Taxon: Raphanus raphanistrum L.

Summary

Genus:

Raphanus

Family:

Brassicaceae (alt. Cruciferae)

Tribe:

Brassiceae

Nomen number:

30856

Place of publication:

Sp. pl. 2:669. 1753

Protologue link:

https://biodiversitylibrary.org/page/358690

Typification:

View in Linnean Typification Project

Verified:

08/06/2008 ARS Systematic Botanists.

Accessions:

11 (4 active, 4 available) in National Plant Germplasm System. (Map it)

Other conspecific taxa

Raphanus raphanistrum L. subsp. landra (Moretti ex DC.) Bonnier & Layens (1 active accession[s])
Raphanus raphanistrum L. subsp. raphanistrum (0 active accession[s])
Raphanus raphanistrum L. subsp. rostratus (DC.) Thell. (0 active accession[s])

Autonyms (not in current use), synonyms and invalid designations

Images (1 total. Click on image for more.)

Links

Seeds or fruits – Seed ID Workshop image, from Department of Horticulture and Crop Science, Ohio State University

Conservation Status

Reference(s)

University of Georgia Center for Invasive Species and Ecosystem Health
Acevedo-Rodríguez, P. & M. T. Strong. 2012. Catalogue of seed plants of the West Indies. Smithsonian Contr. Bot. 98
Akhani, H. 2003. Notes on the flora of Iran: 4. Two new records and synopsis of the new data on Iranian Cruciferae since Flora Iranica. Candollea 58:379.
Aldén, B., S. Ryman, & M. Hjertson. 2012. Svensk Kulturväxtdatabas, SKUD (Swedish Cultivated and Utility Plants Database; online resource)
Ali, S. I. & S. M. H. Jafri, eds. 1976-. Flora of Libya.
Allan, H. H. B. et al. 1961-. Flora of New Zealand.
APHIS. Agricultural Risk Management System United States Department of Agriculture.
Arechavaleta, M. et al., coords. 2009. Lista de especies silvestres de Canarias: hongos, plantas y animales terrestres
Bosser, J. M. et al., eds. 1976-. Flores des Mascareignes.
Brako, L. & J. L. Zarucchi. 1993. Catalogue of the flowering plants and gymnosperms of Peru. Monogr. Syst. Bot. Missouri Bot. Gard. 45
Chinese Academy of Sciences. 1959-. Flora reipublicae popularis sinicae.
CIBA-GEIGY, Basel, Switzerland. Documenta CIBA-GEIGY (Grass weeds 1. 1980, 2. 1981; Monocot weeds 3. 1982; Dicot weeds 1. 1988) Note: four books on weeds worldwide in scope
Cufodontis, G. 1953-1972. Enumeratio plantarum aethiopiae: Spermatophyta.
Darbyshire, S. J. 2003. Inventory of Canadian Agricultural Weeds Agriculture and Agri-Food Canada. 118.
Davis, P. H., ed. 1965-1988. Flora of Turkey and the east Aegean islands.
Duke, J. A. et al. 2002. CRC Handbook of medicinal herbs
Erhardt, W. et al. 2008. Der große Zander: Enzyklopädie der Pflanzennamen
Eriksson, O. et al. 1979. Flora of Macaronesia: checklist of vascular plants, ed. 2.
Exell, A. W. et al., eds. 1960-. Flora zambesiaca.
FitzJohn, R. G. et al. 2007. Hybridisation within Brassica and allied genera: evaluation of potential for transgene escape. Euphytica 158:209-230. Note: Netherlands journal of plant breeding
FNA Editorial Committee. 1993-. Flora of North America.
Forzza, R. C. et al., coord. Lista de espécies da flora do Brasil (on-line resource).
George, A. S., ed. 1980-. Flora of Australia.
Germishuizen, G. & N. L. Meyer, eds. 2003. Plants of southern Africa: an annotated checklist. Strelitzia 14 National Botanical Institute, Pretoria, South Africa.
Gleason, H. A. & A. Cronquist. 1963. Manual of vascular plants of northeastern United States and adjacent Canada.
Greuter, W. et al., eds. 1984-. Med-Checklist.
Grierson, A. J. C. & D. J. Long. 1984-. Flora of Bhutan including a record of plants from Sikkim.
Groth, D. 2005. pers. comm. Note: re. Brazilian common names
Hedberg, I. & S. Edwards. 1989-. Flora of Ethiopia Note: Eritrea added in 2000
Hegi, G. et al. 1906-. Illustrierte Flora von Mittel-Europa Note: ed. 1:1906-1931; ed. 2:1936-68; ed. 3:1966-
Holm, L. et al. 1979. A geographical atlas of world weeds
Instituto de Botánica Darwinion. 2008. Flora del Conosur. Catálogo de las plantas vasculares.
Integrated Botanical Information System (IBIS). Australian plant common name database (on-line resource).
Iwatsuki, K. et al. 1993-. Flora of Japan.
Jalas, J. & J. Suominen. 1972-. Atlas florae europaeae.
