Taxon: Hordeum marinum Huds. subsp. gussoneanum (Parl.) Thell.

Summary

Genus:

Hordeum

Subgenus:

Hordeastrum

Section:

Marina

Family:

Poaceae (alt. Gramineae)

Subfamily:

Pooideae

Tribe:

Triticeae

Nomen number:

102508

Place of publication:

Vierteljahrsschr. Naturf. Ges. Zürich 52:441. 1908

Verified:

11/05/2007 ARS Systematic Botanists.

Accessions:

19 (14 active, 14 available) in National Plant Germplasm System. (Map it)

Other conspecific taxa

Hordeum marinum Huds. (8 active accession[s])
Hordeum marinum Huds. subsp. marinum (1 active accession[s])

Autonyms (not in current use), synonyms and invalid designations

Basionym
Hordeum gussoneanum Parl.

Heterotypic Synonym(s)
Critesion hystrix (Roth) Á. Löve
Hordeum geniculatum All.
Hordeum hystrix Roth

Images (2 total. Click on image for more.)

Conservation Status

Reference(s)

University of Georgia Center for Invasive Species and Ecosystem Health
Aldén, B., S. Ryman, & M. Hjertson. 2012. Svensk Kulturväxtdatabas, SKUD (Swedish Cultivated and Utility Plants Database; online resource)
Blattner, F. R. 2004. Phylogenetic analysis of Hordeum (Poaceae) as inferred by nuclear rDNA ITS sequences. Molec. Phylogenet. Evol. 33:289-299.
Bothmer, R. von et al. 1989. Variation and differentiation in Hordeum marinum (Poaceae). Nordic J. Bot. 9:8.
Cronquist, A. et al. 1972-. Intermountain flora. Note: = Hordeum geniculatum All.
Cuadrado, Á. et al. 2017. On the allopolyploid origin and genome structure of the closely related species Hordeum secalinum and Hordeum capense inferred by molecular karyotyping. Ann. Bot. (Oxford) 120:245-255. DOI: 10.1093/aob/mcw270. Note: donor of Xa genome to allotetraploid Hordeum secalinum
Davis, P. H., ed. 1965-1988. Flora of Turkey and the east Aegean islands. Note: = Hordeum geniculatum All.
Doebley, J. et al. 1992. Chloroplast DNA variation and the phylogeny of Hordeum (Poaceae). Amer. J. Bot. 79:576-584.
FNA Editorial Committee. 1993-. Flora of North America.
George, A. S., ed. 1980-. Flora of Australia. Note: = Hordeum hystrix Roth
Jakob, S. S. & F. R. Blattner. 2006. A chloroplast genealogy of Hordeum (Poaceae): long-term persisting haplotypes, incomplete lineage sorting, regional extinction, and the consequences for phylogenetic inference. Molec. Biol. Evol. 23:1602-1612.
Jakob, S. S. et al. 2007. Combined ecological niche modelling and molecular phylogeography revealed the evolutionary history of Hordeum marinum (Poaceae)-niche differentiation, loss of genetic diversity, and speciation in Mediterranean Quaternary refugia. Molec. Ecol. 16:1713-1727.
Komatsuda, T. et al. 1999. Phylogeny in the genus Hordeum based on nucleotide sequences closely linked to the vrs1 locus (row number of spikelets). Genome 42:973-981.
Meikle, R. D. 1977-1985. Flora of Cyprus. Note: = Hordeum geniculatum All.
Nasir, E. & S. I. Ali, eds. 1970-. Flora of [West] Pakistan.
Nishikawa, T. et al. 2002. Molecular phylogeny of the genus Hordeum using three chloroplast DNA sequences. Genome 45:1157-1166.
Petersen, G. & O. Seberg. 2003. Phylogenetic analysis of the diploid species of Hordeum (Poaceae) and a revised classification of the genus. Syst. Bot. 28:293-306. www.aspt.net/systematic-botany
Sahebi, J. et al. 2004. A taxonomic review of the Hordeum marinum sensu lato (Poaceae: Triticeae) in Iran. Iran. J. Sci. Technol., Trans. A. Sci. 28:127-135.
Skvortsov, A. K., ed. 2006-. Flora of the Lower Volga region Note: treats as Hordeum marinum Huds.
Soreng, R. J. et al. 2003. Catalogue of New World grasses (Poaceae): IV. Subfamily Pooideae. Contr. U.S. Natl. Herb.
Townsend, C. C. & E. Guest. 1966-. Flora of Iraq. Note: = Hordeum geniculatum All.
Tutin, T. G. et al., eds. 1964-1980. Flora europaea. Note: = Hordeum hystrix Roth
Tzvelev, N. N. 1976. Zlaki SSSR

