of chlorina is observed in seedling leaves. Plants are weak or non-viable, dependent on level of environmental stress. After much confusion about this gene, the locus was named zebra stripe 2=zb2. Inheritance: Monofactorial recessive. Located in chromosome 4HL, about 17.0 cM proximal from the glf3 (glossy leaf 3) locus. Previous nomenclature and gene symbolization: Zoned leaf=zd. Zoned leaf=z. Zoned leaf c-zbc. Zebra leaf or zoned Colorado=zc. Zebra striped leaves=zd. Zebra stripe 2=zb2. Chlorina seedling 10=f10. Mutational events: zeb2.b in an unknown cultivar; fch10.s in an unknown cultivar (GSHO 1234); both stocks may have originated from the same mutant used by Woodward and first isolated by Immer and Henderson. Mutant used for description and seed stocks: zeb2.b in an unknown cultivar (GSHO 93); zeb2.b in Bowman (PI 483237)*4 (GSHO 2012).1510745GSHO 93
416GSHO 968Hsh1.aHordeum vulgare L. subsp. vulgare JapanNSGCKERNELS1994DEVELOPEDGenetic materialShort hairs (1 to 3 mm) are scattered or in rows on leaf sheaths of the basal part of the plant. The density of hairs varies considerably among cultivars and with changes in growing conditions. With few exceptions, no hairs are observed on the sheath of upper leaves. Heterozygotes and smooth awned cultivars seem to have fewer hairs. Inheritance: Monofactorial dominant. Located in chromosome 4HL, over 8.7 cM proximal from the yhd1 (yellow head 1) locus, and over 22.5 cM distal from the mlo (reaction to Erysiphe graminis hordei-o) locus. Previous nomenclature and gene symbolization: Hairy leaf sheath=Hs. Mutational events: Hsh1.a introduced into cultivated barley from its wild progenitor. Mutant used for description and seed stocks: Hsh1.a in Kimugi (OUL012, GSHO 968); Hsh1.a from Wolfe's Multiple Dominant Marker Stock in Bowman (PI 483237)*10 (GSHO 2026).1514400GSHO 968
417GSHO 98glf1.aHordeum vulgare L. subsp. vulgare Colorado, United StatesNSGCKERNELS1994DEVELOPEDGenetic materialSurface wax coating on the leaf blade appears to be absent from the seedling stage to near maturity, and leaves have a shiny appearance (wax code ++ ++ -). Plants are semidwarf, relatively weak, and late in heading. The stock in the Bonus is highly sterile, but the Bowman backcross-derived line has nearly complete fertility. Inheritance: Monofactorial recessive. Located in chromosome 4HL, about 7.5 cM distal from the lbi2 (long basal rachis internode 2) locus, and about 4.8 cM distal from the Mlg (reg2, reaction to Erysiphe graminis) locus. Previous nomenclature and gene symbolization: Waxless bloom on leaves=w1. Glossy=gl. Glossy leaves=gl. Glossy leaf=gl. Glossy seedling 2=gl2. Eceriferum-zh=cer-zh. Mutational events: glf1.a, glf1.b (gl2, GSHO 22) in Himalaya; glf1.f in 34-119-1, glf1.g in II-34-199-7-2 (GSHO 89), cer-zh.54 in Bonus; cer-zh.266, -zh.308, -zh.357, -zh.366, -zh.432, -zh.433 in Foma (CIho 11333); cer-zh.325 in Foma; cer-zh.373 in Foma; cer-zh.865 in Bonus. Mutant used for description and seed stocks: glf1.a in Himalaya (GSHO 98); cer-zh.54 in Bonus (GSHO 455) is used for allelism tests; glf1.a in Bowman (PI 483237)*8 (GSHO 2015).1510793GSHO 98
