Taxon:  Physalis grisea (Waterf.) M. Martínez



Summary
Genus:
Physalis
Family:
Solanaceae
Subfamily:
Solanoideae
Tribe:
Physaleae
Subtribe:
Physalinae
Nomen number:
318474
Place of publication:
Taxon 42:104.  1993
Verified:
10/02/1995 ARS Systematic Botanists.
Accessions:
1 (1  active, 0  available) in National Plant Germplasm System.  (Map it)
Other conspecific taxa
Autonyms (not in current use), synonyms and invalid designations
Basionym
Physalis pubescens L. var. grisea Waterf.

Invalid Designation(s)
Physalis pruinosa auct. N. Amer.
No images


Conservation Status

Reference(s)
  1. Aldén, B., S. Ryman, & M. Hjertson. 2012. Svensk Kulturväxtdatabas, SKUD (Swedish Cultivated and Utility Plants Database; online resource)
  2. Chester, E. W. et al. 1993-1997. Atlas of Tennessee vascular plants. Note: = Physalis pruinosa auctt. non L.
  3. Dale, S. M. et al. 2024. Engineering the future of Physalis grisea: A focus on agricultural challenges, model species status, and applied improvements. Plants, People, Planet 1-12. DOI: 10.1002/ppp3.10536.
  4. Duncan, W. H. & J. T. Kartesz. 1981. Vascular flora of Georgia: an annotated checklist. Note: = Physalis pubescens var. grisea Waterfall
  5. Gleason, H. A. & A. Cronquist. 1991. Manual of vascular plants of northeastern United States and adjacent Canada, ed. 2 Note: = Physalis pubescens var. grisea Waterfall
  6. He, J. et al. 2023. Establishing Physalis as a Solanaceae model system enables genetic reevaluation of the inflated calyx syndrome. Pl. Cell 35:351-368. DOI: 10.1093/plcell/koac305 . Note: assembled chromosome-scale genome of 1.37 Gb in size
  7. Hickman, J. C., ed. 1993. The Jepson manual: higher plants of California Note: = Physalis pubescens var. grisea Waterfall
  8. Liberty Hyde Bailey Hortorium. 1976. Hortus third. A concise dictionary of plants cultivated in the United States and Canada Note: = Physalis pruinosa
  9. Magee, D. W. & H. E. Ahles. 1999. Flora of the Northeast. A manual of the vascular flora of New England and adjacent New York. Note: = Physalis pruinosa
  10. Markle, G. M. et al., eds. 1998. Food and feed crops of the United States, ed. 2
  11. Martínez, M. 1993. The correct application of Physalis pruinosa L. (Solanaceae). Taxon 42:103-104. DOI: 10.2307/1223312.
  12. Porcher, M. H. et al. Searchable World Wide Web Multilingual Multiscript Plant Name Database (MMPND) (on-line resource).
  13. Pretz, C. & R. Deanna. 2020. Typifications and nomenclatural notes in Physalis (Solanaceae) from the United States. Taxon 69:170-192. DOI: 10.1002/tax.12159.
  14. Radford, A. E. et al. 1964. Manual of the vascular flora of the Carolinas. Note: = Physalis pubescens var. grisea Waterfall
  15. Rehm, S. 1994. Multilingual dictionary of agronomic plants Note: = Physalis pruinosa
  16. Scoggan, H. J. 1978-1979. The flora of Canada, 4 vol. Note: = Physalis pubescens var. grisea Waterfall
  17. Steyermark, J. A. 1977. Flora of Missouri. Note: = Physalis pubescens var. grisea Waterfall
  18. Strausbaugh, T. D. & E. L. Core. 1978. Flora of West Virginia, ed. 2. Note: = Physalis pruinosa
  19. Sullivan, J. R. 2004. The genus Physalis (Solanaceae) in the southeastern United States. Rhodora 106:305-326.
  20. Ward, D. B. 2008. Keys to the flora of Florida: 19, Physalis (Solanaceae). Phytologia 90:198-207.
  21. Waterfall, U. T. 1958. A taxonomic study of the genus Physalis in North America north of Mexico. Rhodora 60:167. Note: = Physalis pubescens var. grisea Waterfall
  22. Whitson, M. & P. S. Manos. 2005. Untangling Physalis (Solanaceae) from the Physaloids: a two-gene phylogeny of the Physalinae. Syst. Bot. 30:216-230. Note: it used an accession (NIJ 894750256) from Radboud University and Botanical Garden that corresponds instead to P. pruinosa
Common names
LanguageNameAlternate namenoteseqCitation
Englishdwarf Cape-gooseberry as Physalis pruinosa1Liberty Hyde Bailey Hortorium. 1976. Hortus third. A concise dictionary of plants cultivated in the United States and Canada Note: = Physalis pruinosa
Englishhairy ground-cherry  1Rehm, S. 1994. Multilingual dictionary of agronomic plants Note: = Physalis pruinosa
Englishstrawberry-tomato  1Rehm, S. 1994. Multilingual dictionary of agronomic plants Note: = Physalis pruinosa
GermanErdkirsche as Physalis pruinosa2Rehm, S. 1994. Multilingual dictionary of agronomic plants Note: = Physalis pruinosa
Spanishmuyaca as Physalis pruinosa2Rehm, S. 1994. Multilingual dictionary of agronomic plants Note: = Physalis pruinosa