Common names

Language	Name	seq	Citation
English	geniculate barley	1	FNA Editorial Committee. 1993-. Flora of North America.
English	Mediterranean barley	1	FNA Editorial Committee. 1993-. Flora of North America.
Swedish	ullkorn	2	Aldén, B., S. Ryman, & M. Hjertson. 2012. Svensk Kulturväxtdatabas, SKUD (Swedish Cultivated and Utility Plants Database; online resource)

Distribution

order_code	Status	Continent	Subcontinent	Country	State	Note	Reference
1	Native	Africa	Northern Africa	Algeria		n.	Tutin, T. G. et al., eds. 1964-1980. Flora europaea. Note: = Hordeum hystrix Roth
1	Native	Africa	Northern Africa	Egypt		n.	Tutin, T. G. et al., eds. 1964-1980. Flora europaea. Note: = Hordeum hystrix Roth
1	Native	Africa	Northern Africa	Libya		n.	Sahebi, J. et al. 2004. A taxonomic review of the Hordeum marinum sensu lato (Poaceae: Triticeae) in Iran. Iran. J. Sci. Technol., Trans. A. Sci. 28:127-135.
1	Native	Africa	Northern Africa	Morocco			Jakob, S. S. et al. 2007. Combined ecological niche modelling and molecular phylogeography revealed the evolutionary history of Hordeum marinum (Poaceae)-niche differentiation, loss of genetic diversity, and speciation in Mediterranean Quaternary refugia. Molec. Ecol. 16:1713-1727.
1	Native	Africa	Northern Africa	Tunisia			Sahebi, J. et al. 2004. A taxonomic review of the Hordeum marinum sensu lato (Poaceae: Triticeae) in Iran. Iran. J. Sci. Technol., Trans. A. Sci. 28:127-135.
1	Native	Asia-Temperate	Caucasus	Azerbaijan			Tzvelev, N. N. 1976. Zlaki SSSR
1	Native	Asia-Temperate	Caucasus	Georgia			Tzvelev, N. N. 1976. Zlaki SSSR
1	Native	Asia-Temperate	Caucasus	Russian Federation	Dagestan		Tzvelev, N. N. 1976. Zlaki SSSR
1	Native	Asia-Temperate	Caucasus	Russian Federation-Ciscaucasia	Ciscaucasia	w.	Tzvelev, N. N. 1976. Zlaki SSSR
1	Native	Asia-Temperate	Middle Asia	Kazakhstan		s.w.	Tzvelev, N. N. 1976. Zlaki SSSR
1	Native	Asia-Temperate	Middle Asia	Tajikistan			Tzvelev, N. N. 1976. Zlaki SSSR
1	Native	Asia-Temperate	Middle Asia	Turkmenistan			Tzvelev, N. N. 1976. Zlaki SSSR
1	Native	Asia-Temperate	Middle Asia	Uzbekistan			Tzvelev, N. N. 1976. Zlaki SSSR
1	Native	Asia-Temperate	Western Asia	Afghanistan			Nishikawa, T. et al. 2002. Molecular phylogeny of the genus Hordeum using three chloroplast DNA sequences. Genome 45:1157-1166.
1	Native	Asia-Temperate	Western Asia	Cyprus			Meikle, R. D. 1977-1985. Flora of Cyprus. Note: = Hordeum geniculatum All.
1	Native	Asia-Temperate	Western Asia	Iran			Sahebi, J. et al. 2004. A taxonomic review of the Hordeum marinum sensu lato (Poaceae: Triticeae) in Iran. Iran. J. Sci. Technol., Trans. A. Sci. 28:127-135.
1	Native	Asia-Temperate	Western Asia	Iraq			Townsend, C. C. & E. Guest. 1966-. Flora of Iraq. Note: = Hordeum geniculatum All.
1	Native	Asia-Temperate	Western Asia	Israel			Euro+Med Editorial Committee. Euro+Med Plantbase: the information resource for Euro-Mediterranean plant diversity (on-line resource).
