Taxon: Veronica anagallis-aquatica L.

Summary

Genus:

Veronica

Family:

Plantaginaceae

Tribe:

Veroniceae

Nomen number:

80203

Place of publication:

Sp. pl. 1:12. 1753 "anagallis Δ"

Comment:

see Shenzhen ICN Art. 23.3, Ex. 2 regarding transcription of original spelling (i.e., the triangle symbol "Δ" following "anagallis" is to be transcribed as "aquatica")

Typification:

View in Linnean Typification Project

Verified:

04/21/2003 ARS Systematic Botanists.

Accessions:

3 (1 active, 1 available) in National Plant Germplasm System. (Map it)

Other conspecific taxa

Veronica anagallis-aquatica L. subsp. anagallis-aquatica (0 active accession[s])
Veronica anagallis-aquatica L. subsp. lysimachioides (Boiss.) M. A. Fisch. (0 active accession[s])
Veronica anagallis-aquatica L. subsp. michauxii (Lam.) Jelen. (0 active accession[s])
Veronica anagallis-aquatica L. subsp. oxycarpa (Boiss.) Jelen. (0 active accession[s])

Autonyms (not in current use), synonyms and invalid designations

Homotypic Synonym(s)
Veronica anagallis Bong., orth. var.

Images (2 total. Click on image for more.)

Conservation Status

Reference(s)

University of Georgia Center for Invasive Species and Ecosystem Health
Arechavaleta, M. et al., coords. 2009. Lista de especies silvestres de Canarias: hongos, plantas y animales terrestres
Arnold, T. H. & B. C. De Wet, eds. 1993. Plants of southern Africa: names and distribution. Mem. Bot. Surv. S. Africa no. 62
Boulos, L. 1995. Flora of Egypt checklist.
Brako, L. & J. L. Zarucchi. 1993. Catalogue of the flowering plants and gymnosperms of Peru. Monogr. Syst. Bot. Missouri Bot. Gard. 45
Chinese Academy of Sciences. 1959-. Flora reipublicae popularis sinicae.
Chowdhery, H. J. & B. M. Wadhwa. 1984. Flora of Himachal Pradesh.
Correll, D. S. & M. C. Johnston. 1970. Manual of the vascular plants of Texas Note: introduced
Cronquist, A. et al. 1972-. Intermountain flora.
Cufodontis, G. 1953-1972. Enumeratio plantarum aethiopiae: Spermatophyta.
Czerepanov, S. K. 1995. Vascular plants of Russia and adjacent states (the former USSR) Cambridge University Press.
Davis, P. H., ed. 1965-1988. Flora of Turkey and the east Aegean islands.
Euro+Med Editorial Committee. Euro+Med Plantbase: the information resource for Euro-Mediterranean plant diversity (on-line resource).
Exell, A. W. et al., eds. 1960-. Flora zambesiaca.
Foreman, D. B. & N. G. Walsh. 1993-. Flora of Victoria.
Gleason, H. A. & A. Cronquist. 1991. Manual of vascular plants of northeastern United States and adjacent Canada, ed. 2 Note: introduced
Greuter, W. et al., eds. 2000. International code of botanical nomenclature (Saint Louis Code) Art. 23.3, Ex. 2.
Groth, D. 2005. pers. comm. Note: re. Brazilian common names
Grubov, V. I. 2001. Key to the vascular plants of Mongolia
Hansen, A. & P. Sunding. 1993. Flora of Macaronesia: checklist of vascular plants, ed. 4. Sommerfeltia vol. 17.
Hara, H. et al. 1978-1982. An enumeration of the flowering plants of Nepal.
Harden, G. J., ed. 1990-1993. Flora of New South Wales.
Hickman, J. C., ed. 1993. The Jepson manual: higher plants of California
Hitchcock, C. L. et al. 1955-1969. Vascular plants of the Pacific Northwest.
Holm, L. et al. 1979. A geographical atlas of world weeds
Iwatsuki, K. et al. 1993-. Flora of Japan.
Jahandiez, E. & R. Maire. 1931-1941. Catalogue des plantes du Maroc.
Jiangsu Inst. Bot., ed. v. 1; Shan. R. H., ed. v. 2. 1977-1982. Jiangsu Zhiwuzhi (Flora of Jiangsu).
Kartesz, J. T. 1994. A synonymized checklist of the vascular flora of the United States, Canada, and Greenland.
Komarov, V. L. et al., eds. 1934-1964. Flora SSSR.
Marticorena, C. & M. Quezada. 1985. Catálogo de la flora vascular de Chile.
Martin, W. C. & C. R. Hutchins. 1980. A flora of New Mexico.
McGregor, R. L. et al. (The Great Plains Flora Association). 1986. Flora of the Great Plains.
Meikle, R. D. 1977-1985. Flora of Cyprus.
Merxmüller, H., ed. 1966-1972. Prodromus einer flora von Sudwestafrika.
Missouri Botanical Garden. W³Tropicos (on-line resource).
Mouterde, P. 1966-. Nouvelle flore du Liban et de la Syrie.
Öztürk, F. & A. Öztürk. 2000. A revision of the section Beccabunga Dum. belong to Veronica L. (Scrophulariaceae) genus in east anatolia. Sist. Bot. Dergisi 7:20.
Pottier-Alapetite, G. 1979-1981. Flore de la Tunisie: Angiospermes-Dicotyledones. Note: lists as "Veronica anagallis aquatica" L.
Quézel, P. & S. Santa. 1962-1963. Nouvelle flore de l'Algerie.
Rechinger, K. H., ed. 1963-. Flora iranica.
Rehm, S. 1994. Multilingual dictionary of agronomic plants
Scoggan, H. J. 1978-1979. The flora of Canada, 4 vol.
Silva, L. et al. 2005. Listagem da fauna e flora terrestres dos Açores. Lista des plantas vasculares (Pteridophyta e Spermatophyta).
Stace, C. 1995. New flora of the British Isles.
Täckholm, V. 1974. Students' flora of Egypt, ed. 2.
Tanaka, T. et al. 1999. Notes on Veronica sect. Beccabunga (Scrophulariaceae) found in yunnan province, China. J. Jap. Bot. 74:197.
Tutin, T. G. et al., eds. 1964-1980. Flora europaea.
Welsh, S. L. 1974. Anderson's flora of Alaska and adjacent parts of Canada.
Willis, J. H. 1970-1972. A handbook to plants in Victoria.
Wu Zheng-yi & P. H. Raven et al., eds. 1994-. Flora of China (English edition).
Wunderlin, R. P. 1998. Guide to the vascular plants of Florida
Zohary, M. & N. Feinbrun-Dothan. 1966-. Flora palaestina.
Zuloaga, F. O. & O. Morrone, eds. 1996, 1999. Catálogo de las plantas vasculares de la República Argentina. I. Pteridophyta, Gymnospermae y Angiospermae (Monocotyledonae), II. Dicotyledonae. Monogr. Syst. Bot. Missouri Bot. Gard. 60, 74.

