Taxon: Samolus valerandi L.

Summary

Genus:

Samolus

Family:

Primulaceae

Subfamily:

Theophrastoideae

Tribe:

Samoleae

Nomen number:

459207

Place of publication:

Sp. pl. 1:171. 1753

Typification:

View in Linnean Typification Project

Verified:

04/30/2009 ARS Systematic Botanists.

Accessions:

1 (1 active, 1 available) in National Plant Germplasm System. (Map it)

Other conspecific taxa

Autonyms (not in current use), synonyms and invalid designations

Autonym(s)
Samolus valerandi L. subsp. valerandi

Heterotypic Synonym(s)
Samolus floribundus Kunth
Samolus parviflorus Raf.
Samolus valerandi L. subsp. parviflorus (Raf.) Hultén

No images

Conservation Status

Reference(s)

Aldén, B., S. Ryman, & M. Hjertson. 2012. Svensk Kulturväxtdatabas, SKUD (Swedish Cultivated and Utility Plants Database; online resource)
Botanical Society of the British Isles. BSBI taxon database (on-line resource).
Brako, L. & J. L. Zarucchi. 1993. Catalogue of the flowering plants and gymnosperms of Peru. Monogr. Syst. Bot. Missouri Bot. Gard. 45
Cronquist, A. et al. 1972-. Intermountain flora.
Dyer, R. A. et al., eds. 1963-. Flora of southern Africa.
Exell, A. W. et al., eds. 1960-. Flora zambesiaca.
Food and Agriculture Organization of the United Nations (FAO). 2010. Ecocrop (on-line resource).
Ghazanfar, S. A. 1992. An annotated catalogue of the vascular plants of Oman. Scripta Botanica Belgica 2
Greuter, W. et al., eds. 1984-. Med-Checklist.
Harling, G. & B. Sparre, eds. 1973-. Flora of Ecuador.
Jørgensen, P. M. & S. León-Yánez, eds. 1999. Catalogue of the vascular plants of Ecuador. Monogr. Syst. Bot. Missouri Bot. Gard. 75
Keay, R. W. J. & F. N. Hepper. 1953-1972. Flora of west tropical Africa, ed. 2.
Komarov, V. L. et al., eds. 1934-1964. Flora SSSR.
Migahid, A. M. 1988-1990. Flora of Saudi Arabia, ed. 3.
Nasir, E. & S. I. Ali, eds. 1970-. Flora of [West] Pakistan.
Rechinger, K. H., ed. 1963-. Flora iranica.
Robyns, W. et al., eds. 1948-1963. Flore du Congo belge et du Ruanda-Urundi
Thulin, M., ed. 1993-. Flora of Somalia.
Turrill, W. B. et al., eds. 1952-. Flora of tropical East Africa.
Tutin, T. G. et al., eds. 1964-1980. Flora europaea.
Wood, J. R. I. 1997. A handbook of the Yemen flora.
Wu Zheng-yi & P. H. Raven et al., eds. 1994-. Flora of China (English edition).
Zuloaga, F. O. & O. Morrone, eds. 1996, 1999. Catálogo de las plantas vasculares de la República Argentina. I. Pteridophyta, Gymnospermae y Angiospermae (Monocotyledonae), II. Dicotyledonae. Monogr. Syst. Bot. Missouri Bot. Gard. 60, 74.

Common names

Language	Name	Alternate name	note	seq	Citation
English	brookweed			1	Botanical Society of the British Isles. BSBI taxon database (on-line resource).
Swedish	bunge			2	Aldén, B., S. Ryman, & M. Hjertson. 2012. Svensk Kulturväxtdatabas, SKUD (Swedish Cultivated and Utility Plants Database; online resource)