Spanishuvilla as Physalis pruinosa2Rehm, S. 1994. Multilingual dictionary of agronomic plants Note: = Physalis pruinosa
Swedishgyllenbär  2Aldén, B., S. Ryman, & M. Hjertson. 2012. Svensk Kulturväxtdatabas, SKUD (Swedish Cultivated and Utility Plants Database; online resource)
Distribution
order_codeStatusContinentSubcontinentCountryStateNoteReference
1NativeNorthern AmericaNortheastern U.S.A.United StatesConnecticut Magee, D. W. & H. E. Ahles. 1999. Flora of the Northeast. A manual of the vascular flora of New England and adjacent New York. Note: = Physalis pruinosa
1NativeNorthern AmericaNortheastern U.S.A.United StatesMaine Magee, D. W. & H. E. Ahles. 1999. Flora of the Northeast. A manual of the vascular flora of New England and adjacent New York. Note: = Physalis pruinosa
1NativeNorthern AmericaNortheastern U.S.A.United StatesMassachusetts Magee, D. W. & H. E. Ahles. 1999. Flora of the Northeast. A manual of the vascular flora of New England and adjacent New York. Note: = Physalis pruinosa
1NativeNorthern AmericaNortheastern U.S.A.United StatesNew Jersey Gleason, H. A. & A. Cronquist. 1991. Manual of vascular plants of northeastern United States and adjacent Canada, ed. 2 Note: = Physalis pubescens var. grisea Waterfall
1NativeNorthern AmericaNortheastern U.S.A.United StatesNew York Gleason, H. A. & A. Cronquist. 1991. Manual of vascular plants of northeastern United States and adjacent Canada, ed. 2 Note: = Physalis pubescens var. grisea Waterfall
1NativeNorthern AmericaNortheastern U.S.A.United StatesPennsylvania Gleason, H. A. & A. Cronquist. 1991. Manual of vascular plants of northeastern United States and adjacent Canada, ed. 2 Note: = Physalis pubescens var. grisea Waterfall
1NativeNorthern AmericaNortheastern U.S.A.United StatesVermont Magee, D. W. & H. E. Ahles. 1999. Flora of the Northeast. A manual of the vascular flora of New England and adjacent New York. Note: = Physalis pruinosa
1NativeNorthern AmericaNortheastern U.S.A.United StatesWest Virginia Strausbaugh, T. D. & E. L. Core. 1978. Flora of West Virginia, ed. 2. Note: = Physalis pruinosa
1NativeNorthern AmericaSoutheastern U.S.A.United StatesAlabama Gleason, H. A. & A. Cronquist. 1991. Manual of vascular plants of northeastern United States and adjacent Canada, ed. 2 Note: = Physalis pubescens var. grisea Waterfall
1NativeNorthern AmericaSoutheastern U.S.A.United StatesDelaware Gleason, H. A. & A. Cronquist. 1991. Manual of vascular plants of northeastern United States and adjacent Canada, ed. 2 Note: = Physalis pubescens var. grisea Waterfall
1NativeNorthern AmericaSoutheastern U.S.A.United StatesGeorgia Duncan, W. H. & J. T. Kartesz. 1981. Vascular flora of Georgia: an annotated checklist. Note: = Physalis pubescens var. grisea Waterfall
1NativeNorthern AmericaSoutheastern U.S.A.United StatesKentucky Gleason, H. A. & A. Cronquist. 1991. Manual of vascular plants of northeastern United States and adjacent Canada, ed. 2 Note: = Physalis pubescens var. grisea Waterfall
1NativeNorthern AmericaSoutheastern U.S.A.United StatesMaryland Gleason, H. A. & A. Cronquist. 1991. Manual of vascular plants of northeastern United States and adjacent Canada, ed. 2 Note: = Physalis pubescens var. grisea Waterfall
1NativeNorthern AmericaSoutheastern U.S.A.United StatesNorth Carolinaw. & c.Radford, A. E. et al. 1964. Manual of the vascular flora of the Carolinas. Note: = Physalis pubescens var. grisea Waterfall
1NativeNorthern AmericaSoutheastern U.S.A.United StatesSouth Carolina Radford, A. E. et al. 1964. Manual of the vascular flora of the Carolinas. Note: = Physalis pubescens var. grisea Waterfall
1NativeNorthern AmericaSoutheastern U.S.A.United StatesTennessee Chester, E. W. et al. 1993-1997. Atlas of Tennessee vascular plants. Note: = Physalis pruinosa auctt. non L.
1NativeNorthern AmericaSoutheastern U.S.A.United StatesVirginia Gleason, H. A. & A. Cronquist. 1991. Manual of vascular plants of northeastern United States and adjacent Canada, ed. 2 Note: = Physalis pubescens var. grisea Waterfall
4Naturalized    natzd. elsewhere 
Native
Northern America
  • NORTHEASTERN U.S.A.: United States [Connecticut, Maine, Massachusetts, New Jersey, New York, Pennsylvania, Vermont, West Virginia]
  • SOUTHEASTERN U.S.A.: United States [Alabama, Delaware, Georgia, Kentucky, Maryland, North Carolina (w. & c.), South Carolina, Tennessee, Virginia]
Naturalized (natzd. elsewhere)