1	Native	Asia-Temperate	Western Asia	Jordan			Euro+Med Editorial Committee. Euro+Med Plantbase: the information resource for Euro-Mediterranean plant diversity (on-line resource).
1	Native	Asia-Temperate	Western Asia	Lebanon			Sahebi, J. et al. 2004. A taxonomic review of the Hordeum marinum sensu lato (Poaceae: Triticeae) in Iran. Iran. J. Sci. Technol., Trans. A. Sci. 28:127-135.
1	Native	Asia-Temperate	Western Asia	Syria			Sahebi, J. et al. 2004. A taxonomic review of the Hordeum marinum sensu lato (Poaceae: Triticeae) in Iran. Iran. J. Sci. Technol., Trans. A. Sci. 28:127-135.
1	Native	Asia-Temperate	Western Asia	Turkey			Davis, P. H., ed. 1965-1988. Flora of Turkey and the east Aegean islands. Note: = Hordeum geniculatum All.
1	Native	Asia-Tropical	Indian Subcontinent	Pakistan		n.	Nasir, E. & S. I. Ali, eds. 1970-. Flora of [West] Pakistan.
1	Native	Europe	Eastern Europe	Moldova			Tzvelev, N. N. 1976. Zlaki SSSR
1	Native	Europe	Eastern Europe	Russian Federation	Astrakhan		Skvortsov, A. K., ed. 2006-. Flora of the Lower Volga region Note: treats as Hordeum marinum Huds.
1	Native	Europe	Eastern Europe	Russian Federation	Volgogradskaja oblast'		Skvortsov, A. K., ed. 2006-. Flora of the Lower Volga region Note: treats as Hordeum marinum Huds.
1	Native	Europe	Eastern Europe	Ukraine		incl. Krym	Tzvelev, N. N. 1976. Zlaki SSSR
1	Native	Europe	Middle Europe	Austria			Tutin, T. G. et al., eds. 1964-1980. Flora europaea. Note: = Hordeum hystrix Roth
1	Native	Europe	Middle Europe	Czech Republic			Tutin, T. G. et al., eds. 1964-1980. Flora europaea. Note: = Hordeum hystrix Roth
1	Native	Europe	Middle Europe	Hungary			Tutin, T. G. et al., eds. 1964-1980. Flora europaea. Note: = Hordeum hystrix Roth
1	Native	Europe	Middle Europe	Slovakia			Euro+Med Editorial Committee. Euro+Med Plantbase: the information resource for Euro-Mediterranean plant diversity (on-line resource).
1	Native	Europe	Southeastern Europe	Albania			Tutin, T. G. et al., eds. 1964-1980. Flora europaea. Note: = Hordeum hystrix Roth
1	Native	Europe	Southeastern Europe	Bulgaria			Tutin, T. G. et al., eds. 1964-1980. Flora europaea. Note: = Hordeum hystrix Roth
1	Native	Europe	Southeastern Europe	Croatia			Euro+Med Editorial Committee. Euro+Med Plantbase: the information resource for Euro-Mediterranean plant diversity (on-line resource).
1	Native	Europe	Southeastern Europe	Greece		incl. Crete	Tutin, T. G. et al., eds. 1964-1980. Flora europaea. Note: = Hordeum hystrix Roth
1	Native	Europe	Southeastern Europe	Italy		incl. Sardinia, Sicily	Tutin, T. G. et al., eds. 1964-1980. Flora europaea. Note: = Hordeum hystrix Roth
1	Native	Europe	Southeastern Europe	Montenegro			Euro+Med Editorial Committee. Euro+Med Plantbase: the information resource for Euro-Mediterranean plant diversity (on-line resource).
