| 0 | GSHO 112 | fch1.a | Hordeum vulgare L. subsp. vulgare | Colorado, United States | NSGC |  | | 1994 | DONATED | 05/1994 | | | | | Genetic material | Seedling and immature leaves are very pale yellow-green (chlorophyll deficient). The leaves gradually change to a green color beginning with the leaf tip. Plants may grow to maturity in the field, but they have a slightly yellow-green color and are stunted easily by stress. Inheritance: Monofactorial recessive. Located in chromosome 2HS, in the centromeric region and about 4.0 cM distal from the eog (elongated outer glume) locus. Previous nomenclature and gene symbolization: Chlorina seedling=f f. Light green=lg. Light green 8=lg8. Light green 10=lg10. Mutational events: fch1.a in Gull; fch1.p (lg) (Minn 80, GSHO 548); fch1.q (lg8) (No 155, GSHO 8); and fch1.r (lg10)(No 157, GSHO 154) in Himalaya (CIho 1312). Mutant used for description and seed stocks: fch1.a in Minn 84-7 (GSHO 112); fch1.a in Bowman (PI 483237)*9 (GSHO 1886). | 1510685 | GSHO 112 |
| 1 | GSHO 1379 | gpa1.b | Hordeum vulgare L. subsp. vulgare | Idaho, United States | NSGC |  | | 1994 | DEVELOPED | | | | | | Genetic material | Seedlings display a pattern of transverse or alternating white and green bands on the first, second, and occasionally the third foliage leaves. Plants have a slightly pale green color prior to heading. Grandpa plants are sensitive to flooding and produce an albino flag leaf, peduncle, and spike. Plants are viable in the field, but the seed is thin and yields are low. Inheritance: Monofactorial recessive. Located in chromosome 2HL, about 24.5 cM distal from the lig (liguleless) locus. Previous nomenclature and gene symbolization: Grandpa=gp. Grandpa 2=gp2. Mutational events: gpa1.a in Lyallpur (GSHO 519), CIho 6854); gpa1.b (gp2) in Montcalm (CIho 7149). Mutant used for description and seed stocks: gpa1.b in Montcalm (GSHO 1379); gpa1.b in Bowman (PI 483237)*7 (GSHO 1934). | 1514266 | GSHO 1379 |
| 2 | GSHO 153 | vrs1.c | Hordeum vulgare L. subsp. vulgare | Colorado, United States | NSGC |  | | 1994 | DONATED | 05/1994 | | | | | Genetic material | The spike has long awns on the lemma of the central spikelet and awn-like appendages on the lemma of the lateral spikelets. Lateral spikelets have perfect flowers and complete fertility. The lr trait is expressed only in six-rowed barley, and heterozygotes have a short awn on the lemma of lateral spikelets. Since recombination with neither vrs1.a(v) nor Vrs1.b (V) alleles was observed, the lr or vrs1.c phenotype is likely controlled by an allele at the vrs1 locus. The Bowman backcross- derived line for the vrs1.c gene has short, stiff awns on the central spikelets (about 1/3 normal length) and no awns on the lateral spikelets.Inheritance: An incomplete dominant in the multiple allelic series at the vrs1 (six-rowed spike 1) locus. Located in chromosome 2HL, about 15.4 cM distal from the Gth (toothed lemma) locus, and 26.5 cM distal from the eog (elongated outer glume) locus. Previous nomenclature and gene symbolization: Reduced lateral spikelet appendage on the lemma=lr. Reduced lateral spikelet appendage on the lemma=vlr. Mutational events: vrs1.c or lr in Hordeum intermedium var Nudihaxtoni. Mutant used for description and seed stocks: vrs1.c or lr in Hordeum intermedium var Nudihaxtoni (PI 32368, GSHO 153); vrs1.c in Bowman (PI 483237)*7(GSHO 1908). | 1510717 | GSHO 153 |