Jørgensen, P. M. & S. León-Yánez, eds. 1999. Catalogue of the vascular plants of Ecuador. Monogr. Syst. Bot. Missouri Bot. Gard. 75
Kartesz, J. T. 1994. A synonymized checklist of the vascular flora of the United States, Canada, and Greenland.
Kellerman, T. S. et al. 1988. Plant poisonings and mycotoxicoses of livestock in Southern Africa Note: poisonous
Kingsbury, J. M. 1964. Poisonous plants of the United States and Canada. Note: poisonous
Koelling, V. A. & K. Karoly. 2007. Self-pollen interference is absent in wild radish (Raphanus raphanistrum, Brassicaceae), a species with sporophytic self-incompatibility. Amer. J. Bot. 94:896-900.
Komarov, V. L. et al., eds. 1934-1964. Flora SSSR.
Krasnoborov, I. M., ed. 2000-. Flora of Siberia (English translation)
Kunakh, V. A. et al. 2008. Mixoploidy in wild and cultivated species of Cruciferae capable of hybridizing with rapeseed Brassica napus. Cytol. & Genet. 42:204-209. Note: Translation of: Tsitologiya i genetika. Kiev
Lefol, E. et al. 1997. Sexual hybridisation in crosses of cultivated Brassica species with the crucifers Erucastrum gallicum and Raphanus raphanistrum: potential for gene introgression. Euphytica 95:127-139. Note: Netherlands journal of plant breeding
Miller, A. G. & T. A. Cope. 1996-. Flora of the Arabian Peninsula and Socotra.
Mouterde, P. 1966-. Nouvelle flore du Liban et de la Syrie.
Munro, D. B. Canadian poisonous plants information system (on-line resource).
Nasir, E. & S. I. Ali, eds. 1970-. Flora of [West] Pakistan.
Parsons, W. T. & E. G. Cuthbertson. 1992. Noxious weeds of Australia Inkata Press, Melbourne, Australia.
Pistrick, K. 1987. Untersuchungen zur Systematik der Gattung Raphanus L. Kulturpflanze 35:225-321.
Porcher, M. H. et al. Searchable World Wide Web Multilingual Multiscript Plant Name Database (MMPND) (on-line resource).
Rechinger, K. H., ed. 1963-. Flora iranica.
Rehm, S. 1994. Multilingual dictionary of agronomic plants
Sahli, H. F. et al. 2008. Adaptive differentiation of quantitative traits in the globally distributed weed, wild radish (Raphanus raphanistrum). Genetics 180:945-955.
Seed Regulatory and Testing Division, Agricultural Marketing Service, U.S.D.A. State noxious-weed seed requirements recognized in the administration of the Federal Seed Act (updated annually).
Sharma, B. D. et al., eds. 1993-. Flora of India.
Silva, L. et al. 2005. Listagem da fauna e flora terrestres dos Açores. Lista des plantas vasculares (Pteridophyta e Spermatophyta).
Snow, A. A. et al. 2001. Fitness of hybrids betweem weedy and cultivated radish: implications for weed evolution. Ecol. Applic. 11:934-939.
Standley, P. C. & J. A. Steyermark. 1946-1976. Flora of Guatemala.
Stevens, W. D. et al., eds. 2001. Flora de Nicaragua. Monogr. Syst. Bot. Missouri Bot. Gard. 85(1-3).
Terrell, E. E. et al. 1986. Agricultural Handbook no. 505
Townsend, C. C. & E. Guest. 1966-. Flora of Iraq.
Turrill, W. B. et al., eds. 1952-. Flora of tropical East Africa.
Tutin, T. G. et al., eds. 1993. Flora europaea, ed. 2.
USDA/ARS. INVADERS database system. The University of Montana (on-line resource).
Vibrans, H., ed. Malezas de México (on-line resource).
Wagner, W. L. et al. 1990. Manual of the flowering plants of Hawai'i
Warwick, S. I. & A. Francis. 2005. The biology of Canadian weeds. 132. Raphanus raphanistrum L. Canad. J. Pl. Sci. 85:709-733.
Wu Zheng-yi & P. H. Raven et al., eds. 1994-. Flora of China (English edition).
Yamagishi, H. & T. Terachi. 2003. Multiple origins of cultivated radishes as evidenced by a comparison of the structural variations in mitochondrial DNA of Raphanus. Genome 46:89-94.
Yamagishi, H. et al. 2009. Inter- and intraspecific sequence variations of the chloroplast genome in wild and cultivated Raphanus. Pl. Breed. (New York) 128:172-177.
Yamame, K. et al. 2005. Chloroplast DNA variations of cultivated radish and its wild relatives. Pl. Sci. (Elsevier) 168:627-634.
Yamane, K. et al. 2009. Multiple origins and high genetic diversity of cultivated radish inferred from polymorphism in chloroplast simple sequence repeats. Breed. Sci. 59:55-65.
Zohary, M. & N. Feinbrun-Dothan. 1966-. Flora palaestina.
2018. Mid-Atlantic invasive plant species (on-line resource)