Common names

Language	Name	Alternate name	seq	Citation
English	blue water speedwell		1	Foreman, D. B. & N. G. Walsh. 1993-. Flora of Victoria.
English	brook-pimpernel		1	Scoggan, H. J. 1978-1979. The flora of Canada, 4 vol.
English	water speedwell		1	Gleason, H. A. & A. Cronquist. 1991. Manual of vascular plants of northeastern United States and adjacent Canada, ed. 2 Note: introduced
Chinese	bei shui ku mai	北水苦荬	2	Wu Zheng-yi & P. H. Raven et al., eds. 1994-. Flora of China (English edition).
French	mouron d'eau		2	Rehm, S. 1994. Multilingual dictionary of agronomic plants
German	Uferehrenpreis		2	Rehm, S. 1994. Multilingual dictionary of agronomic plants
Japanese Rōmaji	ō-kawajisha		2	Iwatsuki, K. et al. 1993-. Flora of Japan.
Portuguese (Brazil)	verônica		2	Groth, D. 2005. pers. comm. Note: re. Brazilian common names
Spanish	bérula		2	Rehm, S. 1994. Multilingual dictionary of agronomic plants

Distribution

order_code	Status	Continent	Subcontinent	Country	State	Note	Reference
1	Native	Africa	East Tropical Africa	Kenya
1	Native	Africa	East Tropical Africa	Tanzania
1	Native	Africa	East Tropical Africa	Uganda
1	Native	Africa	Macaronesia	Portugal	Azores		Silva, L. et al. 2005. Listagem da fauna e flora terrestres dos Açores. Lista des plantas vasculares (Pteridophyta e Spermatophyta).
1	Native	Africa	Macaronesia	Portugal	Madeira Islands
1	Native	Africa	Macaronesia	Spain	Canarias		Arechavaleta, M. et al., coords. 2009. Lista de especies silvestres de Canarias: hongos, plantas y animales terrestres
1	Native	Africa	Northeast Tropical Africa	Ethiopia
1	Native	Africa	Northeast Tropical Africa	Sudan
1	Native	Africa	Northern Africa	Algeria		n.	Quézel, P. & S. Santa. 1962-1963. Nouvelle flore de l'Algerie.
1	Native	Africa	Northern Africa	Egypt			Boulos, L. 1995. Flora of Egypt checklist.
1	Native	Africa	Northern Africa	Libya		n.	Euro+Med Editorial Committee. Euro+Med Plantbase: the information resource for Euro-Mediterranean plant diversity (on-line resource).
1	Native	Africa	Northern Africa	Morocco			Jahandiez, E. & R. Maire. 1931-1941. Catalogue des plantes du Maroc.
1	Native	Africa	Northern Africa	Tunisia			Pottier-Alapetite, G. 1979-1981. Flore de la Tunisie: Angiospermes-Dicotyledones. Note: lists as "Veronica anagallis aquatica" L.
1	Native	Africa	South Tropical Africa	Zambia			Exell, A. W. et al., eds. 1960-. Flora zambesiaca.
1	Native	Africa	South Tropical Africa	Zimbabwe			Exell, A. W. et al., eds. 1960-. Flora zambesiaca.
1	Native	Africa	Southern Africa	Lesotho			Arnold, T. H. & B. C. De Wet, eds. 1993. Plants of southern Africa: names and distribution. Mem. Bot. Surv. S. Africa no. 62
1	Native	Africa	Southern Africa	Namibia			Arnold, T. H. & B. C. De Wet, eds. 1993. Plants of southern Africa: names and distribution. Mem. Bot. Surv. S. Africa no. 62
1	Native	Africa	Southern Africa	South Africa			Arnold, T. H. & B. C. De Wet, eds. 1993. Plants of southern Africa: names and distribution. Mem. Bot. Surv. S. Africa no. 62
1	Native	Asia-Temperate	Caucasus	Armenia			Komarov, V. L. et al., eds. 1934-1964. Flora SSSR.
1	Native	Asia-Temperate	Caucasus	Azerbaijan			Komarov, V. L. et al., eds. 1934-1964. Flora SSSR.
1	Native	Asia-Temperate	Caucasus	Georgia			Komarov, V. L. et al., eds. 1934-1964. Flora SSSR.
1	Native	Asia-Temperate	Caucasus	Russian Federation	Dagestan		Komarov, V. L. et al., eds. 1934-1964. Flora SSSR.
1	Native	Asia-Temperate	Caucasus	Russian Federation-Ciscaucasia	Ciscaucasia		Komarov, V. L. et al., eds. 1934-1964. Flora SSSR.
1	Native	Asia-Temperate	China	China	Anhui Sheng		Wu Zheng-yi & P. H. Raven et al., eds. 1994-. Flora of China (English edition).
1	Native	Asia-Temperate	China	China	Gansu Sheng		Wu Zheng-yi & P. H. Raven et al., eds. 1994-. Flora of China (English edition).
1	Native	Asia-Temperate	China	China	Guizhou Sheng		Wu Zheng-yi & P. H. Raven et al., eds. 1994-. Flora of China (English edition).
1	Native	Asia-Temperate	China	China	Hebei Sheng		Wu Zheng-yi & P. H. Raven et al., eds. 1994-. Flora of China (English edition).
1	Native	Asia-Temperate	China	China	Heilongjiang Sheng		Wu Zheng-yi & P. H. Raven et al., eds. 1994-. Flora of China (English edition).