418GSHO 592des1.aHordeum vulgare L. subsp. vulgare Minnesota, United StatesNSGCKERNELS1994DEVELOPEDGenetic materialThe name long chromosomes refers to the frequent occurrence of rod bivalents at metaphase I. The chromsomes are paired during pachytene, but undergo desynapsis during diplotene. The degree of desynapsis is d=2.6+1.6 ranging from 7 ring bivalents (d=9) to 5 rod bivalents plus 4 univalents (d=9). Lagging chromosomes and micronuclei are observed frequently at Telophase I. Microspore tetrads contain an average of 0.3 micronuclei per tetrad with a range of 0 to 6. Ovule fertility is about 45%. Inheritance: Monofactorial recessive. Located in chromosome 7H[1]. Previous nomenclature and gene symbolization: Long chromosome=lc. Mutational events: des1.a in Mars; des1.v in Freja (CIho 7130). Mutant used for description and seed stocks: des1.a in Mars (GSHO 592).1512385GSHO 592
419GSHO 593des2.bHordeum vulgare L. subsp. vulgare Manitoba, CanadaNSGCKERNELS1994DEVELOPEDGenetic materialSynapsed homologous chromosome disjoin precociously because of an apparent lack of chiasmata. The chromosomes are paired during pachytene and undergo desynapsis during diplotene. The degree of desynapsis is d=6.0+3.3 with a range from 7 ring bivalents (d=0) to 2 rod bivalents plus 10 univalents (d=12). Lagging chromosomes and micronuclei are observed frequently at Telophase I. Microspore tetrads contain an average of 1.8 micronuclei per tetrad and a range of 0 to 7. Ovule fertility is about 1% and highly variable from spike to spike on the same plant. The very low fertility trait occurred infrequently in the F2 progeny from the cross to Bowman, but about 1/4 the plants had about 60% seed set. Inheritance: Monofactorial recessive. Located in chromosome 3H. Previous nomenclature and gene symbolization: Desynaptic chromosomes=ds. Mutational events: des2.b in Husky. Mutant used for description and seed stocks: des2.b in Husky (GSHO 593).1512429GSHO 593
420GSHO 594des3.cHordeum vulgare L. subsp. vulgare Arizona, United StatesNSGCKERNELS1994DEVELOPEDGenetic materialThe chromosome are paired during the pachyene stage of microsporogenesis, but they undergo desynapsis during diplotene and diakinesis. The degree of desynapsis is 0.9 + 1.7, ranging from 7 ring bivalents (d=0) to 4 ring bivalents plus 2 rod bivalents and 2 univalents (d=4). Ovule fertility ranges from 1% to 33%, much too low to be explained by the low degree of desynapsis observed in microsporocytes. Inheritance: Monofactorial recessive. Location is unknown. Previous nomenclature and gene symbolization: None. Mutational events: des3.c in Betzes. Mutant used for description and seed stocks: des3.c in Betzes (GSHO 594).1512194GSHO 594
421GSHO 595des4.dHordeum vulgare L. subsp. vulgare Arizona, United StatesNSGCKERNELS1994DEVELOPEDGenetic materialThe chromosome are paired during pachytene and undergo desynapsis during diplotene. The degree of desynapsis is d=3.3+2.2 with a range from 7 ring bivalents (d=0) to 3 rod bivalents plus 8 univalents (d=11). Many of the univalents split l longitudinally during Anaphase I, and lagging chromosomes and micronuclei are observed frequently at Telophase I. Mircrospore tetrads contain an average of 1.0 micronuclei per tetrad and a range of 0 to 10. Ovule fertility is about 18%. Inheritance: Monofactorial recessive. Located in chromosome 7H[1]. Previous nomenclature and gene symbolization: None. Mutational events: des4.d, des4.h in Betzes; des4.z, des4.aa, des4.ab, des4.ac, des4.ad, des4.ae, des4.af, des4.ag in Klages (CIho 15487); all the Klages mutants may be identical because they were isolated from the same field. Mutant used for description and seed stock: des4.d in Betzes (GSHO 595).1512410GSHO 595