Common names

Language	Name	note	seq	Citation
English	jointed charlock		1	FNA Editorial Committee. 1993-. Flora of North America.
English	jointed radish		1	FNA Editorial Committee. 1993-. Flora of North America.
English	jointed wild radish		1	FNA Editorial Committee. 1993-. Flora of North America.
English	sea radish		1	Erhardt, W. et al. 2008. Der große Zander: Enzyklopädie der Pflanzennamen
English	white charlock		1	Integrated Botanical Information System (IBIS). Australian plant common name database (on-line resource).
English	wild radish		1	USDA/ARS. INVADERS database system. The University of Montana (on-line resource).
English (Canada)	wild radish	Agriculture & Agri-Food Canada official name	2	Darbyshire, S. J. 2003. Inventory of Canadian Agricultural Weeds Agriculture and Agri-Food Canada. 118.
French	ravenelle		2	Rehm, S. 1994. Multilingual dictionary of agronomic plants
French (Canada)	radis sauvage	Agriculture & Agroalimentaire Canada nom officiel	2	Darbyshire, S. J. 2003. Inventory of Canadian Agricultural Weeds Agriculture and Agri-Food Canada. 118.
German	Hederich		2	Rehm, S. 1994. Multilingual dictionary of agronomic plants
Portuguese (Brazil)	nabiça		2	Groth, D. 2005. pers. comm. Note: re. Brazilian common names
Portuguese (Brazil)	nabo		2	Groth, D. 2005. pers. comm. Note: re. Brazilian common names
Portuguese (Brazil)	rabanete-de-cavalo		2	Groth, D. 2005. pers. comm. Note: re. Brazilian common names
Portuguese (Brazil)	rabanete-selvagem		2	Groth, D. 2005. pers. comm. Note: re. Brazilian common names
Portuguese (Brazil)	rabanete-silvestre		2	Groth, D. 2005. pers. comm. Note: re. Brazilian common names
Spanish	rabanillo blanco		2	Rehm, S. 1994. Multilingual dictionary of agronomic plants
Spanish	rabaniza		2	Rehm, S. 1994. Multilingual dictionary of agronomic plants
Spanish	rábano silvestre		2	Rehm, S. 1994. Multilingual dictionary of agronomic plants
Swedish	åkerrättika		2	Aldén, B., S. Ryman, & M. Hjertson. 2012. Svensk Kulturväxtdatabas, SKUD (Swedish Cultivated and Utility Plants Database; online resource)