1	Native	Asia-Temperate	China	China	Henan Sheng		Wu Zheng-yi & P. H. Raven et al., eds. 1994-. Flora of China (English edition).
1	Native	Asia-Temperate	China	China	Hubei Sheng		Wu Zheng-yi & P. H. Raven et al., eds. 1994-. Flora of China (English edition).
1	Native	Asia-Temperate	China	China	Jiangsu Sheng		Wu Zheng-yi & P. H. Raven et al., eds. 1994-. Flora of China (English edition).
1	Native	Asia-Temperate	China	China	Jilin Sheng		Wu Zheng-yi & P. H. Raven et al., eds. 1994-. Flora of China (English edition).
1	Native	Asia-Temperate	China	China	Liaoning Sheng		Wu Zheng-yi & P. H. Raven et al., eds. 1994-. Flora of China (English edition).
1	Native	Asia-Temperate	China	China	Ningxia Huizi Zizhiqu		Wu Zheng-yi & P. H. Raven et al., eds. 1994-. Flora of China (English edition).
1	Native	Asia-Temperate	China	China	Qinghai Sheng		Wu Zheng-yi & P. H. Raven et al., eds. 1994-. Flora of China (English edition).
1	Native	Asia-Temperate	China	China	Shaanxi Sheng		Wu Zheng-yi & P. H. Raven et al., eds. 1994-. Flora of China (English edition).
1	Native	Asia-Temperate	China	China	Shandong Sheng		Wu Zheng-yi & P. H. Raven et al., eds. 1994-. Flora of China (English edition).
1	Native	Asia-Temperate	China	China	Shanxi Sheng		Wu Zheng-yi & P. H. Raven et al., eds. 1994-. Flora of China (English edition).
1	Native	Asia-Temperate	China	China	Sichuan Sheng		Wu Zheng-yi & P. H. Raven et al., eds. 1994-. Flora of China (English edition).
1	Native	Asia-Temperate	China	China	Xinjiang Uygur Zizhiqu		Wu Zheng-yi & P. H. Raven et al., eds. 1994-. Flora of China (English edition).
1	Native	Asia-Temperate	China	China	Xizang Zizhiqu		Wu Zheng-yi & P. H. Raven et al., eds. 1994-. Flora of China (English edition).
1	Native	Asia-Temperate	China	China	Yunnan Sheng		Wu Zheng-yi & P. H. Raven et al., eds. 1994-. Flora of China (English edition).
1	Native	Asia-Temperate	Eastern Asia	Korea
1	Native	Asia-Temperate	Middle Asia	Kazakhstan			Komarov, V. L. et al., eds. 1934-1964. Flora SSSR.
1	Native	Asia-Temperate	Middle Asia	Kyrgyzstan			Komarov, V. L. et al., eds. 1934-1964. Flora SSSR.
1	Native	Asia-Temperate	Middle Asia	Tajikistan			Komarov, V. L. et al., eds. 1934-1964. Flora SSSR.
1	Native	Asia-Temperate	Middle Asia	Turkmenistan			Komarov, V. L. et al., eds. 1934-1964. Flora SSSR.
1	Native	Asia-Temperate	Middle Asia	Uzbekistan			Komarov, V. L. et al., eds. 1934-1964. Flora SSSR.
1	Native	Asia-Temperate	Mongolia	Mongolia			Grubov, V. I. 2001. Key to the vascular plants of Mongolia
1	Native	Asia-Temperate	Siberia	Russian Federation-Eastern Siberia	Eastern Siberia		Komarov, V. L. et al., eds. 1934-1964. Flora SSSR.
1	Native	Asia-Temperate	Siberia	Russian Federation-Western Siberia	Western Siberia		Komarov, V. L. et al., eds. 1934-1964. Flora SSSR.
1	Native	Asia-Temperate	Western Asia	Afghanistan			Rechinger, K. H., ed. 1963-. Flora iranica.
1	Native	Asia-Temperate	Western Asia	Cyprus			Meikle, R. D. 1977-1985. Flora of Cyprus.
1	Native	Asia-Temperate	Western Asia	Iran			Rechinger, K. H., ed. 1963-. Flora iranica.
1	Native	Asia-Temperate	Western Asia	Iraq			Rechinger, K. H., ed. 1963-. Flora iranica.
1	Native	Asia-Temperate	Western Asia	Israel			Zohary, M. & N. Feinbrun-Dothan. 1966-. Flora palaestina.
1	Native	Asia-Temperate	Western Asia	Jordan			Zohary, M. & N. Feinbrun-Dothan. 1966-. Flora palaestina.
1	Native	Asia-Temperate	Western Asia	Lebanon			Mouterde, P. 1966-. Nouvelle flore du Liban et de la Syrie.
1	Native	Asia-Temperate	Western Asia	Syria			Mouterde, P. 1966-. Nouvelle flore du Liban et de la Syrie.
1	Native	Asia-Temperate	Western Asia	Turkey	Adana		Davis, P. H., ed. 1965-1988. Flora of Turkey and the east Aegean islands.
1	Native	Asia-Temperate	Western Asia	Turkey	Ankara		Davis, P. H., ed. 1965-1988. Flora of Turkey and the east Aegean islands.
1	Native	Asia-Temperate	Western Asia	Turkey	Antalya		Davis, P. H., ed. 1965-1988. Flora of Turkey and the east Aegean islands.
1	Native	Asia-Temperate	Western Asia	Turkey	Aydın		Davis, P. H., ed. 1965-1988. Flora of Turkey and the east Aegean islands.