1	Native	Europe	Southeastern Europe	North Macedonia			Euro+Med Editorial Committee. Euro+Med Plantbase: the information resource for Euro-Mediterranean plant diversity (on-line resource).
1	Native	Europe	Southeastern Europe	Romania			Tutin, T. G. et al., eds. 1964-1980. Flora europaea. Note: = Hordeum hystrix Roth
1	Native	Europe	Southeastern Europe	Serbia			Euro+Med Editorial Committee. Euro+Med Plantbase: the information resource for Euro-Mediterranean plant diversity (on-line resource).
1	Native	Europe	Southeastern Europe	Slovenia			Euro+Med Editorial Committee. Euro+Med Plantbase: the information resource for Euro-Mediterranean plant diversity (on-line resource).
1	Native	Europe	Southwestern Europe	France		s. & Corsica	Euro+Med Editorial Committee. Euro+Med Plantbase: the information resource for Euro-Mediterranean plant diversity (on-line resource).
1	Native	Europe	Southwestern Europe	Portugal			Tutin, T. G. et al., eds. 1964-1980. Flora europaea. Note: = Hordeum hystrix Roth
1	Native	Europe	Southwestern Europe	Spain		incl. Baleares	Tutin, T. G. et al., eds. 1964-1980. Flora europaea. Note: = Hordeum hystrix Roth
3	Adventive	Europe	Northern Europe	United Kingdom			Euro+Med Editorial Committee. Euro+Med Plantbase: the information resource for Euro-Mediterranean plant diversity (on-line resource).
4	Naturalized	Africa	Macaronesia	Portugal	Azores		Euro+Med Editorial Committee. Euro+Med Plantbase: the information resource for Euro-Mediterranean plant diversity (on-line resource).
4	Naturalized	Africa	Southern Africa	South Africa			Germishuizen, G. & N. L. Meyer, eds. 2003. Plants of southern Africa: an annotated checklist. Strelitzia 14 National Botanical Institute, Pretoria, South Africa.
4	Naturalized	Asia-Temperate	Eastern Asia	Japan			Koyama, T. 1987. Grasses of Japan and its neighboring regions
4	Naturalized	Australasia	Australia	Australia		s.	George, A. S., ed. 1980-. Flora of Australia.
4	Naturalized	Australasia	New Zealand	New Zealand			Allan, H. H. B. et al. 1961-. Flora of New Zealand.
4	Naturalized	Europe	Middle Europe	Germany			Euro+Med Editorial Committee. Euro+Med Plantbase: the information resource for Euro-Mediterranean plant diversity (on-line resource).
4	Naturalized	Europe	Northern Europe	Denmark			Euro+Med Editorial Committee. Euro+Med Plantbase: the information resource for Euro-Mediterranean plant diversity (on-line resource).
4	Naturalized	Northern America		Canada			Soreng, R. J. et al. 2003. Catalogue of New World grasses (Poaceae): IV. Subfamily Pooideae. Contr. U.S. Natl. Herb.
4	Naturalized	Northern America		Mexico			Soreng, R. J. et al. 2003. Catalogue of New World grasses (Poaceae): IV. Subfamily Pooideae. Contr. U.S. Natl. Herb.
4	Naturalized	Northern America		United States			FNA Editorial Committee. 1993-. Flora of North America.
4	Naturalized	Southern America	Southern South America	Chile			Soreng, R. J. et al. 2003. Catalogue of New World grasses (Poaceae): IV. Subfamily Pooideae. Contr. U.S. Natl. Herb.