Distribution

order_code	Status	Continent	Subcontinent	Country	State	Note	Reference
1	Native	Africa	Macaronesia	Portugal	Madeira Islands
1	Native	Africa	Macaronesia	Spain	Canarias		Arechavaleta, M. et al., coords. 2009. Lista de especies silvestres de Canarias: hongos, plantas y animales terrestres
1	Native	Africa	Northern Africa	Algeria		n.	Greuter, W. et al., eds. 1984-. Med-Checklist.
1	Native	Africa	Northern Africa	Egypt		n.	Greuter, W. et al., eds. 1984-. Med-Checklist.
1	Native	Africa	Northern Africa	Libya		n.	Greuter, W. et al., eds. 1984-. Med-Checklist.
1	Native	Africa	Northern Africa	Morocco			Greuter, W. et al., eds. 1984-. Med-Checklist.
1	Native	Africa	Northern Africa	Tunisia			Greuter, W. et al., eds. 1984-. Med-Checklist.
1	Native	Asia-Temperate	Caucasus	Armenia			Komarov, V. L. et al., eds. 1934-1964. Flora SSSR.
1	Native	Asia-Temperate	Caucasus	Azerbaijan			Komarov, V. L. et al., eds. 1934-1964. Flora SSSR.
1	Native	Asia-Temperate	Caucasus	Georgia			Komarov, V. L. et al., eds. 1934-1964. Flora SSSR.
1	Native	Asia-Temperate	Caucasus	Russian Federation-Ciscaucasia	Ciscaucasia		Komarov, V. L. et al., eds. 1934-1964. Flora SSSR.
1	Native	Asia-Temperate	Western Asia	Cyprus			Greuter, W. et al., eds. 1984-. Med-Checklist.
1	Native	Asia-Temperate	Western Asia	Iran			Rechinger, K. H., ed. 1963-. Flora iranica.
1	Native	Asia-Temperate	Western Asia	Iraq			Townsend, C. C. & E. Guest. 1966-. Flora of Iraq.
1	Native	Asia-Temperate	Western Asia	Israel			Zohary, M. & N. Feinbrun-Dothan. 1966-. Flora palaestina.
1	Native	Asia-Temperate	Western Asia	Jordan			Zohary, M. & N. Feinbrun-Dothan. 1966-. Flora palaestina.
1	Native	Asia-Temperate	Western Asia	Lebanon			Mouterde, P. 1966-. Nouvelle flore du Liban et de la Syrie.
1	Native	Asia-Temperate	Western Asia	Syria			Mouterde, P. 1966-. Nouvelle flore du Liban et de la Syrie.
1	Native	Asia-Temperate	Western Asia	Turkey			Davis, P. H., ed. 1965-1988. Flora of Turkey and the east Aegean islands.
1	Native	Europe	Eastern Europe	Belarus			Jalas, J. & J. Suominen. 1972-. Atlas florae europaeae.
1	Native	Europe	Eastern Europe	Estonia			Jalas, J. & J. Suominen. 1972-. Atlas florae europaeae.
1	Native	Europe	Eastern Europe	Latvia			Jalas, J. & J. Suominen. 1972-. Atlas florae europaeae.
1	Native	Europe	Eastern Europe	Lithuania			Jalas, J. & J. Suominen. 1972-. Atlas florae europaeae.
1	Native	Europe	Eastern Europe	Moldova			Jalas, J. & J. Suominen. 1972-. Atlas florae europaeae.
1	Native	Europe	Eastern Europe	Russian Federation-European part	European part		Jalas, J. & J. Suominen. 1972-. Atlas florae europaeae.
1	Native	Europe	Eastern Europe	Ukraine		incl. Krym	Jalas, J. & J. Suominen. 1972-. Atlas florae europaeae.
1	Native	Europe	Middle Europe	Austria			Jalas, J. & J. Suominen. 1972-. Atlas florae europaeae.
1	Native	Europe	Middle Europe	Belgium			Jalas, J. & J. Suominen. 1972-. Atlas florae europaeae.
1	Native	Europe	Middle Europe	Czech Republic			Jalas, J. & J. Suominen. 1972-. Atlas florae europaeae.
1	Native	Europe	Middle Europe	Germany			Jalas, J. & J. Suominen. 1972-. Atlas florae europaeae.
1	Native	Europe	Middle Europe	Hungary			Jalas, J. & J. Suominen. 1972-. Atlas florae europaeae.
1	Native	Europe	Middle Europe	Netherlands			Jalas, J. & J. Suominen. 1972-. Atlas florae europaeae.
1	Native	Europe	Middle Europe	Poland			Jalas, J. & J. Suominen. 1972-. Atlas florae europaeae.
1	Native	Europe	Middle Europe	Slovakia			Jalas, J. & J. Suominen. 1972-. Atlas florae europaeae.
1	Native	Europe	Middle Europe	Switzerland			Jalas, J. & J. Suominen. 1972-. Atlas florae europaeae.
1	Native	Europe	Northern Europe	Denmark			Jalas, J. & J. Suominen. 1972-. Atlas florae europaeae.
1	Native	Europe	Northern Europe	Finland			Jalas, J. & J. Suominen. 1972-. Atlas florae europaeae.
1	Native	Europe	Northern Europe	Ireland			Jalas, J. & J. Suominen. 1972-. Atlas florae europaeae.
1	Native	Europe	Northern Europe	Norway			Jalas, J. & J. Suominen. 1972-. Atlas florae europaeae.