Distribution

order_code	Status	Continent	Subcontinent	Country	State	Note	Reference
1	Native	Africa	East Tropical Africa	Kenya			Turrill, W. B. et al., eds. 1952-. Flora of tropical East Africa.
1	Native	Africa	Northeast Tropical Africa	Eritrea			Thulin, M., ed. 1993-. Flora of Somalia.
1	Native	Africa	Northeast Tropical Africa	Ethiopia			Thulin, M., ed. 1993-. Flora of Somalia.
1	Native	Africa	Northeast Tropical Africa	Somalia			Thulin, M., ed. 1993-. Flora of Somalia.
1	Native	Africa	Northern Africa	Algeria			Greuter, W. et al., eds. 1984-. Med-Checklist.
1	Native	Africa	Northern Africa	Egypt			Greuter, W. et al., eds. 1984-. Med-Checklist.
1	Native	Africa	Northern Africa	Libya			Greuter, W. et al., eds. 1984-. Med-Checklist.
1	Native	Africa	Northern Africa	Morocco			Greuter, W. et al., eds. 1984-. Med-Checklist.
1	Native	Africa	Northern Africa	Tunisia			Greuter, W. et al., eds. 1984-. Med-Checklist.
1	Native	Africa	South Tropical Africa	Mozambique			Exell, A. W. et al., eds. 1960-. Flora zambesiaca.
1	Native	Africa	South Tropical Africa	Zambia			Exell, A. W. et al., eds. 1960-. Flora zambesiaca.
1	Native	Africa	South Tropical Africa	Zimbabwe			Exell, A. W. et al., eds. 1960-. Flora zambesiaca.
1	Native	Africa	Southern Africa	Botswana			Exell, A. W. et al., eds. 1960-. Flora zambesiaca.
1	Native	Africa	Southern Africa	Namibia			Dyer, R. A. et al., eds. 1963-. Flora of southern Africa.
1	Native	Africa	Southern Africa	South Africa	Cape Province		Dyer, R. A. et al., eds. 1963-. Flora of southern Africa.
1	Native	Africa	Southern Africa	South Africa	KwaZulu-Natal		Dyer, R. A. et al., eds. 1963-. Flora of southern Africa.
1	Native	Africa	Southern Africa	South Africa	Transvaal		Dyer, R. A. et al., eds. 1963-. Flora of southern Africa.
1	Native	Africa	West Tropical Africa	Nigeria			Keay, R. W. J. & F. N. Hepper. 1953-1972. Flora of west tropical Africa, ed. 2.
1	Native	Africa	West-Central Tropical Africa	Democratic Republic of the Congo			Robyns, W. et al., eds. 1948-1963. Flore du Congo belge et du Ruanda-Urundi
1	Native	Asia-Temperate	Arabian Peninsula	Oman			Ghazanfar, S. A. 1992. An annotated catalogue of the vascular plants of Oman. Scripta Botanica Belgica 2
1	Native	Asia-Temperate	Arabian Peninsula	Saudi Arabia			Migahid, A. M. 1988-1990. Flora of Saudi Arabia, ed. 3.
1	Native	Asia-Temperate	Arabian Peninsula	Yemen			Wood, J. R. I. 1997. A handbook of the Yemen flora.
1	Native	Asia-Temperate	Caucasus	Armenia			Komarov, V. L. et al., eds. 1934-1964. Flora SSSR.
1	Native	Asia-Temperate	Caucasus	Azerbaijan			Komarov, V. L. et al., eds. 1934-1964. Flora SSSR.
1	Native	Asia-Temperate	Caucasus	Georgia			Komarov, V. L. et al., eds. 1934-1964. Flora SSSR.
1	Native	Asia-Temperate	Caucasus	Russian Federation	Dagestan		Komarov, V. L. et al., eds. 1934-1964. Flora SSSR.
1	Native	Asia-Temperate	Caucasus	Russian Federation-Ciscaucasia	Ciscaucasia		Komarov, V. L. et al., eds. 1934-1964. Flora SSSR.
1	Native	Asia-Temperate	China	China	Guangdong Sheng		Wu Zheng-yi & P. H. Raven et al., eds. 1994-. Flora of China (English edition).