1	Native	Asia-Temperate	Western Asia	Turkey	Bolu		Davis, P. H., ed. 1965-1988. Flora of Turkey and the east Aegean islands.
1	Native	Asia-Temperate	Western Asia	Turkey	Çankırı		Davis, P. H., ed. 1965-1988. Flora of Turkey and the east Aegean islands.
1	Native	Asia-Temperate	Western Asia	Turkey	Denizli		Davis, P. H., ed. 1965-1988. Flora of Turkey and the east Aegean islands.
1	Native	Asia-Temperate	Western Asia	Turkey	Diyarbakır		Davis, P. H., ed. 1965-1988. Flora of Turkey and the east Aegean islands.
1	Native	Asia-Temperate	Western Asia	Turkey	Erzurum		Davis, P. H., ed. 1965-1988. Flora of Turkey and the east Aegean islands.
1	Native	Asia-Temperate	Western Asia	Turkey	Icel		Davis, P. H., ed. 1965-1988. Flora of Turkey and the east Aegean islands.
1	Native	Asia-Temperate	Western Asia	Turkey	Isparta		Davis, P. H., ed. 1965-1988. Flora of Turkey and the east Aegean islands.
1	Native	Asia-Temperate	Western Asia	Turkey	İzmir		Davis, P. H., ed. 1965-1988. Flora of Turkey and the east Aegean islands.
1	Native	Asia-Temperate	Western Asia	Turkey	Kars		Davis, P. H., ed. 1965-1988. Flora of Turkey and the east Aegean islands.
1	Native	Asia-Temperate	Western Asia	Turkey	Kastamonu		Davis, P. H., ed. 1965-1988. Flora of Turkey and the east Aegean islands.
1	Native	Asia-Temperate	Western Asia	Turkey	Kayseri		Davis, P. H., ed. 1965-1988. Flora of Turkey and the east Aegean islands.
1	Native	Asia-Temperate	Western Asia	Turkey	Kütahya		Davis, P. H., ed. 1965-1988. Flora of Turkey and the east Aegean islands.
1	Native	Asia-Temperate	Western Asia	Turkey	Rize		Davis, P. H., ed. 1965-1988. Flora of Turkey and the east Aegean islands.
1	Native	Asia-Temperate	Western Asia	Turkey	Samsun		Davis, P. H., ed. 1965-1988. Flora of Turkey and the east Aegean islands.
1	Native	Asia-Temperate	Western Asia	Turkey	Sivas		Davis, P. H., ed. 1965-1988. Flora of Turkey and the east Aegean islands.
1	Native	Asia-Temperate	Western Asia	Turkey	Trabzon		Davis, P. H., ed. 1965-1988. Flora of Turkey and the east Aegean islands.
1	Native	Asia-Temperate	Western Asia	Turkey	Urfa		Davis, P. H., ed. 1965-1988. Flora of Turkey and the east Aegean islands.
1	Native	Asia-Temperate	Western Asia	Turkey	Van		Davis, P. H., ed. 1965-1988. Flora of Turkey and the east Aegean islands.
1	Native	Asia-Tropical	Indian Subcontinent	Bhutan
1	Native	Asia-Tropical	Indian Subcontinent	India
1	Native	Asia-Tropical	Indian Subcontinent	Nepal			Hara, H. et al. 1978-1982. An enumeration of the flowering plants of Nepal.
1	Native	Asia-Tropical	Indian Subcontinent	Pakistan
1	Native	Europe	Eastern Europe	Belarus			Euro+Med Editorial Committee. Euro+Med Plantbase: the information resource for Euro-Mediterranean plant diversity (on-line resource).
1	Native	Europe	Eastern Europe	Estonia			Euro+Med Editorial Committee. Euro+Med Plantbase: the information resource for Euro-Mediterranean plant diversity (on-line resource).
1	Native	Europe	Eastern Europe	Latvia			Euro+Med Editorial Committee. Euro+Med Plantbase: the information resource for Euro-Mediterranean plant diversity (on-line resource).
1	Native	Europe	Eastern Europe	Lithuania			Euro+Med Editorial Committee. Euro+Med Plantbase: the information resource for Euro-Mediterranean plant diversity (on-line resource).
1	Native	Europe	Eastern Europe	Moldova			Euro+Med Editorial Committee. Euro+Med Plantbase: the information resource for Euro-Mediterranean plant diversity (on-line resource).
1	Native	Europe	Eastern Europe	Russian Federation-European part	European part		Komarov, V. L. et al., eds. 1934-1964. Flora SSSR.
1	Native	Europe	Eastern Europe	Ukraine		incl. Krym	Euro+Med Editorial Committee. Euro+Med Plantbase: the information resource for Euro-Mediterranean plant diversity (on-line resource).
1	Native	Europe	Middle Europe	Austria			Tutin, T. G. et al., eds. 1964-1980. Flora europaea.
1	Native	Europe	Middle Europe	Belgium			Tutin, T. G. et al., eds. 1964-1980. Flora europaea.
1	Native	Europe	Middle Europe	Czech Republic			Euro+Med Editorial Committee. Euro+Med Plantbase: the information resource for Euro-Mediterranean plant diversity (on-line resource).