1	Native	Europe	Northern Europe	Sweden			Jalas, J. & J. Suominen. 1972-. Atlas florae europaeae.
1	Native	Europe	Northern Europe	United Kingdom			Jalas, J. & J. Suominen. 1972-. Atlas florae europaeae.
1	Native	Europe	Southeastern Europe	Albania			Jalas, J. & J. Suominen. 1972-. Atlas florae europaeae.
1	Native	Europe	Southeastern Europe	Bosnia and Herzegovina			Jalas, J. & J. Suominen. 1972-. Atlas florae europaeae.
1	Native	Europe	Southeastern Europe	Bulgaria			Jalas, J. & J. Suominen. 1972-. Atlas florae europaeae.
1	Native	Europe	Southeastern Europe	Croatia			Jalas, J. & J. Suominen. 1972-. Atlas florae europaeae.
1	Native	Europe	Southeastern Europe	Greece		incl. Crete	Jalas, J. & J. Suominen. 1972-. Atlas florae europaeae.
1	Native	Europe	Southeastern Europe	Italy		incl. Sardinia, Sicily	Jalas, J. & J. Suominen. 1972-. Atlas florae europaeae.
1	Native	Europe	Southeastern Europe	Montenegro			Jalas, J. & J. Suominen. 1972-. Atlas florae europaeae.
1	Native	Europe	Southeastern Europe	North Macedonia			Jalas, J. & J. Suominen. 1972-. Atlas florae europaeae.
1	Native	Europe	Southeastern Europe	Romania			Jalas, J. & J. Suominen. 1972-. Atlas florae europaeae.
1	Native	Europe	Southeastern Europe	Serbia			Jalas, J. & J. Suominen. 1972-. Atlas florae europaeae.
1	Native	Europe	Southeastern Europe	Slovenia			Jalas, J. & J. Suominen. 1972-. Atlas florae europaeae.
1	Native	Europe	Southwestern Europe	France		incl. Corsica	Jalas, J. & J. Suominen. 1972-. Atlas florae europaeae.
1	Native	Europe	Southwestern Europe	Portugal			Jalas, J. & J. Suominen. 1972-. Atlas florae europaeae.
1	Native	Europe	Southwestern Europe	Spain		incl. Baleares	Jalas, J. & J. Suominen. 1972-. Atlas florae europaeae.
4	Naturalized	Africa	East Tropical Africa	Kenya			Turrill, W. B. et al., eds. 1952-. Flora of tropical East Africa.
4	Naturalized	Africa	East Tropical Africa	Tanzania			Turrill, W. B. et al., eds. 1952-. Flora of tropical East Africa.
4	Naturalized	Africa	Macaronesia	Portugal	Azores		Silva, L. et al. 2005. Listagem da fauna e flora terrestres dos Açores. Lista des plantas vasculares (Pteridophyta e Spermatophyta).
4	Naturalized	Africa	Northeast Tropical Africa	Eritrea			Hedberg, I. & S. Edwards. 1989-. Flora of Ethiopia Note: Eritrea added in 2000
4	Naturalized	Africa	Northeast Tropical Africa	Ethiopia			Hedberg, I. & S. Edwards. 1989-. Flora of Ethiopia Note: Eritrea added in 2000
4	Naturalized	Africa	South Tropical Africa	Zimbabwe			Exell, A. W. et al., eds. 1960-. Flora zambesiaca.
4	Naturalized	Africa	Southern Africa	Lesotho			Germishuizen, G. & N. L. Meyer, eds. 2003. Plants of southern Africa: an annotated checklist. Strelitzia 14 National Botanical Institute, Pretoria, South Africa.
4	Naturalized	Africa	Southern Africa	South Africa			Germishuizen, G. & N. L. Meyer, eds. 2003. Plants of southern Africa: an annotated checklist. Strelitzia 14 National Botanical Institute, Pretoria, South Africa.
4	Naturalized	Africa	Western Indian Ocean	Mauritius			Bosser, J. M. et al., eds. 1976-. Flores des Mascareignes.
4	Naturalized	Africa	Western Indian Ocean	Reunion			Bosser, J. M. et al., eds. 1976-. Flores des Mascareignes.
4	Naturalized	Asia-Temperate	Arabian Peninsula	Bahrain			Miller, A. G. & T. A. Cope. 1996-. Flora of the Arabian Peninsula and Socotra.
4	Naturalized	Asia-Temperate	Arabian Peninsula	Oman			Miller, A. G. & T. A. Cope. 1996-. Flora of the Arabian Peninsula and Socotra.
4	Naturalized	Asia-Temperate	Arabian Peninsula	Saudi Arabia			Miller, A. G. & T. A. Cope. 1996-. Flora of the Arabian Peninsula and Socotra.
4	Naturalized	Asia-Temperate	Arabian Peninsula	Yemen			Miller, A. G. & T. A. Cope. 1996-. Flora of the Arabian Peninsula and Socotra.
4	Naturalized	Asia-Temperate	China	China	Qinghai Sheng		Wu Zheng-yi & P. H. Raven et al., eds. 1994-. Flora of China (English edition).
4	Naturalized	Asia-Temperate	China	China	Sichuan Sheng		Wu Zheng-yi & P. H. Raven et al., eds. 1994-. Flora of China (English edition).
