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ACCESSIONPLANT NAMETAXONOMYORIGINGENEBANKIMAGEAVAILABILITYRECEIVEDSOURCE TYPESOURCE DATECOLLECTION SITECOORDINATESELEVATIONHABITATIMPROVEMENT LEVELNARRATIVE
0GSHO 196vrs1.aHordeum vulgare L. subsp. vulgare Colorado, United StatesNSGCKERNELS1994DONATED05/1994Genetic materialAlleles at this complex locus modify development of the lateral spikelets and the associated lemma awn. The vrs1.a allele (v gene) is present in most six-rowed cultivars and produces well-developed lateral spikelets. The lemma awn of lateral spikelets will vary from 3/4 to nearly as long as those of central spikelets, depending upon alleles present at other loci. The Vrs1.b allele (V gene, distichon) is present in many two-rowed cultivars and reduces lateral spikelets to sterile bracts with a rounded tip. The Vrs1.t allele (Vt gene, deficiens) causes an extreme reduction in the size of lateral spikelets. The lr or vlr (vrs1.c) gene in Nudihaxtoni and Bozu types will not recombine with the vrs1.a allele and produces phenotypes similar to the Vrs1.d allele (Vd gene) of Svanhals. The series of induced mutants in two-rowed barley called hex-v and Int-d mutants differ in the size of lateral spikelets, but they interact with the vrs1.a allele as incomplete dominants. Many heterozygous combinations with vrs1.a have a pointed tip on the lemma of sterile lateral spikelets. Alleles at the int-c (intermedium spike-c) locus modify lateral size in the presence of vrs1.a, Vrs1.b, and Vrs1.d, but not when Vrs1.t is present. Inheritance: A multiple allelic series, incomplete dominant allele interactions based on the size and shape of lateral spikelets. Located in chromosome 2HL, about 30.5 cM distal from the eog (elongated outer glume) locus. Previous nomenclature and gene symbolization:Two-row vs six-row=Zz. Six-row vs two-row=Aa. Two-row=D. Six-row vs two-row=Vv. Six-row vs two-row (distichon) vs two-row (deficiens)=A,as,af. Redcd lateral spikelet append. on the lemma=lr. Allelic series v,Vd,V,Vt. Hex. mut. Intermed. spike-d=Int-d. Reduced lat. spike. append. on lemma-vlr. Mut.events:vrs1.a ... vrs1.o(v1b) in New Golden.Mut. used for descr.:vrs1.a...(GSHO 1907).1510659GSHO 196
1GSHO 773vrs2.eHordeum vulgare L. subsp. vulgare JapanNSGCKERNELS1994DEVELOPEDGenetic materialThe spike appears similar to the ordinary six-rowed spike, but the lateral spikelets on the upper and lower portion of the spike are reduced in size, less fertile, and more or less pedicellated. Laterals are pointed with reduced awn length or awnless. Inheritance: Monofactorial recessive. Located in chromosome 5HL [7L], over 19.0 cM proximal from the srh (short rachilla hair) locus. Previous nomenclature and gene symbolization: Six-rowed spike 2=v2. Mutational events: vrs2.e in Svanhals (Kmut 27, OUM284). Mutant used for description and seed stocks: vrs2.e in Svanhals (GSHO 773); vrs2.e in Bowman (PI 483237)*4 (GSHO 2111).1513794GSHO 773
2GSHO 774vrs3.fHordeum vulgare L. subsp. vulgare JapanNSGCKERNELS1994DEVELOPEDGenetic materialThe upper half of the spike appears to be six-rowed, but the lateral spikelets in the lower half are reduced in size with the basal portion being two-rowed. Awns on the lateral spikelets range from normal near the top, to awnletted or pointed in the middle, to awnless at the base of the spike. Lateral spikelets set seed only in the upper two-thirds of the spike. Lateral spikelets may be pedicellated in some stocks. Central spikelets often have double awn, one on the lemma and one on the palea. The rachilla may be deformed. Inheritance: Monofactorial recessive. Located in chromosome 1HL [5L]; near the centromere and proximal from the ert-b (erectoide-b) locus. Previous nomenclature and gene symbolization: Six-rowed spike 3=v3. Intermedium spike-a=int-1. Intermedium spike-g=int-g. Mutational events: vrs3.f in Hakata 2 (Kmut 213, OUM292), vrs3.g (Xb 388.8, Piro-1), vrs3.h (Xd 160.46, Piro-3), vrs3.i (X15 1463, Piro-5), vrs3.j (X17 3555, Piro-8) in Piroline (PI 539132); int-a.1 in Bonus (PI 189763); int-a.2 in Bonus; int-a.8, -a.9, -a.10, -a.64 in Bonus, -a.14, -a.17, -a.21, -a.27, -a.30, -a.31, -a.32, -a.34, -a.35, -a.37 in Foma (CIho 11333), -a.46, -a.51, -a.52, -a.54, -a.55, -a.59, -a.61 in Kristina; int-a.71, -a.74, -a.77, -a.79, -a.86, -a.88 in Bonus, -a.102, -a.103 in Hege. Mutant used for description and seed stocks: vrs3.f in Hakata 2 (GSHO 774); int-a.1 in Bonus (GSHO 170); vrs3.f in Bowman (PI 483237)*7 (GSHO 2056); int-a.1 in Bowman*5 (GSHO 2055).1513988GSHO 774
3GSHO 775vrs4.kHordeum vulgare L. subsp. vulgare Japan Historic1994DEVELOPEDGenetic materialAlleles at this locus enhance the development of lateral spikelets and formation of additonal spikelets to various degrees. The int-e mutants, which were isolated in two-rowed cultivars, have enlarged lateral spikelets that may set seed in the upper two-thirds of the spike. Kernels in lateral spikelets are smaller than those from central spikelets in int-e mutants. The rachilla may be deformed by partial formation of an extra spikelet. The awn size on lateral spikelets range from a pointed apex to 3/4 normal length. The mul1.a and vrs4.k alleles, which were isolated in six-rowed cultivars, may produce numerous extra spikelets at the base of the lateral spikelets and on the rachilla. Phenotypic differences among alleles are retained in the Bowman backcross-derived lines. Inheritance: Monofactorial recessive. Located in chromosome 3HL, about 27.5 cM distal from the uzu (uzu) locus. Previous nomenclature and gene symbolization: Multiflorus=mul. Intermedium spike-e=int-e. Six-rowed spike 4=v4. Mutational events: mul1.a (Alb Acc 321, GSHO 561) in Montcalm (CIho 7149); vrs4.k (Gamma III 3B, MFB-2) in MFB 104 (PI 232921), vrs4.l (Xc 41.5, Piro-2) in Piroline (PI 262210); vrs4.m (X17 2497, Piro-7) in Piroline; int-e.4 (trans) in Bonus (PI 189763), -e.65 in Bonus; int-e.72, -e.87 in Bonus, -e.89, -e.90, -e.91, -e.92, -e.101 in Hege. Mutant used for description and seed stocks: vrs4.k in Gamma III 3B (GSHO 775); int-e.58 in Kristina (GSHO 1766); mul1.a in Bowman (PI 483237)*6 (GSHO 1985); vrs4.k in Bowman*6 (GSHO 1986); int-e.58 in Bowman*6 (GSHO 1987).1513925GSHO 775