1	Native	Asia-Temperate	China	China	Guangxi Zhuangzu Zizhiqu		Wu Zheng-yi & P. H. Raven et al., eds. 1994-. Flora of China (English edition).
1	Native	Asia-Temperate	China	China	Guizhou Sheng		Wu Zheng-yi & P. H. Raven et al., eds. 1994-. Flora of China (English edition).
1	Native	Asia-Temperate	China	China	Hunan Sheng		Wu Zheng-yi & P. H. Raven et al., eds. 1994-. Flora of China (English edition).
1	Native	Asia-Temperate	China	China	Yunnan Sheng		Wu Zheng-yi & P. H. Raven et al., eds. 1994-. Flora of China (English edition).
1	Native	Asia-Temperate	Middle Asia	Kyrgyzstan			Komarov, V. L. et al., eds. 1934-1964. Flora SSSR.
1	Native	Asia-Temperate	Middle Asia	Tajikistan			Komarov, V. L. et al., eds. 1934-1964. Flora SSSR.
1	Native	Asia-Temperate	Middle Asia	Turkmenistan			Komarov, V. L. et al., eds. 1934-1964. Flora SSSR.
1	Native	Asia-Temperate	Middle Asia	Uzbekistan			Komarov, V. L. et al., eds. 1934-1964. Flora SSSR.
1	Native	Asia-Temperate	Western Asia	Afghanistan			Rechinger, K. H., ed. 1963-. Flora iranica.
1	Native	Asia-Temperate	Western Asia	Cyprus			Greuter, W. et al., eds. 1984-. Med-Checklist.
1	Native	Asia-Temperate	Western Asia	Egypt	Sinai		Greuter, W. et al., eds. 1984-. Med-Checklist.
1	Native	Asia-Temperate	Western Asia	Iran			Rechinger, K. H., ed. 1963-. Flora iranica.
1	Native	Asia-Temperate	Western Asia	Israel			Greuter, W. et al., eds. 1984-. Med-Checklist.
1	Native	Asia-Temperate	Western Asia	Jordan			Greuter, W. et al., eds. 1984-. Med-Checklist.
1	Native	Asia-Temperate	Western Asia	Lebanon			Greuter, W. et al., eds. 1984-. Med-Checklist.
1	Native	Asia-Temperate	Western Asia	Syria			Greuter, W. et al., eds. 1984-. Med-Checklist.
1	Native	Asia-Temperate	Western Asia	Turkey			Greuter, W. et al., eds. 1984-. Med-Checklist.
1	Native	Asia-Tropical	Indian Subcontinent	India			Nasir, E. & S. I. Ali, eds. 1970-. Flora of [West] Pakistan.
1	Native	Asia-Tropical	Indian Subcontinent	Pakistan			Nasir, E. & S. I. Ali, eds. 1970-. Flora of [West] Pakistan.
1	Native	Europe	Eastern Europe	Belarus			Tutin, T. G. et al., eds. 1964-1980. Flora europaea.
1	Native	Europe	Eastern Europe	Estonia			Tutin, T. G. et al., eds. 1964-1980. Flora europaea.
1	Native	Europe	Eastern Europe	Latvia			Tutin, T. G. et al., eds. 1964-1980. Flora europaea.
1	Native	Europe	Eastern Europe	Lithuania			Tutin, T. G. et al., eds. 1964-1980. Flora europaea.
1	Native	Europe	Eastern Europe	Moldova			Tutin, T. G. et al., eds. 1964-1980. Flora europaea.
1	Native	Europe	Eastern Europe	Russian Federation-European part	European part		Komarov, V. L. et al., eds. 1934-1964. Flora SSSR.
1	Native	Europe	Eastern Europe	Ukraine		incl. Krym	Tutin, T. G. et al., eds. 1964-1980. Flora europaea.
1	Native	Europe	Middle Europe	Austria			Tutin, T. G. et al., eds. 1964-1980. Flora europaea.
1	Native	Europe	Middle Europe	Belgium			Tutin, T. G. et al., eds. 1964-1980. Flora europaea.
1	Native	Europe	Middle Europe	Czech Republic			Tutin, T. G. et al., eds. 1964-1980. Flora europaea.
1	Native	Europe	Middle Europe	Germany			Tutin, T. G. et al., eds. 1964-1980. Flora europaea.