4	Naturalized	Asia-Temperate	Eastern Asia	Japan	Hokkaidô		Iwatsuki, K. et al. 1993-. Flora of Japan.
4	Naturalized	Asia-Temperate	Eastern Asia	Japan	Honshu		Iwatsuki, K. et al. 1993-. Flora of Japan.
4	Naturalized	Asia-Temperate	Eastern Asia	Japan	Shikoku		Iwatsuki, K. et al. 1993-. Flora of Japan.
4	Naturalized	Asia-Temperate	Eastern Asia	Taiwan			Wu Zheng-yi & P. H. Raven et al., eds. 1994-. Flora of China (English edition).
4	Naturalized	Asia-Temperate	Mongolia	Mongolia			Iwatsuki, K. et al. 1993-. Flora of Japan.
4	Naturalized	Asia-Temperate	Russian Far East	Russian Federation-Far East	Far East		Krasnoborov, I. M., ed. 2000-. Flora of Siberia (English translation)
4	Naturalized	Asia-Temperate	Siberia	Russian Federation-Eastern Siberia	Eastern Siberia		Krasnoborov, I. M., ed. 2000-. Flora of Siberia (English translation)
4	Naturalized	Asia-Temperate	Siberia	Russian Federation-Western Siberia	Western Siberia		Krasnoborov, I. M., ed. 2000-. Flora of Siberia (English translation)
4	Naturalized	Asia-Tropical	Indian Subcontinent	India	Sikkim		Sharma, B. D. et al., eds. 1993-. Flora of India.
4	Naturalized	Australasia	Australia	Australia			George, A. S., ed. 1980-. Flora of Australia.
4	Naturalized	Australasia	New Zealand	New Zealand			Allan, H. H. B. et al. 1961-. Flora of New Zealand.
4	Naturalized	Northern America		Canada			FNA Editorial Committee. 1993-. Flora of North America.
4	Naturalized	Northern America		Mexico			FNA Editorial Committee. 1993-. Flora of North America.
4	Naturalized	Northern America		United States			Gleason, H. A. & A. Cronquist. 1963. Manual of vascular plants of northeastern United States and adjacent Canada.
4	Naturalized	Northern America	Eastern Canada	St. Pierre and Miquelon			FNA Editorial Committee. 1993-. Flora of North America.
4	Naturalized	Northern America	Subarctic America	Greenland			FNA Editorial Committee. 1993-. Flora of North America.
4	Naturalized	Pacific	North-Central Pacific	United States	Hawaii		Wagner, W. L. et al. 1990. Manual of the flowering plants of Hawai'i
4	Naturalized	Southern America	Brazil	Brazil			Forzza, R. C. et al., coord. Lista de espécies da flora do Brasil (on-line resource).
4	Naturalized	Southern America	Caribbean	Bahamas			Acevedo-Rodríguez, P. & M. T. Strong. 2012. Catalogue of seed plants of the West Indies. Smithsonian Contr. Bot. 98
4	Naturalized	Southern America	Caribbean	Cuba			Acevedo-Rodríguez, P. & M. T. Strong. 2012. Catalogue of seed plants of the West Indies. Smithsonian Contr. Bot. 98
4	Naturalized	Southern America	Caribbean	Hispaniola			Acevedo-Rodríguez, P. & M. T. Strong. 2012. Catalogue of seed plants of the West Indies. Smithsonian Contr. Bot. 98
4	Naturalized	Southern America	Central America	Guatemala			Standley, P. C. & J. A. Steyermark. 1946-1976. Flora of Guatemala.
4	Naturalized	Southern America	Central America	Nicaragua			Stevens, W. D. et al., eds. 2001. Flora de Nicaragua. Monogr. Syst. Bot. Missouri Bot. Gard. 85(1-3).
4	Naturalized	Southern America	Southern South America	Argentina			Instituto de Botánica Darwinion. 2008. Flora del Conosur. Catálogo de las plantas vasculares.
4	Naturalized	Southern America	Southern South America	Chile			Instituto de Botánica Darwinion. 2008. Flora del Conosur. Catálogo de las plantas vasculares.
4	Naturalized	Southern America	Southern South America	Paraguay			Instituto de Botánica Darwinion. 2008. Flora del Conosur. Catálogo de las plantas vasculares.
4	Naturalized	Southern America	Southern South America	Uruguay			Instituto de Botánica Darwinion. 2008. Flora del Conosur. Catálogo de las plantas vasculares.
4	Naturalized	Southern America	Western South America	Ecuador			Jørgensen, P. M. & S. León-Yánez, eds. 1999. Catalogue of the vascular plants of Ecuador. Monogr. Syst. Bot. Missouri Bot. Gard. 75
4	Naturalized	Southern America	Western South America	Peru			Brako, L. & J. L. Zarucchi. 1993. Catalogue of the flowering plants and gymnosperms of Peru. Monogr. Syst. Bot. Missouri Bot. Gard. 45