| 3 | GSHO 171 | lgn3.c | Hordeum vulgare L. subsp. vulgare | Minnesota, United States | NSGC |  | | 1994 | DEVELOPED | | | | | | Genetic material | Seedlings have light or pale green color. The pale green color persists until near maturity, and the plants are very weak or non-viable in the field. Plants can be grown to maturity in greenhouses, but tillering is much reduced and seeds are very thin. Inheritance: Monofactorial recessive. Located in chromosome 4HL, over 10.5 cM distal from the Kap (hooded lemma) locus. Previous nomenclature and gene symbolization: Light green 7=lg7. Light green 3=lg3, the lg3 stock studied by Hanson and Kramer may not be the same as the lg3 (GSHO 171) stock. Mutational events: lgn3.c in an unknown cultivar (No 150); lgn3.g (lg7) in an unknown cultivar (GSHO 173, No 154). Mutant used for description and seed stocks: lgn3.c in an unknown cultivar (GSHO 171); lgn3.c in Bowman (PI 483237)*5 (GSHO 2010). | 1510675 | GSHO 171 |
| 4 | GSHO 196 | vrs1.a | Hordeum vulgare L. subsp. vulgare | Colorado, United States | NSGC |  | | 1994 | DONATED | 05/1994 | | | | | Genetic material | Alleles at this complex locus modify development of the lateral spikelets and the associated lemma awn. The vrs1.a allele (v gene) is present in most six-rowed cultivars and produces well-developed lateral spikelets. The lemma awn of lateral spikelets will vary from 3/4 to nearly as long as those of central spikelets, depending upon alleles present at other loci. The Vrs1.b allele (V gene, distichon) is present in many two-rowed cultivars and reduces lateral spikelets to sterile bracts with a rounded tip. The Vrs1.t allele (Vt gene, deficiens) causes an extreme reduction in the size of lateral spikelets. The lr or vlr (vrs1.c) gene in Nudihaxtoni and Bozu types will not recombine with the vrs1.a allele and produces phenotypes similar to the Vrs1.d allele (Vd gene) of Svanhals. The series of induced mutants in two-rowed barley called hex-v and Int-d mutants differ in the size of lateral spikelets, but they interact with the vrs1.a allele as incomplete dominants. Many heterozygous combinations with vrs1.a have a pointed tip on the lemma of sterile lateral spikelets. Alleles at the int-c (intermedium spike-c) locus modify lateral size in the presence of vrs1.a, Vrs1.b, and Vrs1.d, but not when Vrs1.t is present. Inheritance: A multiple allelic series, incomplete dominant allele interactions based on the size and shape of lateral spikelets. Located in chromosome 2HL, about 30.5 cM distal from the eog (elongated outer glume) locus. Previous nomenclature and gene symbolization:Two-row vs six-row=Zz. Six-row vs two-row=Aa. Two-row=D. Six-row vs two-row=Vv. Six-row vs two-row (distichon) vs two-row (deficiens)=A,as,af. Redcd lateral spikelet append. on the lemma=lr. Allelic series v,Vd,V,Vt. Hex. mut. Intermed. spike-d=Int-d. Reduced lat. spike. append. on lemma-vlr. Mut.events:vrs1.a ... vrs1.o(v1b) in New Golden.Mut. used for descr.:vrs1.a...(GSHO 1907). | 1510659 | GSHO 196 |
| 5 | GSHO 210 | trp1.a | Hordeum vulgare L. subsp. vulgare | Colorado, United States | NSGC |  | | 1994 | DONATED | 05/1994 | | | | | Genetic material | Mutants may have three awns on the lemma instead of one, but expression is variable. The awn or hood on the lemma of the central spikelet forks to produce one normal central appendage and one or two shorter lateral ones. Expression in the Bowman backcross-derived line is reduced to an occasional branch on the awn. Inheritance: Monofactorial recessive. Located in chromosome 2HL, over 35.5 cM distal from the wst7 (white streak 7) locus. Previous nomenclature and gene symbolization: Triple- bearded variation. Triple awned lemma=tr. Mutational events: trp1.a in Triple-awn lemma (CIho 6630). Mutant used for description and seed stocks: trp1. in CIho 6630 (GSHO 210); trp1.a from CIho 6880 (Ahang/ Twisted Flag//Triple-awn Lemma) in Bowman (PI 483237)*7 (GSHO 1936). | 1510639 | GSHO 210 |