1	Native	Europe	Middle Europe	Hungary			Tutin, T. G. et al., eds. 1964-1980. Flora europaea.
1	Native	Europe	Middle Europe	Netherlands			Tutin, T. G. et al., eds. 1964-1980. Flora europaea.
1	Native	Europe	Middle Europe	Poland			Tutin, T. G. et al., eds. 1964-1980. Flora europaea.
1	Native	Europe	Middle Europe	Slovakia			Tutin, T. G. et al., eds. 1964-1980. Flora europaea.
1	Native	Europe	Middle Europe	Switzerland			Tutin, T. G. et al., eds. 1964-1980. Flora europaea.
1	Native	Europe	Northern Europe	Denmark			Tutin, T. G. et al., eds. 1964-1980. Flora europaea.
1	Native	Europe	Northern Europe	Finland			Tutin, T. G. et al., eds. 1964-1980. Flora europaea.
1	Native	Europe	Northern Europe	Ireland			Tutin, T. G. et al., eds. 1964-1980. Flora europaea.
1	Native	Europe	Northern Europe	Sweden			Tutin, T. G. et al., eds. 1964-1980. Flora europaea.
1	Native	Europe	Northern Europe	United Kingdom			Tutin, T. G. et al., eds. 1964-1980. Flora europaea.
1	Native	Europe	Southeastern Europe	Albania			Tutin, T. G. et al., eds. 1964-1980. Flora europaea.
1	Native	Europe	Southeastern Europe	Bosnia and Herzegovina			Tutin, T. G. et al., eds. 1964-1980. Flora europaea.
1	Native	Europe	Southeastern Europe	Bulgaria			Tutin, T. G. et al., eds. 1964-1980. Flora europaea.
1	Native	Europe	Southeastern Europe	Croatia			Tutin, T. G. et al., eds. 1964-1980. Flora europaea.
1	Native	Europe	Southeastern Europe	Greece		incl. Crete	Tutin, T. G. et al., eds. 1964-1980. Flora europaea.
1	Native	Europe	Southeastern Europe	Italy		incl. Sardinia, Sicily	Tutin, T. G. et al., eds. 1964-1980. Flora europaea.
1	Native	Europe	Southeastern Europe	Montenegro			Tutin, T. G. et al., eds. 1964-1980. Flora europaea.
1	Native	Europe	Southeastern Europe	North Macedonia			Tutin, T. G. et al., eds. 1964-1980. Flora europaea.
1	Native	Europe	Southeastern Europe	Romania			Tutin, T. G. et al., eds. 1964-1980. Flora europaea.
1	Native	Europe	Southeastern Europe	Serbia			Tutin, T. G. et al., eds. 1964-1980. Flora europaea.
1	Native	Europe	Southeastern Europe	Slovenia			Tutin, T. G. et al., eds. 1964-1980. Flora europaea.
1	Native	Europe	Southwestern Europe	France		incl. Corsica	Tutin, T. G. et al., eds. 1964-1980. Flora europaea.
1	Native	Europe	Southwestern Europe	Spain		incl. Baleares	Tutin, T. G. et al., eds. 1964-1980. Flora europaea.
1	Native	Northern America	Eastern Canada	Canada	New Brunswick		Hinds, H. R. 2000. Flora of New Brunswick, ed. 2.
1	Native	Northern America	Eastern Canada	Canada	Nova Scotia		Scoggan, H. J. 1978-1979. The flora of Canada, 4 vol.
1	Native	Northern America	Eastern Canada	Canada	Ontario	s.e.	Scoggan, H. J. 1978-1979. The flora of Canada, 4 vol.
1	Native	Northern America	Eastern Canada	Canada	Prince Edward Island		Scoggan, H. J. 1978-1979. The flora of Canada, 4 vol.
1	Native	Northern America	Eastern Canada	Canada	Québec	s.w.	Scoggan, H. J. 1978-1979. The flora of Canada, 4 vol.
1	Native	Northern America	North-Central U.S.A.	United States	Illinois		Mohlenbrock, R. H. & D. M. Ladd. 1978. Distribution of Illinois vascular plants