Native
Africa

MACARONESIA: Spain [Canarias], Portugal [Madeira Islands]
NORTHERN AFRICA: Algeria (n.), Egypt (n.), Libya (n.), Morocco, Tunisia

Asia-Temperate

WESTERN ASIA: Cyprus, Iran, Iraq, Israel, Jordan, Lebanon, Syria, Turkey
CAUCASUS: Russian Federation-Ciscaucasia [Ciscaucasia], Armenia, Azerbaijan, Georgia

Europe

NORTHERN EUROPE: Denmark, Finland, United Kingdom, Ireland, Norway, Sweden
MIDDLE EUROPE: Austria, Belgium, Switzerland, Czech Republic, Germany, Hungary, Netherlands, Poland, Slovakia
EASTERN EUROPE: Russian Federation-European part [European part], Belarus, Estonia, Lithuania, Latvia, Moldova, Ukraine (incl. Krym)
SOUTHEASTERN EUROPE: Albania, Bulgaria, Bosnia and Herzegovina, Greece (incl. Crete), Croatia, Italy (incl. Sardinia, Sicily), North Macedonia, Montenegro, Romania, Serbia, Slovenia
SOUTHWESTERN EUROPE: Spain (incl. Baleares), France (incl. Corsica), Portugal

Naturalized
Africa

MACARONESIA: Portugal [Azores]
NORTHEAST TROPICAL AFRICA: Eritrea, Ethiopia
EAST TROPICAL AFRICA: Kenya, Tanzania
SOUTH TROPICAL AFRICA: Zimbabwe
SOUTHERN AFRICA: Lesotho, South Africa
WESTERN INDIAN OCEAN: Mauritius, Reunion

Asia-Temperate

ARABIAN PENINSULA: Bahrain, Oman, Saudi Arabia, Yemen
SIBERIA: Russian Federation-Western Siberia [Western Siberia], Russian Federation-Eastern Siberia [Eastern Siberia]
MONGOLIA: Mongolia
RUSSIAN FAR EAST: Russian Federation-Far East [Far East]
CHINA: China [Qinghai Sheng, Sichuan Sheng]
EASTERN ASIA: Japan [Hokkaidô, Honshu, Shikoku], Taiwan

Asia-Tropical

INDIAN SUBCONTINENT: India [Sikkim]

Australasia

AUSTRALIA: Australia
NEW ZEALAND: New Zealand

Northern America

SUBARCTIC AMERICA: Greenland
EASTERN CANADA: St. Pierre and Miquelon
REGION: Canada, Mexico, United States

Pacific

NORTH-CENTRAL PACIFIC: United States [Hawaii]

Southern America

CARIBBEAN: Hispaniola, Bahamas, Cuba
CENTRAL AMERICA: Guatemala, Nicaragua
BRAZIL: Brazil
WESTERN SOUTH AMERICA: Ecuador, Peru
SOUTHERN SOUTH AMERICA: Argentina, Chile, Paraguay, Uruguay