| 6 | GSHO 29 | eog1.a | Hordeum vulgare L. subsp. vulgare | California, United States | NSGC |  | | 1994 | DEVELOPED | | | | | | Genetic material | A multiple allelic series exists at this locus which controls increased size of the outer glumes (glumes) of the spikelet. Glume width varies from 2.5 to 4.0 mm depending on genetic background and specific allele. Glume awn length varies from awnless to nearly as long as the lemma awn (92 mm) depending on the eog allele present and genes at other loci, but some confusion has occurred regarding alleles at the eog locus and expression of other glume characteristics. The size of the glume may range from less than twice normal width in CIho 14955 to lemma-like glumes in Triple Bearded Mariout. Inheritance: Monofactorial recessive. Located in chromosome 2HL, about 30.5 cM proximal from the vrs1 (six-rowed spike 1) locus. Previous nomenclature and gene symboliation: Extended outer glumes=e. Wide glumes=w. Macrolepis=lep-e(lep-a). Long outer glume=log. Elongated outer glume=e. Mutational events: eog1.a in Triple Bearded Mariout (CIho 2523); lep-e.1, -e.2 in Gull (CIho 1145), -e.3, -e.4, -e.5, -e.6, -e.7, -e.8, -e.9, -e.10, -e.11, -e.12, -e.13, -e.29, -e.34, -e.42, -e.45, -e.49, -e.50, -e.51, -e.52, -e.53, -e.54 in Bonus (PI 189763), -e.14, -e.15, -e.16, -e.17, -e.18, -e.19, -e.20, -e.21, -e.22, -e.23, -e.24, -e.25, -e.26, -e.27, -e.28, -e.30, -e.31, -e.32, -e.33, -e.38 in Foma (CIho 11333), -e.35, -e.36, -e.37, -e.39, -e.40, -e.41, -e.42, -e.43, -e.44, -e.46, -e.47, -e.48 in Kristina; eog 1.b in PI 173567 (G 8); eog1.c in CIho 14955; eog1.d (Mut 3447/64) in Proctor (PI 280420). Mutant used for description and seed stocks: eog1.a in Triple Bearded mariout (GSHO 29); lep-e.1 in Gull (GSHO 960); eog1.a in Bowman (PI 483237)*7(GSHO 1891); eog1.c from CIho 14955 in Bowman*7 (GSHO 1892). | 1510696 | GSHO 29 |
| 7 | GSHO 68 | yvs1.a | Hordeum vulgare L. subsp. vulgare | Colorado, United States | NSGC |  | | 1994 | DONATED | 05/1994 | | | | | Genetic material | Virescent seedlings have a slightly green tip, but fail to survive beyond the seedling stage. Recessive alleles at the yvs1 locus must be maintained as heterozygous stocks. Inheritance: Monofactorial recessive. Located in chromosome 2HS, over 13.7 cM proximal from the fch15 (chlorina seedling 15) locus and about 2.0 cM proximal from the fch1 (chlorina seedling 1) locus. Previous nomenclature and gene symbolization: Yellow seedling 1=y. Xantha (yellow) seedling x=yx. Yellow seedling x=yx. Albino seedling-c=alb-c. Mutational events: yvs1.a in a Swedish cultivar; yvs1.b (yx)(GSHO 62) in Faust I (CIho 4579); alb-c.7 in Gull (CIho 1145). Mutant used for description and seed stocks: yvs1.a in Minn 72-8 (GSHO 68). | 1510837 | GSHO 68 |
| 8 | GSHO 681 | lgn4.d | Hordeum vulgare L. subsp. vulgare | Minnesota, United States | NSGC |  | | 1994 | DEVELOPED | | | | | | Genetic material | Seedlings are light green in color, and the light green color persists until near maturity. In the Bowman backcross-derived lines, the plants are weaker than normal sibs and produce thinner seeds. Inheritance: Monofactorial recessive. Located in chromosome 4HL, about 5.0 cM distal from the Kap (hooded lemma) locus and about 4.0 cM proximal from the glf1 (glossy leaf 1) locus. Previous nomenclature and gene symbolization: Light green 1=lg1. Light green 4=lg4. Light green 9=lg9. Mutational events: lgn4.d (lg1) in Himalaya X Ingrescens; lgn4.i (lg9) in No 156, an unknown cultivar (GSHO 1, GSHO 94). Mutant used for description and seed stocks: lgn4.d in Himalaya X Ingrescens (GSHO 681); lgn4.d in Bowman (PI 483237)*7 (GSHO 2011). | 1512411 | GSHO 681 |