Reference(s)
  1. Chester, E. W. et al. 1993-1997. Atlas of Tennessee vascular plants. Note: = Physalis pruinosa auctt. non L.
  2. Duncan, W. H. & J. T. Kartesz. 1981. Vascular flora of Georgia: an annotated checklist. Note: = Physalis pubescens var. grisea Waterfall
  3. Gleason, H. A. & A. Cronquist. 1991. Manual of vascular plants of northeastern United States and adjacent Canada, ed. 2 Note: = Physalis pubescens var. grisea Waterfall
  4. Magee, D. W. & H. E. Ahles. 1999. Flora of the Northeast. A manual of the vascular flora of New England and adjacent New York. Note: = Physalis pruinosa
  5. Radford, A. E. et al. 1964. Manual of the vascular flora of the Carolinas. Note: = Physalis pubescens var. grisea Waterfall
  6. Strausbaugh, T. D. & E. L. Core. 1978. Flora of West Virginia, ed. 2. Note: = Physalis pruinosa
Economic Uses
UsageTypeNoteReference
Human foodfruit Martínez, M. 1993. The correct application of Physalis pruinosa L. (Solanaceae). Taxon 42:104.
Human foodfruitof unknown domestication status; locally cultivatedDale, S. M. et al. 2024. Engineering the future of Physalis grisea: A focus on agricultural challenges, model species status, and applied improvements. Plants, People, Planet 1-12. DOI: 10.1002/ppp3.10536.
Weed of cultivated fields in FloridaWard, D. B. 2008. Keys to the flora of Florida: 19, Physalis (Solanaceae). Phytologia 90:198-207.
Cite as: USDA, Agricultural Research Service, National Plant Germplasm System. 2026. Germplasm Resources Information Network (GRIN Taxonomy). National Germplasm Resources Laboratory, Beltsville, Maryland.
URL: https://npgstest.ars-grin.gov/gringlobal/taxon/taxonomydetail?id=318474. Accessed 11 May 2026.