1	Native	Northern America	North-Central U.S.A.	United States	Kansas		Cronquist, A. et al. 1972-. Intermountain flora.
1	Native	Northern America	North-Central U.S.A.	United States	Missouri		Steyermark, J. A. 1977. Flora of Missouri.
1	Native	Northern America	North-Central U.S.A.	United States	Oklahoma		Cronquist, A. et al. 1972-. Intermountain flora.
1	Native	Northern America	North-Central U.S.A.	United States	Wisconsin	s.e.	Wetter, M. A. et al. WISFLORA: Wisconsin vascular plant species (on-line resource).
1	Native	Northern America	Northeastern U.S.A.	United States	Connecticut		Magee, D. W. & H. E. Ahles. 1999. Flora of the Northeast. A manual of the vascular flora of New England and adjacent New York.
1	Native	Northern America	Northeastern U.S.A.	United States	Indiana		Deam, C. C. 1940. Flora of Indiana.
1	Native	Northern America	Northeastern U.S.A.	United States	Maine		Magee, D. W. & H. E. Ahles. 1999. Flora of the Northeast. A manual of the vascular flora of New England and adjacent New York.
1	Native	Northern America	Northeastern U.S.A.	United States	Massachusetts		Magee, D. W. & H. E. Ahles. 1999. Flora of the Northeast. A manual of the vascular flora of New England and adjacent New York.
1	Native	Northern America	Northeastern U.S.A.	United States	Michigan	s.	Voss, E. 1972-. Michigan flora.
1	Native	Northern America	Northeastern U.S.A.	United States	New Hampshire		Magee, D. W. & H. E. Ahles. 1999. Flora of the Northeast. A manual of the vascular flora of New England and adjacent New York.
1	Native	Northern America	Northeastern U.S.A.	United States	New Jersey		Gleason, H. A. & A. Cronquist. 1991. Manual of vascular plants of northeastern United States and adjacent Canada, ed. 2
1	Native	Northern America	Northeastern U.S.A.	United States	New York		Magee, D. W. & H. E. Ahles. 1999. Flora of the Northeast. A manual of the vascular flora of New England and adjacent New York.
1	Native	Northern America	Northeastern U.S.A.	United States	Ohio		Ohio Flora Committee (E. L. Braun, T. S. Cooperrider, T. R. Fisher, J. J. Furlow). 1967-. The vascular flora of Ohio.
1	Native	Northern America	Northeastern U.S.A.	United States	Pennsylvania		Rhoads, A. F. & T. A. Block. 2000. The plants of Pennsylvannia. An illustrated manual
1	Native	Northern America	Northeastern U.S.A.	United States	Rhode Island		Magee, D. W. & H. E. Ahles. 1999. Flora of the Northeast. A manual of the vascular flora of New England and adjacent New York.
1	Native	Northern America	Northeastern U.S.A.	United States	Vermont		Magee, D. W. & H. E. Ahles. 1999. Flora of the Northeast. A manual of the vascular flora of New England and adjacent New York.
1	Native	Northern America	Northeastern U.S.A.	United States	West Virginia		Harmon, P. J. et al. 2006. Checklist and atlas of the vascular flora of West Virginia.
1	Native	Northern America	Northern Mexico	Mexico	Baja California (Norte)		Gómez-Pompa, A. & V. Sosa, eds. 1978-. Flora de Veracruz.
1	Native	Northern America	Northern Mexico	Mexico	Chihuahua		Gómez-Pompa, A. & V. Sosa, eds. 1978-. Flora de Veracruz.