| 9 | GSHO 851 | fch3.d | Hordeum vulgare L. subsp. vulgare | Minnesota, United States | NSGC |  | | 1994 | DEVELOPED | | | | | | Genetic material | Seedlings have bright yellow leaves, which gradually change color to a light green as the plant grows. Plants are vigorous, but remain chlorina or light green until maturity. Plants expressing the fch3.d gene can be distinguished readily from normal plants at all stages of growth. Inheritance: Monofactorial recessive. Located in chromosome 1HS [5S]. Previous nomenclature and gene symbolization: Chlorina seedling 3=f3. Mutational events: fch3.d in an unknown cultivar (Minn 89-4). Mutant used for description and seed stocks: fch3.d in an unknown cultivar (GSHO 851); fch3.d in Bowman (PI 483237)*6 (GSHO 2033). | 1513938 | GSHO 851 |
| 10 | GSHO 93 | zeb2.b | Hordeum vulgare L. subsp. vulgare | Minnesota, United States | NSGC |  | | 1994 | DEVELOPED | | | | | | Genetic material | Leaves are chlorina throughout the life cycle, independent of light and temperature. Plants may be killed by low temperature during the winter when normal plants are unaffected. A zonate pattern of different degrees of chlorina is observed in seedling leaves. Plants are weak or non-viable, dependent on level of environmental stress. After much confusion about this gene, the locus was named zebra stripe 2=zb2. Inheritance: Monofactorial recessive. Located in chromosome 4HL, about 17.0 cM proximal from the glf3 (glossy leaf 3) locus. Previous nomenclature and gene symbolization: Zoned leaf=zd. Zoned leaf=z. Zoned leaf c-zbc. Zebra leaf or zoned Colorado=zc. Zebra striped leaves=zd. Zebra stripe 2=zb2. Chlorina seedling 10=f10. Mutational events: zeb2.b in an unknown cultivar; fch10.s in an unknown cultivar (GSHO 1234); both stocks may have originated from the same mutant used by Woodward and first isolated by Immer and Henderson. Mutant used for description and seed stocks: zeb2.b in an unknown cultivar (GSHO 93); zeb2.b in Bowman (PI 483237)*4 (GSHO 2012). | 1510745 | GSHO 93 |
| 11 | GSHO 98 | glf1.a | Hordeum vulgare L. subsp. vulgare | Colorado, United States | NSGC |  | | 1994 | DEVELOPED | | | | | | Genetic material | Surface wax coating on the leaf blade appears to be absent from the seedling stage to near maturity, and leaves have a shiny appearance (wax code ++ ++ -). Plants are semidwarf, relatively weak, and late in heading. The stock in the Bonus is highly sterile, but the Bowman backcross-derived line has nearly complete fertility. Inheritance: Monofactorial recessive. Located in chromosome 4HL, about 7.5 cM distal from the lbi2 (long basal rachis internode 2) locus, and about 4.8 cM distal from the Mlg (reg2, reaction to Erysiphe graminis) locus. Previous nomenclature and gene symbolization: Waxless bloom on leaves=w1. Glossy=gl. Glossy leaves=gl. Glossy leaf=gl. Glossy seedling 2=gl2. Eceriferum-zh=cer-zh. Mutational events: glf1.a, glf1.b (gl2, GSHO 22) in Himalaya; glf1.f in 34-119-1, glf1.g in II-34-199-7-2 (GSHO 89), cer-zh.54 in Bonus; cer-zh.266, -zh.308, -zh.357, -zh.366, -zh.432, -zh.433 in Foma (CIho 11333); cer-zh.325 in Foma; cer-zh.373 in Foma; cer-zh.865 in Bonus. Mutant used for description and seed stocks: glf1.a in Himalaya (GSHO 98); cer-zh.54 in Bonus (GSHO 455) is used for allelism tests; glf1.a in Bowman (PI 483237)*8 (GSHO 2015). | 1510793 | GSHO 98 |