1	Native	Europe	Middle Europe	Germany			Tutin, T. G. et al., eds. 1964-1980. Flora europaea.
1	Native	Europe	Middle Europe	Hungary			Tutin, T. G. et al., eds. 1964-1980. Flora europaea.
1	Native	Europe	Middle Europe	Netherlands			Tutin, T. G. et al., eds. 1964-1980. Flora europaea.
1	Native	Europe	Middle Europe	Poland			Tutin, T. G. et al., eds. 1964-1980. Flora europaea.
1	Native	Europe	Middle Europe	Slovakia			Euro+Med Editorial Committee. Euro+Med Plantbase: the information resource for Euro-Mediterranean plant diversity (on-line resource).
1	Native	Europe	Middle Europe	Switzerland			Tutin, T. G. et al., eds. 1964-1980. Flora europaea.
1	Native	Europe	Northern Europe	Denmark			Tutin, T. G. et al., eds. 1964-1980. Flora europaea.
1	Native	Europe	Northern Europe	Finland			Tutin, T. G. et al., eds. 1964-1980. Flora europaea.
1	Native	Europe	Northern Europe	Iceland			Tutin, T. G. et al., eds. 1964-1980. Flora europaea.
1	Native	Europe	Northern Europe	Ireland			Tutin, T. G. et al., eds. 1964-1980. Flora europaea.
1	Native	Europe	Northern Europe	Norway			Tutin, T. G. et al., eds. 1964-1980. Flora europaea.
1	Native	Europe	Northern Europe	Sweden			Tutin, T. G. et al., eds. 1964-1980. Flora europaea.
1	Native	Europe	Northern Europe	United Kingdom			Tutin, T. G. et al., eds. 1964-1980. Flora europaea.
1	Native	Europe	Southeastern Europe	Albania			Tutin, T. G. et al., eds. 1964-1980. Flora europaea.
1	Native	Europe	Southeastern Europe	Bulgaria			Tutin, T. G. et al., eds. 1964-1980. Flora europaea.
1	Native	Europe	Southeastern Europe	Croatia			Euro+Med Editorial Committee. Euro+Med Plantbase: the information resource for Euro-Mediterranean plant diversity (on-line resource).
1	Native	Europe	Southeastern Europe	Greece		incl. Crete	Tutin, T. G. et al., eds. 1964-1980. Flora europaea.
1	Native	Europe	Southeastern Europe	Italy		incl. Sardinia, Sicily	Tutin, T. G. et al., eds. 1964-1980. Flora europaea.
1	Native	Europe	Southeastern Europe	Romania			Tutin, T. G. et al., eds. 1964-1980. Flora europaea.
1	Native	Europe	Southeastern Europe	Serbia			Euro+Med Editorial Committee. Euro+Med Plantbase: the information resource for Euro-Mediterranean plant diversity (on-line resource).
1	Native	Europe	Southeastern Europe	Slovenia			Euro+Med Editorial Committee. Euro+Med Plantbase: the information resource for Euro-Mediterranean plant diversity (on-line resource).
1	Native	Europe	Southeastern Europe	Turkey	İstanbul		Tutin, T. G. et al., eds. 1964-1980. Flora europaea.
1	Native	Europe	Southeastern Europe	Turkey	Kırklareli		Tutin, T. G. et al., eds. 1964-1980. Flora europaea.
1	Native	Europe	Southwestern Europe	France		incl. Corsica	Tutin, T. G. et al., eds. 1964-1980. Flora europaea.
1	Native	Europe	Southwestern Europe	Portugal			Tutin, T. G. et al., eds. 1964-1980. Flora europaea.
1	Native	Europe	Southwestern Europe	Spain		incl. Baleares	Tutin, T. G. et al., eds. 1964-1980. Flora europaea.
1	Native	Southern America	Southern South America	Argentina	Buenos Aires		Zuloaga, F. O. & O. Morrone, eds. 1996, 1999. Catálogo de las plantas vasculares de la República Argentina. I. Pteridophyta, Gymnospermae y Angiospermae (Monocotyledonae), II. Dicotyledonae. Monogr. Syst. Bot. Missouri Bot. Gard. 60, 74.
1	Native	Southern America	Southern South America	Argentina	Catamarca		Zuloaga, F. O. & O. Morrone, eds. 1996, 1999. Catálogo de las plantas vasculares de la República Argentina. I. Pteridophyta, Gymnospermae y Angiospermae (Monocotyledonae), II. Dicotyledonae. Monogr. Syst. Bot. Missouri Bot. Gard. 60, 74.
1	Native	Southern America	Southern South America	Argentina	Chubut		Zuloaga, F. O. & O. Morrone, eds. 1996, 1999. Catálogo de las plantas vasculares de la República Argentina. I. Pteridophyta, Gymnospermae y Angiospermae (Monocotyledonae), II. Dicotyledonae. Monogr. Syst. Bot. Missouri Bot. Gard. 60, 74.
1	Native	Southern America	Southern South America	Argentina	Jujuy		Zuloaga, F. O. & O. Morrone, eds. 1996, 1999. Catálogo de las plantas vasculares de la República Argentina. I. Pteridophyta, Gymnospermae y Angiospermae (Monocotyledonae), II. Dicotyledonae. Monogr. Syst. Bot. Missouri Bot. Gard. 60, 74.