1	Native	Northern America	Northern Mexico	Mexico	Coahuila de Zaragoza		Gómez-Pompa, A. & V. Sosa, eds. 1978-. Flora de Veracruz.
1	Native	Northern America	Northern Mexico	Mexico	San Luis Potosí		Gómez-Pompa, A. & V. Sosa, eds. 1978-. Flora de Veracruz.
1	Native	Northern America	Northern Mexico	Mexico	Tamaulipas		Gómez-Pompa, A. & V. Sosa, eds. 1978-. Flora de Veracruz.
1	Native	Northern America	South-Central U.S.A.	United States	New Mexico		Cronquist, A. et al. 1972-. Intermountain flora.
1	Native	Northern America	South-Central U.S.A.	United States	Texas		Cronquist, A. et al. 1972-. Intermountain flora.
1	Native	Northern America	Southeastern U.S.A.	United States	Alabama		Radford, A. E. et al. 1964. Manual of the vascular flora of the Carolinas.
1	Native	Northern America	Southeastern U.S.A.	United States	Arkansas		Smith, E. B. 1978. An atlas and annotated list of the vascular plants of Arkansas
1	Native	Northern America	Southeastern U.S.A.	United States	Delaware		Gleason, H. A. & A. Cronquist. 1991. Manual of vascular plants of northeastern United States and adjacent Canada, ed. 2
1	Native	Northern America	Southeastern U.S.A.	United States	Florida		Radford, A. E. et al. 1964. Manual of the vascular flora of the Carolinas.
1	Native	Northern America	Southeastern U.S.A.	United States	Georgia		Radford, A. E. et al. 1964. Manual of the vascular flora of the Carolinas.
1	Native	Northern America	Southeastern U.S.A.	United States	Kentucky		Jones, R. L. 2005. Plant life of Kentucky: an illustrated guide to the vascular flora.
1	Native	Northern America	Southeastern U.S.A.	United States	Louisiana		Radford, A. E. et al. 1964. Manual of the vascular flora of the Carolinas.
1	Native	Northern America	Southeastern U.S.A.	United States	Maryland		Brown, R. G. & M. L. Brown. 1984. Herbaceous plants of Maryland
1	Native	Northern America	Southeastern U.S.A.	United States	Mississippi		Radford, A. E. et al. 1964. Manual of the vascular flora of the Carolinas.
1	Native	Northern America	Southeastern U.S.A.	United States	North Carolina		Radford, A. E. et al. 1964. Manual of the vascular flora of the Carolinas.
1	Native	Northern America	Southeastern U.S.A.	United States	South Carolina		Radford, A. E. et al. 1964. Manual of the vascular flora of the Carolinas.
1	Native	Northern America	Southeastern U.S.A.	United States	Tennessee		Chester, E. W. et al. 1993-1997. Atlas of Tennessee vascular plants.
1	Native	Northern America	Southeastern U.S.A.	United States	Virginia		Radford, A. E. et al. 1964. Manual of the vascular flora of the Carolinas.
1	Native	Northern America	Southern Mexico	Mexico	Chiapas		Gómez-Pompa, A. & V. Sosa, eds. 1978-. Flora de Veracruz.
1	Native	Northern America	Southern Mexico	Mexico	Guanajuato		Gómez-Pompa, A. & V. Sosa, eds. 1978-. Flora de Veracruz.
1	Native	Northern America	Southern Mexico	Mexico	Hidalgo		Gómez-Pompa, A. & V. Sosa, eds. 1978-. Flora de Veracruz.
1	Native	Northern America	Southern Mexico	Mexico	Veracruz de Ignacio de la Llave		Gómez-Pompa, A. & V. Sosa, eds. 1978-. Flora de Veracruz.
1	Native	Northern America	Southwestern U.S.A.	United States	Arizona		Cronquist, A. et al. 1972-. Intermountain flora.