Reference(s)

Acevedo-Rodríguez, P. & M. T. Strong. 2012. Catalogue of seed plants of the West Indies. Smithsonian Contr. Bot. 98
Allan, H. H. B. et al. 1961-. Flora of New Zealand.
Arechavaleta, M. et al., coords. 2009. Lista de especies silvestres de Canarias: hongos, plantas y animales terrestres
Bosser, J. M. et al., eds. 1976-. Flores des Mascareignes.
Brako, L. & J. L. Zarucchi. 1993. Catalogue of the flowering plants and gymnosperms of Peru. Monogr. Syst. Bot. Missouri Bot. Gard. 45
Davis, P. H., ed. 1965-1988. Flora of Turkey and the east Aegean islands.
Exell, A. W. et al., eds. 1960-. Flora zambesiaca.
FNA Editorial Committee. 1993-. Flora of North America.
Forzza, R. C. et al., coord. Lista de espécies da flora do Brasil (on-line resource).
George, A. S., ed. 1980-. Flora of Australia.
Germishuizen, G. & N. L. Meyer, eds. 2003. Plants of southern Africa: an annotated checklist. Strelitzia 14 National Botanical Institute, Pretoria, South Africa.
Gleason, H. A. & A. Cronquist. 1963. Manual of vascular plants of northeastern United States and adjacent Canada.
Greuter, W. et al., eds. 1984-. Med-Checklist.
Hedberg, I. & S. Edwards. 1989-. Flora of Ethiopia Note: Eritrea added in 2000
Instituto de Botánica Darwinion. 2008. Flora del Conosur. Catálogo de las plantas vasculares.
Iwatsuki, K. et al. 1993-. Flora of Japan.
Jalas, J. & J. Suominen. 1972-. Atlas florae europaeae.
Jørgensen, P. M. & S. León-Yánez, eds. 1999. Catalogue of the vascular plants of Ecuador. Monogr. Syst. Bot. Missouri Bot. Gard. 75
Komarov, V. L. et al., eds. 1934-1964. Flora SSSR.
Krasnoborov, I. M., ed. 2000-. Flora of Siberia (English translation)
Miller, A. G. & T. A. Cope. 1996-. Flora of the Arabian Peninsula and Socotra.
Mouterde, P. 1966-. Nouvelle flore du Liban et de la Syrie.
Rechinger, K. H., ed. 1963-. Flora iranica.
Sharma, B. D. et al., eds. 1993-. Flora of India.
Silva, L. et al. 2005. Listagem da fauna e flora terrestres dos Açores. Lista des plantas vasculares (Pteridophyta e Spermatophyta).
Standley, P. C. & J. A. Steyermark. 1946-1976. Flora of Guatemala.
Stevens, W. D. et al., eds. 2001. Flora de Nicaragua. Monogr. Syst. Bot. Missouri Bot. Gard. 85(1-3).
Townsend, C. C. & E. Guest. 1966-. Flora of Iraq.
Turrill, W. B. et al., eds. 1952-. Flora of tropical East Africa.
Wagner, W. L. et al. 1990. Manual of the flowering plants of Hawai'i
Wu Zheng-yi & P. H. Raven et al., eds. 1994-. Flora of China (English edition).
Zohary, M. & N. Feinbrun-Dothan. 1966-. Flora palaestina.

Economic Uses

Usage	Type	Note	Reference
Medicines	folklore		Duke, J. A. et al. 2002. CRC Handbook of medicinal herbs
Vertebrate poisons	mammals		Kingsbury, J. M. 1964. Poisonous plants of the United States and Canada. Note: poisonous
Vertebrate poisons	mammals		Kellerman, T. S. et al. 1988. Plant poisonings and mycotoxicoses of livestock in Southern Africa Note: poisonous
Weed	seed contaminant	potential	CIBA-GEIGY, Basel, Switzerland. Documenta CIBA-GEIGY (Grass weeds 1. 1980, 2. 1981; Monocot weeds 3. 1982; Dicot weeds 1. 1988) Note: four books on weeds worldwide in scope

Cite as: USDA, Agricultural Research Service, National Plant Germplasm System. 2026. Germplasm Resources Information Network (GRIN Taxonomy). National Germplasm Resources Laboratory, Beltsville, Maryland.
URL: https://npgstest.ars-grin.gov/gringlobal/taxon/taxonomydetail?id=30856. Accessed 15 April 2026.