Native
Africa

NORTHERN AFRICA: Algeria (n.), Egypt (n.), Libya (n.), Morocco, Tunisia

Asia-Temperate

WESTERN ASIA: Afghanistan, Cyprus, Iran, Iraq, Israel, Jordan, Lebanon, Syria, Turkey
CAUCASUS: Russian Federation-Ciscaucasia [Ciscaucasia (w.)], Azerbaijan, Georgia, Russian Federation [Dagestan]
MIDDLE ASIA: Kazakhstan (s.w.), Tajikistan, Turkmenistan, Uzbekistan

Asia-Tropical

INDIAN SUBCONTINENT: Pakistan (n.)

Europe

MIDDLE EUROPE: Austria, Czech Republic, Hungary, Slovakia
EASTERN EUROPE: Moldova, Russian Federation [Astrakhan, Volgogradskaja oblast'], Ukraine (incl. Krym)
SOUTHEASTERN EUROPE: Albania, Bulgaria, Greece (incl. Crete), Croatia, Italy (incl. Sardinia, Sicily), North Macedonia, Montenegro, Romania, Serbia, Slovenia
SOUTHWESTERN EUROPE: Spain (incl. Baleares), France (s. & Corsica), Portugal

Adventive
Europe

NORTHERN EUROPE: United Kingdom

Naturalized
Africa

MACARONESIA: Portugal [Azores]
SOUTHERN AFRICA: South Africa

Asia-Temperate

EASTERN ASIA: Japan

Australasia

AUSTRALIA: Australia (s.)
NEW ZEALAND: New Zealand

Europe

NORTHERN EUROPE: Denmark
MIDDLE EUROPE: Germany

Northern America

REGION: Canada, Mexico, United States

Southern America

SOUTHERN SOUTH AMERICA: Chile

Reference(s)

Allan, H. H. B. et al. 1961-. Flora of New Zealand.
Davis, P. H., ed. 1965-1988. Flora of Turkey and the east Aegean islands. Note: = Hordeum geniculatum All.
Euro+Med Editorial Committee. Euro+Med Plantbase: the information resource for Euro-Mediterranean plant diversity (on-line resource).
FNA Editorial Committee. 1993-. Flora of North America.
George, A. S., ed. 1980-. Flora of Australia.
Germishuizen, G. & N. L. Meyer, eds. 2003. Plants of southern Africa: an annotated checklist. Strelitzia 14 National Botanical Institute, Pretoria, South Africa.
Jakob, S. S. et al. 2007. Combined ecological niche modelling and molecular phylogeography revealed the evolutionary history of Hordeum marinum (Poaceae)-niche differentiation, loss of genetic diversity, and speciation in Mediterranean Quaternary refugia. Molec. Ecol. 16:1713-1727.
Koyama, T. 1987. Grasses of Japan and its neighboring regions
Meikle, R. D. 1977-1985. Flora of Cyprus. Note: = Hordeum geniculatum All.
Nasir, E. & S. I. Ali, eds. 1970-. Flora of [West] Pakistan.
Nishikawa, T. et al. 2002. Molecular phylogeny of the genus Hordeum using three chloroplast DNA sequences. Genome 45:1157-1166.
Sahebi, J. et al. 2004. A taxonomic review of the Hordeum marinum sensu lato (Poaceae: Triticeae) in Iran. Iran. J. Sci. Technol., Trans. A. Sci. 28:127-135.
Skvortsov, A. K., ed. 2006-. Flora of the Lower Volga region Note: treats as Hordeum marinum Huds.
Soreng, R. J. et al. 2003. Catalogue of New World grasses (Poaceae): IV. Subfamily Pooideae. Contr. U.S. Natl. Herb.
Townsend, C. C. & E. Guest. 1966-. Flora of Iraq. Note: = Hordeum geniculatum All.
Tutin, T. G. et al., eds. 1964-1980. Flora europaea. Note: = Hordeum hystrix Roth
Tzvelev, N. N. 1976. Zlaki SSSR

Cite as: USDA, Agricultural Research Service, National Plant Germplasm System. 2026. Germplasm Resources Information Network (GRIN Taxonomy). National Germplasm Resources Laboratory, Beltsville, Maryland.
URL: https://npgstest.ars-grin.gov/gringlobal/taxon/taxonomydetail?id=102508. Accessed 15 April 2026.