1	Native	Northern America	Southwestern U.S.A.	United States	California		Hickman, J. C., ed. 1993. The Jepson manual: higher plants of California
1	Native	Northern America	Southwestern U.S.A.	United States	Nevada		Cronquist, A. et al. 1972-. Intermountain flora.
1	Native	Northern America	Southwestern U.S.A.	United States	Utah		Cronquist, A. et al. 1972-. Intermountain flora.
1	Native	Southern America	Brazil	Brazil			Zuloaga, F. O. & O. Morrone, eds. 1996, 1999. Catálogo de las plantas vasculares de la República Argentina. I. Pteridophyta, Gymnospermae y Angiospermae (Monocotyledonae), II. Dicotyledonae. Monogr. Syst. Bot. Missouri Bot. Gard. 60, 74.
1	Native	Southern America	Caribbean	Cuba			Liogier, H. A. 1984-. Descriptive flora of Puerto Rico and adjacent islands.
1	Native	Southern America	Caribbean	Dominican Republic			Liogier, H. A. 1981-. Flora of Hispaniola: Part 1; La Flora de la Española. Part II-.
1	Native	Southern America	Caribbean	Haiti			Liogier, H. A. 1981-. Flora of Hispaniola: Part 1; La Flora de la Española. Part II-.
1	Native	Southern America	Caribbean	United States	Puerto Rico		Liogier, H. A. 1984-. Descriptive flora of Puerto Rico and adjacent islands.
1	Native	Southern America	Southern South America	Argentina			Zuloaga, F. O. & O. Morrone, eds. 1996, 1999. Catálogo de las plantas vasculares de la República Argentina. I. Pteridophyta, Gymnospermae y Angiospermae (Monocotyledonae), II. Dicotyledonae. Monogr. Syst. Bot. Missouri Bot. Gard. 60, 74.
1	Native	Southern America	Southern South America	Chile			Zuloaga, F. O. & O. Morrone, eds. 1996, 1999. Catálogo de las plantas vasculares de la República Argentina. I. Pteridophyta, Gymnospermae y Angiospermae (Monocotyledonae), II. Dicotyledonae. Monogr. Syst. Bot. Missouri Bot. Gard. 60, 74.
1	Native	Southern America	Southern South America	Paraguay			Zuloaga, F. O. & O. Morrone, eds. 1996, 1999. Catálogo de las plantas vasculares de la República Argentina. I. Pteridophyta, Gymnospermae y Angiospermae (Monocotyledonae), II. Dicotyledonae. Monogr. Syst. Bot. Missouri Bot. Gard. 60, 74.
1	Native	Southern America	Southern South America	Uruguay			Zuloaga, F. O. & O. Morrone, eds. 1996, 1999. Catálogo de las plantas vasculares de la República Argentina. I. Pteridophyta, Gymnospermae y Angiospermae (Monocotyledonae), II. Dicotyledonae. Monogr. Syst. Bot. Missouri Bot. Gard. 60, 74.
1	Native	Southern America	Western South America	Bolivia			Zuloaga, F. O. & O. Morrone, eds. 1996, 1999. Catálogo de las plantas vasculares de la República Argentina. I. Pteridophyta, Gymnospermae y Angiospermae (Monocotyledonae), II. Dicotyledonae. Monogr. Syst. Bot. Missouri Bot. Gard. 60, 74.
1	Native	Southern America	Western South America	Ecuador			Jørgensen, P. M. & S. León-Yánez, eds. 1999. Catalogue of the vascular plants of Ecuador. Monogr. Syst. Bot. Missouri Bot. Gard. 75
1	Native	Southern America	Western South America	Peru			Brako, L. & J. L. Zarucchi. 1993. Catalogue of the flowering plants and gymnosperms of Peru. Monogr. Syst. Bot. Missouri Bot. Gard. 45