422GSHO 596des5.eHordeum vulgare L. subsp. vulgare Arizona, United StatesNSGCKERNELSNot Available1994DEVELOPEDGenetic materialThe chromosomes are paired during pachytene and undergo desynapsis during diplotene. The degree of desynapsis is 11.1+2.6 ranging from 7 ring bivalents (d=0) to 14 univalents (d=14). Many univalents split longitudinally during Anaphase I. Lagging chromosomes and micronuclei are observed frequently at Telophase I. Microspore quartets contain an average of 4.5 micronuclei per quartet with a range of 0 to 18. Ovule fertility is about 7%. When crossed with non-allelic desynaptic lines, the F1's frequently show a low degree of desynapsis (up to 3 rod bivalents per cell). Inheritance: Monofactorial recessive. Located in chromosome 7H [1]. Previous nomenclature and gene symbolization: None. Mutational events: des5.e, des5.f, des5.g in Betzes. Mutant used for description and seed stocks: des5.e in Betzes (GSHO 596).1512195GSHO 596
423GSHO 599des8.kHordeum vulgare L. subsp. vulgare Arizona, United StatesNSGCKERNELS1994DEVELOPEDGenetic materialThe chromosomes are paired during pachytene and undergo desynapsis during diplotene. The degree of desynapsis is 7.9 + 2.3, ranging from 7 ring bivalents (d=0) to 14 univalents (d=14). Many univalents split longitudinally during anaphase I. Lagging chromosomes and micronuclei are frequently observed at telophase I. Microspore quartets contain an average of 1.2 micronuclei per quartet with a range of 0 to 10. Ovule fertility is about 43% in the des8.k stock, but about 1% in the des8.l stock. Inheritance: Monofactorial recessive. Location is unknown. Previous nomenclature and gene symbolization: None. Mutational events: des8.k, des8.l in Betzes. Mutant used for description and seed stocks: des8.k in Betzes (GSHO 599).1512119GSHO 599
424GSHO 600des9.nHordeum vulgare L. subsp. vulgare Arizona, United StatesNSGCKERNELS1994DEVELOPEDGenetic materialThe chromsomes are paired during pachytene and undergo desynapsis during diplotene and diakinesis. The average degree of desynapsis is 0.9, ranging from 7 ring bivalents (d=0) to 5 rod bivalents (d=5). Ovule fertility is about 90%. Inheritance: Monofactorial recessive. Location is unknown. Previous nomenclature and gene symbolization: Desynapsis n=des,,n. Mutational events: des9.n in Betzes. Mutant used for description and seed stocks: des9.n in Betzes (GSHO 600).1512322GSHO 600
425GSHO 601des10.pHordeum vulgare L. subsp. vulgare Arizona, United StatesNSGCKERNELS1994DEVELOPEDGenetic materialThe chromosomes are paired during pachytene and undergo desynapsis during diplotene and diakinesis. The average degree of desynapsis is 2.0, ranging from 7 ring bivalents (d=0) to 3 ring bivalents plus 2 rod bivalents plus 4 univalents (d=6). Ovule fertility is 60 to 80%. Inheritance: Monofactorial recessive. Location is unknown. Previous nomenclature and gene symbolization: Desynapsis p=des,,p. Mutational events: des10.p in Betzes. Mutant used for description and seed stocks: des10.p in Betzes (GSHO 601).1512282GSHO 601
426GSHO 602des11.rHordeum vulgare L. subsp. vulgare Arizona, United StatesNSGCKERNELS1994DEVELOPEDGenetic materialThe chromosome are paired during pachytene and undergo desynapsis during diplotene and diakinesis. The average degree of desynapsis is 5.7, ranging from 6 ring bivalents plus one rod bivalent (d=1) to 2 rod bivalents plus 10 univalents (d=12). Ovule fertility is about 40%. When crossed with non-allelic desynaptic mutants des4.d and des5.e, the F1's frequently show a low degree of desynapsis (up to 3 rod bivalents). When crossed with des7.j, the F1's show a low to medium degree of desynapsis (d=1.8), ranging from 7 ring bivalents (d=0) to 4 ring bivalents plus 4 rod bivalents (d=5), and there are lagging chromosomes and micronuclei at Telophase I. Inheritance: Monofactorial recessive. Location is unknown. Previous nomenclature and gene symbolization: Desynapsis r=des,,r. Mutational events: des11.r in Betzes. Mutant used for description and seed stocks: des11.r in Betzes (GSHO 602).1512196GSHO 602