1	Native	Southern America	Southern South America	Argentina	La Pampa		Zuloaga, F. O. & O. Morrone, eds. 1996, 1999. Catálogo de las plantas vasculares de la República Argentina. I. Pteridophyta, Gymnospermae y Angiospermae (Monocotyledonae), II. Dicotyledonae. Monogr. Syst. Bot. Missouri Bot. Gard. 60, 74.
1	Native	Southern America	Southern South America	Argentina	La Rioja		Zuloaga, F. O. & O. Morrone, eds. 1996, 1999. Catálogo de las plantas vasculares de la República Argentina. I. Pteridophyta, Gymnospermae y Angiospermae (Monocotyledonae), II. Dicotyledonae. Monogr. Syst. Bot. Missouri Bot. Gard. 60, 74.
1	Native	Southern America	Southern South America	Argentina	Mendoza		Zuloaga, F. O. & O. Morrone, eds. 1996, 1999. Catálogo de las plantas vasculares de la República Argentina. I. Pteridophyta, Gymnospermae y Angiospermae (Monocotyledonae), II. Dicotyledonae. Monogr. Syst. Bot. Missouri Bot. Gard. 60, 74.
1	Native	Southern America	Southern South America	Argentina	Neuquén		Zuloaga, F. O. & O. Morrone, eds. 1996, 1999. Catálogo de las plantas vasculares de la República Argentina. I. Pteridophyta, Gymnospermae y Angiospermae (Monocotyledonae), II. Dicotyledonae. Monogr. Syst. Bot. Missouri Bot. Gard. 60, 74.
1	Native	Southern America	Southern South America	Argentina	Río Negro		Zuloaga, F. O. & O. Morrone, eds. 1996, 1999. Catálogo de las plantas vasculares de la República Argentina. I. Pteridophyta, Gymnospermae y Angiospermae (Monocotyledonae), II. Dicotyledonae. Monogr. Syst. Bot. Missouri Bot. Gard. 60, 74.
1	Native	Southern America	Southern South America	Argentina	Salta		Zuloaga, F. O. & O. Morrone, eds. 1996, 1999. Catálogo de las plantas vasculares de la República Argentina. I. Pteridophyta, Gymnospermae y Angiospermae (Monocotyledonae), II. Dicotyledonae. Monogr. Syst. Bot. Missouri Bot. Gard. 60, 74.
1	Native	Southern America	Southern South America	Argentina	San Juan		Zuloaga, F. O. & O. Morrone, eds. 1996, 1999. Catálogo de las plantas vasculares de la República Argentina. I. Pteridophyta, Gymnospermae y Angiospermae (Monocotyledonae), II. Dicotyledonae. Monogr. Syst. Bot. Missouri Bot. Gard. 60, 74.
1	Native	Southern America	Southern South America	Argentina	San Luis		Zuloaga, F. O. & O. Morrone, eds. 1996, 1999. Catálogo de las plantas vasculares de la República Argentina. I. Pteridophyta, Gymnospermae y Angiospermae (Monocotyledonae), II. Dicotyledonae. Monogr. Syst. Bot. Missouri Bot. Gard. 60, 74.
1	Native	Southern America	Southern South America	Argentina	Tierra del Fuego		Zuloaga, F. O. & O. Morrone, eds. 1996, 1999. Catálogo de las plantas vasculares de la República Argentina. I. Pteridophyta, Gymnospermae y Angiospermae (Monocotyledonae), II. Dicotyledonae. Monogr. Syst. Bot. Missouri Bot. Gard. 60, 74.
1	Native	Southern America	Southern South America	Chile
1	Native	Southern America	Southern South America	Uruguay			Zuloaga, F. O. & O. Morrone, eds. 1996, 1999. Catálogo de las plantas vasculares de la República Argentina. I. Pteridophyta, Gymnospermae y Angiospermae (Monocotyledonae), II. Dicotyledonae. Monogr. Syst. Bot. Missouri Bot. Gard. 60, 74.
1	Native	Southern America	Western South America	Peru	Ancash		Brako, L. & J. L. Zarucchi. 1993. Catalogue of the flowering plants and gymnosperms of Peru. Monogr. Syst. Bot. Missouri Bot. Gard. 45
1	Native	Southern America	Western South America	Peru	Arequipa		Brako, L. & J. L. Zarucchi. 1993. Catalogue of the flowering plants and gymnosperms of Peru. Monogr. Syst. Bot. Missouri Bot. Gard. 45
1	Native	Southern America	Western South America	Peru	Cajamarca		Brako, L. & J. L. Zarucchi. 1993. Catalogue of the flowering plants and gymnosperms of Peru. Monogr. Syst. Bot. Missouri Bot. Gard. 45
1	Native	Southern America	Western South America	Peru	Lima		Brako, L. & J. L. Zarucchi. 1993. Catalogue of the flowering plants and gymnosperms of Peru. Monogr. Syst. Bot. Missouri Bot. Gard. 45
4	Naturalized					natzd. in temperate regions
6	Other					exact native range obscure