Native
Africa

NORTHERN AFRICA: Algeria, Egypt, Libya, Morocco, Tunisia
NORTHEAST TROPICAL AFRICA: Eritrea, Ethiopia, Somalia
EAST TROPICAL AFRICA: Kenya
WEST-CENTRAL TROPICAL AFRICA: Democratic Republic of the Congo
WEST TROPICAL AFRICA: Nigeria
SOUTH TROPICAL AFRICA: Mozambique, Zambia, Zimbabwe
SOUTHERN AFRICA: Botswana, Namibia, South Africa [Cape Province, KwaZulu-Natal, Transvaal]

Asia-Temperate

ARABIAN PENINSULA: Oman, Saudi Arabia, Yemen
WESTERN ASIA: Afghanistan, Cyprus, Egypt [Sinai], Iran, Israel, Jordan, Lebanon, Syria, Turkey
CAUCASUS: Russian Federation-Ciscaucasia [Ciscaucasia], Armenia, Azerbaijan, Georgia, Russian Federation [Dagestan]
MIDDLE ASIA: Kyrgyzstan, Tajikistan, Turkmenistan, Uzbekistan
CHINA: China [Guangdong Sheng, Guangxi Zhuangzu Zizhiqu, Guizhou Sheng, Hunan Sheng, Yunnan Sheng]

Asia-Tropical

INDIAN SUBCONTINENT: India, Pakistan

Europe

NORTHERN EUROPE: Denmark, Finland, United Kingdom, Ireland, Sweden
MIDDLE EUROPE: Austria, Belgium, Switzerland, Czech Republic, Germany, Hungary, Netherlands, Poland, Slovakia
EASTERN EUROPE: Russian Federation-European part [European part], Belarus, Estonia, Lithuania, Latvia, Moldova, Ukraine (incl. Krym)
SOUTHEASTERN EUROPE: Albania, Bulgaria, Bosnia and Herzegovina, Greece (incl. Crete), Croatia, Italy (incl. Sardinia, Sicily), North Macedonia, Montenegro, Romania, Serbia, Slovenia
SOUTHWESTERN EUROPE: Spain (incl. Baleares), France (incl. Corsica)

Northern America

EASTERN CANADA: Canada [Québec (s.w.), Nova Scotia, Ontario (s.e.), Prince Edward Island, New Brunswick]
NORTHEASTERN U.S.A.: United States [Connecticut, Indiana, Maine, Massachusetts, Michigan (s.), New Hampshire, New Jersey, New York, Ohio, Pennsylvania, Rhode Island, Vermont, West Virginia]
NORTH-CENTRAL U.S.A.: United States [Illinois, Kansas, Missouri, Oklahoma, Wisconsin (s.e.)]
SOUTHEASTERN U.S.A.: United States [Alabama, Arkansas, Delaware, Florida, Georgia, Kentucky, Louisiana, Maryland, Mississippi, North Carolina, South Carolina, Tennessee, Virginia]
SOUTH-CENTRAL U.S.A.: United States [New Mexico, Texas]
SOUTHWESTERN U.S.A.: United States [Arizona, California, Nevada, Utah]
NORTHERN MEXICO: Mexico [Baja California (Norte), Chihuahua, Coahuila de Zaragoza, San Luis Potosí, Tamaulipas]
SOUTHERN MEXICO: Mexico [Chiapas, Guanajuato, Hidalgo, Veracruz de Ignacio de la Llave]

Southern America

CARIBBEAN: Cuba, Dominican Republic, Haiti, United States [Puerto Rico]
BRAZIL: Brazil
WESTERN SOUTH AMERICA: Bolivia, Ecuador, Peru
SOUTHERN SOUTH AMERICA: Argentina, Chile, Paraguay, Uruguay

Reference(s)

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Cite as: USDA, Agricultural Research Service, National Plant Germplasm System. 2026. Germplasm Resources Information Network (GRIN Taxonomy). National Germplasm Resources Laboratory, Beltsville, Maryland.
URL: https://npgstest.ars-grin.gov/gringlobal/taxon/taxonomydetail?id=459207. Accessed 13 May 2026.