427GSHO 603des12.sHordeum vulgare L. subsp. vulgare Arizona, United StatesNSGCKERNELS1994DEVELOPEDGenetic materialThe chromosomes are paired during pachytene and undergo desynapsis during diplotene and diakinesis. The average degree of desynapsis is 4.7, ranging from 7 ring bivalents (d=0) to 1 rod bivalent plus 12 univalents (d=13). Ovule fertility is about 35%. Inheritance: Monofactorial recessive. Location is unknown. Previous nomenclature and gene symbolization: Desynapsis s=des,,s. Mutational events: des12.s, des12.w in Betzes. Mutant used for description and seed stocks: des12.s in Betzes (GSHO 603).1512226GSHO 603
428GSHO 604des13.tHordeum vulgare L. subsp. vulgare Arizona, United StatesNSGCKERNELS1994DEVELOPEDGenetic materialThere are two types of microspore mother cells in mutant plants. About 58% have normal early prophases and undergo desynapsis during diplotene and diakinesis. The average degree of desynapsis is 0.8, ranging from 7 ring bivalents (d=0) to 4 ring bivalents plus 2 rod bivalents plus 2 univalents (d=4). The othr type of microspore mother cells have chromosomes that appear "sticky" at Metaphase I. The stickiness is accompanined by very elongated ring bivalents and by extensive chromosome fragmentation. Ovule fertility is about 20 to 30%. Inheritance: Monofactorial recessive. Location is unknown. Previous nomenclature and gene symbolization: Desynapsis t=des,,t. Mutational events: des13.t in Betzes. Mutant used for description and seed stocks: des13.t in Betzes (GSHO 604).1512396GSHO 604
429GSHO 605des14.uHordeum vulgare L. subsp. vulgare Arizona, United StatesNSGCKERNELS1994DEVELOPEDGenetic materialThe chromosomes are paired during pachytene and undergo desynapsis during diplotene and diakinesis. The average degree of desynapsis is 4.8, ranging frm 7 ring bivalents (d=0) to 1 rod bivalent plus 12 univalents (d=13). Lagging chromosomes and microsnuclei are observed frequently at Telophase I. Ovule fertility is about 35%. Inheritance: Monofactorial recessive. Location is unknown. Previous nomenclature and gene symbolization: Desynapsis u=des,,u. Mutational events: des14.u in Betzes. Mutant used for description and seed stocks: des14.u in Betzes (GSHO 605).1512456GSHO 605
430GSHO 606des15.xHordeum vulgare L. subsp. vulgare Arizona, United StatesNSGCKERNELS1994DEVELOPEDGenetic materialThe chromosomes are paired during pachytene and undergo desynapsis during diplotene and diakinesis. The average degree of desynapsis is 7.5, ranging from 5 ring bivalents plus 2 rod bivalents (d=2) to 1 rod bivalent plus 12 univalents (d=13). Ovule fertility is about 25%. Inheritance: Monofactorial recessive. Location is unknown. Previous nomenclature and gene symbolization: Desynapsis x=des,,x. Mutational events: des15.x, des15.y in Ingrid. Mutant used for description and seed stocks: des15.x in Ingrid (GSHO 606).1512334GSHO 606
431GSHO 2538ferHordeum vulgare L. subsp. vulgare Colorado, United States Historic1990DONATED12/1990Genetic materialFew roots of seedlings and of plants grown in gravel. Older plants are phenotypically normal, but inclined to overturn. See Barley Genetics Newsletter 8:162. 1006454GSHO 2538