Native
Africa

MACARONESIA: Spain [Canarias], Portugal [Azores, Madeira Islands]
NORTHERN AFRICA: Algeria (n.), Egypt, Libya (n.), Morocco, Tunisia
NORTHEAST TROPICAL AFRICA: Ethiopia, Sudan
EAST TROPICAL AFRICA: Kenya, Tanzania, Uganda
SOUTH TROPICAL AFRICA: Zambia, Zimbabwe
SOUTHERN AFRICA: Lesotho, Namibia, South Africa

Asia-Temperate

WESTERN ASIA: Afghanistan, Cyprus, Iran, Iraq, Israel, Jordan, Lebanon, Syria, Turkey [Adana, Ankara, Antalya, Aydın, Bolu, Çankırı, Denizli, Diyarbakır, Erzurum, Icel, Isparta, İzmir, Kars, Kastamonu, Kayseri, Kütahya, Rize, Samsun, Sivas, Trabzon, Urfa, Van]
CAUCASUS: Russian Federation-Ciscaucasia [Ciscaucasia], Armenia, Azerbaijan, Georgia, Russian Federation [Dagestan]
SIBERIA: Russian Federation-Western Siberia [Western Siberia], Russian Federation-Eastern Siberia [Eastern Siberia]
MIDDLE ASIA: Kazakhstan, Kyrgyzstan, Tajikistan, Turkmenistan, Uzbekistan
MONGOLIA: Mongolia
CHINA: China [Anhui Sheng, Gansu Sheng, Guizhou Sheng, Hebei Sheng, Heilongjiang Sheng, Henan Sheng, Hubei Sheng, Jiangsu Sheng, Jilin Sheng, Liaoning Sheng, Ningxia Huizi Zizhiqu, Qinghai Sheng, Shaanxi Sheng, Shandong Sheng, Shanxi Sheng, Sichuan Sheng, Xinjiang Uygur Zizhiqu, Xizang Zizhiqu, Yunnan Sheng]
EASTERN ASIA: Korea

Asia-Tropical

INDIAN SUBCONTINENT: Bhutan, India, Nepal, Pakistan

Europe

NORTHERN EUROPE: Denmark, Finland, United Kingdom, Ireland, Iceland, Norway, Sweden
MIDDLE EUROPE: Austria, Belgium, Switzerland, Czech Republic, Germany, Hungary, Netherlands, Poland, Slovakia
EASTERN EUROPE: Russian Federation-European part [European part], Belarus, Estonia, Lithuania, Latvia, Moldova, Ukraine (incl. Krym)
SOUTHEASTERN EUROPE: Albania, Bulgaria, Greece (incl. Crete), Croatia, Italy (incl. Sardinia, Sicily), Romania, Serbia, Slovenia, Turkey [İstanbul, Kırklareli]
SOUTHWESTERN EUROPE: Spain (incl. Baleares), France (incl. Corsica), Portugal

Southern America

WESTERN SOUTH AMERICA: Peru [Ancash, Arequipa, Cajamarca, Lima]
SOUTHERN SOUTH AMERICA: Argentina [Buenos Aires, Catamarca, Chubut, Jujuy, La Pampa, La Rioja, Mendoza, Neuquén, Río Negro, Salta, San Juan, San Luis, Tierra del Fuego], Chile, Uruguay

Naturalized (natzd. in temperate regions)
Other (exact native range obscure)

Reference(s)

Arechavaleta, M. et al., coords. 2009. Lista de especies silvestres de Canarias: hongos, plantas y animales terrestres
Arnold, T. H. & B. C. De Wet, eds. 1993. Plants of southern Africa: names and distribution. Mem. Bot. Surv. S. Africa no. 62
Boulos, L. 1995. Flora of Egypt checklist.
Brako, L. & J. L. Zarucchi. 1993. Catalogue of the flowering plants and gymnosperms of Peru. Monogr. Syst. Bot. Missouri Bot. Gard. 45
Davis, P. H., ed. 1965-1988. Flora of Turkey and the east Aegean islands.
Euro+Med Editorial Committee. Euro+Med Plantbase: the information resource for Euro-Mediterranean plant diversity (on-line resource).
Exell, A. W. et al., eds. 1960-. Flora zambesiaca.
Grubov, V. I. 2001. Key to the vascular plants of Mongolia
Hara, H. et al. 1978-1982. An enumeration of the flowering plants of Nepal.
Jahandiez, E. & R. Maire. 1931-1941. Catalogue des plantes du Maroc.
Komarov, V. L. et al., eds. 1934-1964. Flora SSSR.
Meikle, R. D. 1977-1985. Flora of Cyprus.
Mouterde, P. 1966-. Nouvelle flore du Liban et de la Syrie.
Pottier-Alapetite, G. 1979-1981. Flore de la Tunisie: Angiospermes-Dicotyledones. Note: lists as "Veronica anagallis aquatica" L.
Quézel, P. & S. Santa. 1962-1963. Nouvelle flore de l'Algerie.
Rechinger, K. H., ed. 1963-. Flora iranica.
Silva, L. et al. 2005. Listagem da fauna e flora terrestres dos Açores. Lista des plantas vasculares (Pteridophyta e Spermatophyta).
Tutin, T. G. et al., eds. 1964-1980. Flora europaea.
Wu Zheng-yi & P. H. Raven et al., eds. 1994-. Flora of China (English edition).
Zohary, M. & N. Feinbrun-Dothan. 1966-. Flora palaestina.
Zuloaga, F. O. & O. Morrone, eds. 1996, 1999. Catálogo de las plantas vasculares de la República Argentina. I. Pteridophyta, Gymnospermae y Angiospermae (Monocotyledonae), II. Dicotyledonae. Monogr. Syst. Bot. Missouri Bot. Gard. 60, 74.

Economic Uses

Usage	Type	Note	Reference
Medicines	folklore		Wu Zheng-yi & P. H. Raven et al., eds. 1994-. Flora of China (English edition).
Weed	seed contaminant	potential; common or of unknown importance	Holm, L. et al. 1979. A geographical atlas of world weeds

Cite as: USDA, Agricultural Research Service, National Plant Germplasm System. 2026. Germplasm Resources Information Network (GRIN Taxonomy). National Germplasm Resources Laboratory, Beltsville, Maryland.
URL: https://npgstest.ars-grin.gov/gringlobal/taxon/taxonomydetail?id=80203. Accessed 14 May 2026.